146 lines
18 KiB
XML
146 lines
18 KiB
XML
<document id="D1B80DC221876F9B9BD7D194438AC309" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFEAFFE86194FB5A006CF6F2" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus diffusus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="24" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="23" updateTime="1698978858223" updateUser="plazi">
|
||
<mods:mods id="B5E28B98E9D99C4D395826840B246DB4" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CF22A89DF25FA15DDAC778C766C44F10">
|
||
<mods:title id="8BA07F594A3BAF937AF4A738EA5C67E9">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="F7ECBDA9378CC519698571E8EC2D25A6" type="personal">
|
||
<mods:role id="ADB89CA65D6341A15F6CF3EA36E409C8">
|
||
<mods:roleTerm id="F19F8FB374C8699BC8B178EA0EAF942B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="122C1D948FE067C66B1E4CB035E56585">Perkins, Philip D.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3C3107C5A35F401E94766ADA14F6EC77">text</mods:typeOfResource>
|
||
<mods:relatedItem id="AFBE6AA5BB97B783E62A9F7C269D438B" type="host">
|
||
<mods:titleInfo id="A03C8F2AD3C28B9FE7AB5B8C2E9782E3">
|
||
<mods:title id="1171BD1FBF5B76EFC3C73A2A8E4CAC79">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B2837B3DF6A964F5F8C3F63A402B862F">
|
||
<mods:date id="0F3B508C60CFEADF4E0B59EB9FB92920">2021</mods:date>
|
||
<mods:detail id="6F1948DF66FE914715B58403297AC9DB" type="pubDate">
|
||
<mods:number id="60F63463B5A3BBCD9B30B922CB484186">2021-06-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="414ED88811A703ACA3D5D95532BC2C12" type="volume">
|
||
<mods:number id="00770EC99123AA8911437C98C4AA03F3">4994</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0D421C05581A632408A1AB378DF38013" type="issue">
|
||
<mods:number id="8FE76183B3190DAB7D4D211B8B189CF3">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1033E6D20ACF2DCAA0800D7A1953261D" unit="page">
|
||
<mods:start id="39462205B504D0CA82EF4A7D412A496F">1</mods:start>
|
||
<mods:end id="AC7CD220187B524C287B1AC8CFBDB9E0">93</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BC26B5A1F2A3EE77696F94FDE13E2BBA">journal article</mods:classification>
|
||
<mods:identifier id="764B6579CF2A58A93FE307A194568076" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
|
||
<mods:identifier id="9E96A2E8EAD49DFA8B3ECA0B1B72CCAD" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="160BF50CCC26732DE6F3212F46277ED8" type="Zenodo-Dep">5043261</mods:identifier>
|
||
<mods:identifier id="B64C891DD5C7ADD4ECEF197AEA3751BF" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03A187DEFFEAFFE86194FB5A006CF6F2" ID-DOI="http://doi.org/10.5281/zenodo.5043954" ID-Zenodo-Dep="5043954" LSID="urn:lsid:plazi:treatment:03A187DEFFEAFFE86194FB5A006CF6F2" httpUri="http://treatment.plazi.org/id/03A187DEFFEAFFE86194FB5A006CF6F2" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
<subSubSection id="C3126543FFEAFFE96194FB5A067BF3C3" box="[151,548,1194,1221]" pageId="22" pageNumber="23" type="nomenclature">
|
||
<paragraph id="8BB736C8FFEAFFE96194FB5A067BF3C3" blockId="22.[151,548,1194,1257]" box="[151,548,1194,1221]" pageId="22" pageNumber="23">
|
||
<heading id="D0FF81A4FFEAFFE96194FB5A067BF3C3" bold="true" box="[151,548,1194,1221]" fontSize="11" level="2" pageId="22" pageNumber="23" reason="4">
|
||
<emphasis id="B97CEADAFFEAFFE96194FB5A067BF3C3" bold="true" box="[151,548,1194,1221]" pageId="22" pageNumber="23">
|
||
<taxonomicName id="4C084D4BFFEAFFE96194FB5A05DAF3C2" ID-CoL="8Y4PW" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,389,1194,1221]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="diffusus" status="sp. nov.">
|
||
<emphasis id="B97CEADAFFEAFFE96194FB5A05DAF3C2" bold="true" box="[151,389,1194,1221]" italics="true" pageId="22" pageNumber="23">Hydrochus diffusus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A24F57A1FFEAFFE96097FB5A067BF3C3" box="[404,548,1194,1220]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126543FFEAFFE96194FB3E053EF3EF" box="[151,353,1230,1257]" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="8BB736C8FFEAFFE96194FB3E053EF3EF" blockId="22.[151,548,1194,1257]" box="[151,353,1230,1257]" pageId="22" pageNumber="23">
|
||
<figureCitation id="13332A4DFFEAFFE96194FB3E04BAF3EF" box="[151,229,1230,1256]" captionStart="FIGURE 2" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[164,1405,189,1904]" captionTargetId="figure-15@7.[153,1434,181,1919]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043267" httpUri="https://zenodo.org/record/5043267/files/figure.png" pageId="22" pageNumber="23">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13332A4DFFEAFFE961F2FB3E0550F3EF" box="[241,271,1230,1256]" captionStart="FIGURES 26–27" captionStartId="65.[151,264,1944,1968]" captionTargetBox="[164,1427,199,1874]" captionTargetId="figure-15@65.[151,1435,181,1920]" captionTargetPageId="65" captionText="FIGURES 26–27. Habitus and male genitalia of holotypes. 26. Hydrochus robustus n. sp. 27. Hydrochus diffusus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043310" httpUri="https://zenodo.org/record/5043310/files/figure.png" pageId="22" pageNumber="23">27</figureCitation>
|
||
,
|
||
<figureCitation id="13332A4DFFEAFFE96018FB3E0566F3EE" box="[283,313,1230,1257]" captionStart="FIGURES 59–60" captionStartId="82.[151,264,1677,1701]" captionTargetBox="[157,1431,181,1654]" captionTargetId="figure-28@82.[157,1431,181,1654]" captionTargetPageId="82" captionText="FIGURES 59–60. Geographical distributions. 59. Endemics and near endemics. 60. H. formosus." figureDoi="http://doi.org/10.5281/zenodo.5043360" httpUri="https://zenodo.org/record/5043360/files/figure.png" pageId="22" pageNumber="23">59</figureCitation>
|
||
,
|
||
<figureCitation id="13332A4DFFEAFFE96046FB3E053EF3EF" box="[325,353,1230,1256]" captionStart="FIGURE 71" captionStartId="90.[151,250,1796,1820]" captionTargetBox="[166,1425,227,1738]" captionTargetId="figure-28@90.[151,1435,181,1773]" captionTargetPageId="90" captionText="FIGURE 71. Pictorial aid to species groups of Hydrochus from Bolivia, Brazil and Paraguay, using male genitalia. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.5043384" httpUri="https://zenodo.org/record/5043384/files/figure.png" pageId="22" pageNumber="23">71</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126543FFEAFFE96194FAE60558F253" pageId="22" pageNumber="23" type="materials_examined">
|
||
<paragraph id="8BB736C8FFEAFFE96194FAE60558F253" blockId="22.[151,1437,1302,2013]" pageId="22" pageNumber="23">
|
||
<emphasis id="B97CEADAFFEAFFE96194FAE60514F237" bold="true" box="[151,331,1302,1328]" pageId="22" pageNumber="23">Type Material.</emphasis>
|
||
<materialsCitation id="3B603C95FFEAFFE96056FAE60558F253" collectingDate="1985-10-03" collectingDateMax="1985-10-04" collectingDateMin="1985-10-03" collectionCode="NMNH" collectorName="Scott E. Miller" country="Brazil" elevation="1100" location="Paraiso" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
|
||
<emphasis id="B97CEADAFFEAFFE96056FAE6067EF237" bold="true" box="[341,545,1302,1328]" pageId="22" pageNumber="23">
|
||
<typeStatus id="54B3886AFFEAFFE96056FAE6059DF237" box="[341,450,1302,1328]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
|
||
(male):
|
||
</emphasis>
|
||
“
|
||
<collectingCountry id="F31F7658FFEAFFE96336FAE606FEF237" box="[565,673,1302,1328]" name="Brazil" pageId="22" pageNumber="23">BRAZIL</collectingCountry>
|
||
:
|
||
<collectingRegion id="49CCF82AFFEAFFE963ACFAE606ACF237" box="[687,755,1302,1328]" country="Brazil" name="Goias" pageId="22" pageNumber="23">Goias</collectingRegion>
|
||
,
|
||
<quantity id="4CF09B2DFFEAFFE96202FAE6076CF237" box="[769,819,1302,1328]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="km" value="7.0">7km</quantity>
|
||
NE
|
||
<location id="8ED76013FFEAFFE9626FFAE6079EF237" LSID="urn:lsid:plazi:treatment:03A187DEFFEAFFE86194FB5A006CF6F2:8ED76013FFEAFFE9626FFAE6079EF237" box="[876,961,1302,1328]" country="Brazil" name="Paraiso" pageId="22" pageNumber="23" stateProvince="Goias">Paraiso</location>
|
||
,
|
||
<quantity id="4CF09B2DFFEAFFE962D3FAE60040F237" box="[976,1055,1302,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="m" value="1100.0">
|
||
<elevation id="0025D1FBFFEAFFE962D3FAE60040F237" box="[976,1055,1302,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="22" pageNumber="23" unit="m" value="1100.0">1100m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB61008FFEAFFE9652DFAE60089F236" box="[1070,1238,1302,1329]" pageId="22" pageNumber="23" value="1985-10-03" valueMax="1985-10-04" valueMin="1985-10-03">
|
||
<collectingDate id="EFF2E9E0FFEAFFE9652DFAE60089F236" box="[1070,1238,1302,1329]" pageId="22" pageNumber="23" value="1985-10-03" valueMax="1985-10-04" valueMin="1985-10-03">3–4 Oct. 1985</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26FD531EFFEAFFE965E7FAE601CCF237" box="[1252,1427,1302,1329]" pageId="22" pageNumber="23">Scott E. Miller</collectorName>
|
||
” (
|
||
<collectionCode id="ED19AE0DFFEAFFE9619CFACA04A4F253" box="[159,251,1338,1364]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23" type="Museum">NMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126543FFEAFFE961C4FAAE07B8F2E3" pageId="22" pageNumber="23" type="diagnosis">
|
||
<paragraph id="8BB736C8FFEAFFE961C4FAAE07B8F2E3" blockId="22.[151,1437,1302,2013]" pageId="22" pageNumber="23">
|
||
<emphasis id="B97CEADAFFEAFFE961C4FAAE058DF27F" bold="true" box="[199,466,1374,1400]" pageId="22" pageNumber="23">Differential Diagnosis.</emphasis>
|
||
Among Brazilian species, recognized by the combination of moderately large size (ca.
|
||
<quantity id="4CF09B2DFFEAFFE96194FA72055FF29A" box="[151,256,1410,1437]" metricMagnitude="-3" metricUnit="m" metricValue="4.26" pageId="22" pageNumber="23" unit="mm" value="4.26">4.26 mm</quantity>
|
||
), the moderately deep pronotal depressions, the rather uniform pronotal punctation, the elytra with small round widely separated black spots, and the male genitalia (
|
||
<figureCitation id="13332A4DFFEAFFE9622AFA560724F2C7" box="[809,891,1446,1472]" captionStart="FIGURES 26–27" captionStartId="65.[151,264,1944,1968]" captionTargetBox="[164,1427,199,1874]" captionTargetId="figure-15@65.[151,1435,181,1920]" captionTargetPageId="65" captionText="FIGURES 26–27. Habitus and male genitalia of holotypes. 26. Hydrochus robustus n. sp. 27. Hydrochus diffusus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043310" httpUri="https://zenodo.org/record/5043310/files/figure.png" pageId="22" pageNumber="23">Fig. 27</figureCitation>
|
||
; described below). This species has a generalized
|
||
<taxonomicName id="4C084D4BFFEAFFE96194FA3A054EF2E3" authorityName="Leach" authorityYear="1817" box="[151,273,1482,1508]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEADAFFEAFFE96194FA3A054EF2E3" box="[151,273,1482,1508]" italics="true" pageId="22" pageNumber="23">Hydrochus</emphasis>
|
||
</taxonomicName>
|
||
habitus; reliable determinations will require dissection of males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126543FFEAFFE861C4FA1E06D0F6D6" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="8BB736C8FFEAFFE961C4FA1E052CF1BB" blockId="22.[151,1437,1302,2013]" pageId="22" pageNumber="23">
|
||
<emphasis id="B97CEADAFFEAFFE961C4FA1E050EF10F" bold="true" box="[199,337,1518,1544]" pageId="22" pageNumber="23">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="54B3886AFFEAFFE960A7FA1E0659F10F" box="[420,518,1518,1544]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 4.26/1.44; head width 0.94; pronotum l/w 0.90/0.98; PA 0.94; PB 0.82; elytra 2.80/1.44. Habitus as illustrated (
|
||
<figureCitation id="13332A4DFFEAFFE9654DF9E200FBF12B" box="[1102,1188,1554,1580]" captionStart="FIGURES 26–27" captionStartId="65.[151,264,1944,1968]" captionTargetBox="[164,1427,199,1874]" captionTargetId="figure-15@65.[151,1435,181,1920]" captionTargetPageId="65" captionText="FIGURES 26–27. Habitus and male genitalia of holotypes. 26. Hydrochus robustus n. sp. 27. Hydrochus diffusus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043310" httpUri="https://zenodo.org/record/5043310/files/figure.png" pageId="22" pageNumber="23">Fig. 27</figureCitation>
|
||
). Body size large (ca.
|
||
<quantity id="4CF09B2DFFEAFFE96194F9C604A0F156" box="[151,255,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="4.26" pageId="22" pageNumber="23" unit="mm" value="4.26">4.26 mm</quantity>
|
||
). Dorsum grey to reddish grey, without iridescence, elytra with small round widely separated black spots. Legs testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2x that of pronotum. Elytra interstriae ca. 2x strial puncture diameter. Interstria 9
|
||
<superScript id="7C7D9B80FFEAFFE9634BF98E060AF18B" attach="left" box="[584,597,1662,1676]" fontSize="6" pageId="22" pageNumber="23">th</superScript>
|
||
slightly more convex than others, overhanging 10
|
||
<superScript id="7C7D9B80FFEAFFE96587F98E00CEF18B" attach="left" box="[1156,1169,1662,1676]" fontSize="6" pageId="22" pageNumber="23">th</superScript>
|
||
interstria. Usual area of callus on 5
|
||
<superScript id="7C7D9B80FFEAFFE96013F9520542F1B7" attach="left" box="[272,285,1698,1712]" fontSize="6" pageId="22" pageNumber="23">th</superScript>
|
||
absent.
|
||
</paragraph>
|
||
<paragraph id="8BB736C8FFEAFFE961C4F9370067F02E" blockId="22.[151,1437,1302,2013]" pageId="22" pageNumber="23">Pronotum slightly wider than long (ratio ca. 27/23), widest at anterior margin, narrowed at base, sides very weakly sinuate, smooth; anterior margin slightly bisinuate; lateral and anterior margins with row of short setae; depressions shallow to moderately deep; punctation rather uniform over pronotum.</paragraph>
|
||
<paragraph id="8BB736C8FFEAFFE961C4F8C20700F077" blockId="22.[151,1437,1302,2013]" pageId="22" pageNumber="23">Elytra with spaces between strial punctures ca. 1x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with slight angulation.</paragraph>
|
||
<paragraph id="8BB736C8FFEAFFE961C4F88A0556F0BF" blockId="22.[151,1437,1302,2013]" pageId="22" pageNumber="23">Ventral characters: Mentum punctate, with central depression; submentum with two foveae, small and widely separated.</paragraph>
|
||
<paragraph id="8BB736C8FFEAFFE861C4F83200A7F662" blockId="22.[151,1437,1302,2013]" lastBlockId="23.[151,1437,150,501]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
Male genitalia (
|
||
<figureCitation id="13332A4DFFEAFFE9607EF832058EF0DB" box="[381,465,1986,2012]" captionStart="FIGURES 26–27" captionStartId="65.[151,264,1944,1968]" captionTargetBox="[164,1427,199,1874]" captionTargetId="figure-15@65.[151,1435,181,1920]" captionTargetPageId="65" captionText="FIGURES 26–27. Habitus and male genitalia of holotypes. 26. Hydrochus robustus n. sp. 27. Hydrochus diffusus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043310" httpUri="https://zenodo.org/record/5043310/files/figure.png" pageId="22" pageNumber="23">Fig. 27</figureCitation>
|
||
) general characters: genitalia moderately stout, length ca. 3.8x greatest width, basal piece longer than parameres (ratio ca. 5/4); paramere tips and aedeagus tip at same level, paramere tips arcing toward midline, tips almost meeting, plsa absent, lateral margins of tips strongly arcuate; in lateral view parameres wide basally, tapering to rounded apex; aedeagus very wide in both dorsal/ventral and lateral views, oval, laterally heavily sclerotized, medial margins of sclerotized part together forming deep V-shape, agw at ca. midlength; basal piece in dorsal/ventral views tapering from distal end to base, in lateral view also tapering toward base, and arcuate, greatest width in dorsal/ventral views slightly greater than lateral width, orifice large, asymmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BB736C8FFEBFFE861C4FE9F0017F68E" blockId="23.[151,1437,150,501]" box="[199,1096,367,393]" pageId="23" pageNumber="24">Dorsal surface: adtl subapical; pdmm approximating alm; adbl small, triangular.</paragraph>
|
||
<paragraph id="8BB736C8FFEBFFE861C4FE6306D0F6D6" blockId="23.[151,1437,150,501]" pageId="23" pageNumber="24">Ventral surface: avtl not apparent; pvmm similar to pdmm in distal ½, in proximal ½ widely overlapping aedeagus, near base pvmm narrowly separated.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126543FFEBFFE861C4FE2B006CF6F2" box="[199,1075,475,501]" pageId="23" pageNumber="24" type="etymology">
|
||
<paragraph id="8BB736C8FFEBFFE861C4FE2B006CF6F2" blockId="23.[151,1437,150,501]" box="[199,1075,475,501]" pageId="23" pageNumber="24">
|
||
<emphasis id="B97CEADAFFEBFFE861C4FE2B0517F6F2" bold="true" box="[199,328,475,501]" pageId="23" pageNumber="24">Etymology</emphasis>
|
||
. Named in reference to the widely spaced black dots on the elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |