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<document id="2B346A13976A43473C7308CBF346FED9" ID-CLB-Dataset="22056" ID-DOI="10.5852/ejt.2016.177" ID-GBIF-Dataset="33a17d32-75b4-423c-bc63-4bd22ae66a4f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832770" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589832188929" checkinUser="valdenar" docAuthor="Enghoff, Henrik" docDate="2016" docId="0137952B753FCB116A02C4D8FB0740A8" docLanguage="en" docName="ejt-2016-177.pdf.imf" docOrigin="European Journal of Taxonomy 177" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Geotypodon submontanus Enghoff, 2016, gen. et sp. nov." docType="treatment" docUuid="02DBFF6A-56A1-4EB3-9001-AF063F22C1B6" docUuidSource="ZooBank" docVersion="9" lastPageNumber="14" masterDocId="FD0EED537534CB1C6847C63BFFC9414A" masterDocTitle="A mountain of millipedes III: A new genus for three new species from the Udzungwa Mountains and surroundings, Tanzania, as well as several ‘ orphaned’ species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="12" updateTime="1698840199972" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="EDAD1B845802426E56781AA62C9B3B9E">
<mods:title id="9078EC21B8DC84848965FBB5CF8ECE09">A mountain of millipedes III: A new genus for three new species from the Udzungwa Mountains and surroundings, Tanzania, as well as several ‘ orphaned’ species previously assigned to Odontopyge Brandt, 1841 (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
</mods:titleInfo>
<mods:name id="B847DD827E6C936F7904CC50DE059E57" type="personal">
<mods:role id="16161A0AC6CEFA2485754597FFC57211">
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<mods:namePart id="F26E50F3050B6A8EB6976427848FB3FB">Enghoff, Henrik</mods:namePart>
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<mods:title id="685A20A7149720A1C9AF867A96EEEC13">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="6EE11188C750ED89C1A071015211804E">
<mods:date id="9DEF527D95968E93F3EA8D1DE4ABC369">2016</mods:date>
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<mods:number id="BA857C6388A87AE9B7B08E87466FC528">2016-02-26</mods:number>
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<mods:number id="CBEFA049ED56DFA20FE025C888D7D661">177</mods:number>
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<mods:classification id="2E4494F42D690FA6F317B5D3016A7765">journal article</mods:classification>
<mods:identifier id="A620980DEA37D68BB6F1EAA2EF887D14" type="CLB-Dataset">22056</mods:identifier>
<mods:identifier id="F60FB28A593FE96EE459074103412FF3" type="DOI">10.5852/ejt.2016.177</mods:identifier>
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<mods:identifier id="A0B8DC24B540793F2AF0CA1063FD33E1" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="FB683DD8ABFDD04FF6FF7CF3AE8FD5D6" type="Zenodo-Dep">3832770</mods:identifier>
<mods:identifier id="0232E7C3655D279D199C7C99F15F97FF" type="ZooBank">A9E7A041-A454-4BC1-BCF4-F1E021BFDCD5</mods:identifier>
</mods:mods>
<treatment id="0137952B753FCB116A02C4D8FB0740A8" ID-DOI="http://doi.org/10.5281/zenodo.3850255" ID-GBIF-Taxon="164167483" ID-Zenodo-Dep="3850255" LSID="urn:lsid:zoobank.org:act:02DBFF6A-56A1-4EB3-9001-AF063F22C1B6" httpUri="http://treatment.plazi.org/id/0137952B753FCB116A02C4D8FB0740A8" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="C18477B6753FCB176A02C4D8FB224268" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8921243D753FCB176A02C4D8FBF143B4" blockId="11.[402,1259,739,837]" box="[581,1080,739,766]" pageId="11" pageNumber="12">
<heading id="D2699351753FCB176A02C4D8FBF143B4" box="[581,1080,739,766]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="4E9E5FBE753FCB176A02C4D8FC4843B4" ID-CoL="3FTNZ" authorityName="Enghoff" authorityYear="2016" box="[581,897,739,766]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="submontanus" status="gen. et sp. nov.">
<emphasis id="BBEAF82F753FCB176A02C4D8FC4843B4" bold="true" box="[581,897,739,766]" italics="true" pageId="11" pageNumber="12">Geotypodon submontanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D94554753FCB176BCFC4DFFBF143B4" box="[904,1080,740,766]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="FD0F283F753FCB1769D5C53CFB224268" box="[402,1259,775,802]" pageId="11" pageNumber="12">
urn:lsid:zoobank.org:act:
<uuid id="FD381EE8753FCB176AF2C53CFB224268" box="[693,1259,775,802]" pageId="11" pageNumber="12">02DBFF6A-56A1-4EB3-9001-AF063F22C1B6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C18477B6753FCB176ABAC511FC48420F" box="[765,897,810,837]" pageId="11" pageNumber="12" type="description">
<paragraph id="8921243D753FCB176ABAC511FC48420F" blockId="11.[402,1259,739,837]" box="[765,897,810,837]" pageId="11" pageNumber="12">
<figureCitation id="11A538B8753FCB176ABAC511FC8F420E" box="[765,838,810,836]" captionStart="Fig" captionStartId="1.[227,270,1894,1920]" captionTargetBox="[236,1429,925,1845]" captionTargetId="figure@1.[227,1436,918,1853]" captionTargetPageId="1" captionText="Fig. 1. Map of the Udzungwa Mountains, showing the collecting sites for the three new Geotypodon species, as well as names of the Forest Reserves in question and names of individual mountains in West Kilombero FR. Red diamonds = G. millemanus gen. et sp. nov., yellow dot: G. submontanus gen. et sp. nov., blue triangle: G. iringensis gen. et sp. nov. Based on fig. 1 in Marshall et al. (2010) and information in Doody et al. (2001)." figureDoi="http://doi.org/10.5281/zenodo.3832772" httpUri="https://zenodo.org/record/3832772/files/figure.png" pageId="11" pageNumber="12">Figs 1</figureCitation>
,
<figureCitation id="11A538B8753FCB176B13C511FCAD420E" box="[852,868,810,836]" captionStart="Fig" captionStartId="8.[227,270,1966,1992]" captionTargetBox="[329,1330,1159,1900]" captionTargetId="figure@8.[301,1353,1155,1914]" captionTargetPageId="8" captionText="Fig. 3. Body size of males of Geotypodon spp. Bold symbols indicate numbers of podous rings and midbody vertical diameter of the new species described here. Small circles and shaded areas indicate" figureDoi="http://doi.org/10.5281/zenodo.3832776" httpUri="https://zenodo.org/record/3832776/files/figure.png" pageId="11" pageNumber="12">3</figureCitation>
,
<figureCitation id="11A538B8753FCB176B35C511FC48420F" box="[882,897,810,837]" captionStart="Fig" captionStartId="12.[227,270,1862,1888]" captionTargetBox="[272,1377,302,1825]" captionTargetId="figure@12.[272,1377,302,1825]" captionTargetPageId="12" captionText="Fig. 5. Geotypodon submontanus gen. et sp. nov., holotype. A–E. Left gonopod. A. Anterior view. B. Posterior view. C. Apical part of coxa and proximal part of telopodite, anterior view. D. (Anterior-) mesal view. E. Posterior distal lobe of telomere; insertion highlights spine-like process. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mla = metaplical lamella; msp = metaplical spine-like process; ptl = posterior distal" figureDoi="http://doi.org/10.5281/zenodo.3832780" httpUri="https://zenodo.org/record/3832780/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C18477B6753FCB1768A6C550FCB0452E" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8921243D753FCB1768A6C550FE9342CF" blockId="11.[225,346,875,901]" box="[225,346,875,901]" pageId="11" pageNumber="12">
<heading id="D2699351753FCB1768A6C550FE9342CF" bold="true" box="[225,346,875,901]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="BBEAF82F753FCB1768A6C550FE9342CF" bold="true" box="[225,346,875,901]" pageId="11" pageNumber="12">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8921243D753FCB1768A6C5A1FCB0452E" blockId="11.[225,1437,922,1125]" pageId="11" pageNumber="12">
A species of
<taxonomicName id="4E9E5FBE753FCB17693CC5A1FDC342FE" authorityName="Enghoff" authorityYear="2016" box="[379,522,922,948]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BBEAF82F753FCB17693CC5A1FDC342FE" box="[379,522,922,948]" italics="true" pageId="11" pageNumber="12">Geotypodon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D94554753FCB176A54C5A0FDB642FF" box="[531,639,923,949]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
in which the gonopod coxal metaplica is produced mesad in a thin lamella, a small longitudinal lamella, perpendicular to the other, is present on the anterior part of the metaplica, the telomere is apically divided into two lobes of approximately equal size, and the limbus lobes are multi-cusped, hand-like. Shares these characters with
<taxonomicName id="4E9E5FBE753FCB176BABC23FFB524554" authorityName="Enghoff" authorityYear="2016" box="[1004,1179,1028,1054]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
<emphasis id="BBEAF82F753FCB176BABC23FFB524554" box="[1004,1179,1028,1054]" italics="true" pageId="11" pageNumber="12">G. millemanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D94554753FCB176CE0C23FFAAE4554" box="[1191,1383,1028,1054]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, but differs from that species in smaller size, bicuspid mesal margin of metaplical lamella and presence of a spine-like process on the posterior apical telomeral lobe.
</paragraph>
</subSubSection>
<subSubSection id="C18477B6753FCB1768A6C2B7FE0E44E5" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8921243D753FCB1768A6C2B7FEA145EC" blockId="11.[225,360,1164,1190]" box="[225,360,1164,1190]" pageId="11" pageNumber="12">
<heading id="D2699351753FCB1768A6C2B7FEA145EC" bold="true" box="[225,360,1164,1190]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="BBEAF82F753FCB1768A6C2B7FEA145EC" bold="true" box="[225,360,1164,1190]" pageId="11" pageNumber="12">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8921243D753FCB1768A6C281FC9B459E" blockId="11.[225,850,1210,1237]" box="[225,850,1210,1237]" pageId="11" pageNumber="12">The name is a Latin adjective referring to the habitat.</paragraph>
<paragraph id="8921243D753FCB1768A6C2C0FDF5445C" blockId="11.[225,572,1274,1302]" box="[225,572,1274,1302]" pageId="11" pageNumber="12">
<emphasis id="BBEAF82F753FCB1768A6C2C0FE78445F" bold="true" box="[225,433,1275,1301]" pageId="11" pageNumber="12">Material studied</emphasis>
(total: 1 ♂)
</paragraph>
<materialsCitation id="39F62E60753FCB176948C311FE0E44E5" ID-GBIF-Occurrence="2620529302" collectingDate="2000-11" collectionCode="ZMUC" collectorName="West Kilombero Scarp FR &amp; Ukami &amp; Frontier Tanzania" country="Tanzania" elevation="1145" latitude="-7.88875" location="West Kilombero Scarp" longLatPrecision="1" longitude="36.38656" municipality="Udzungwa Mts" pageId="11" pageNumber="12" specimenCode="ZMUC00046992" specimenCount="1" stateProvince="Iringa" typeStatus="holotype">
<paragraph id="8921243D753FCB176948C311FE48440E" blockId="11.[225,1437,1322,1455]" box="[271,385,1322,1348]" pageId="11" pageNumber="12">
<emphasis id="BBEAF82F753FCB176948C311FE48440E" bold="true" box="[271,385,1322,1348]" pageId="11" pageNumber="12">
<typeStatus id="56259A9F753FCB176948C311FE48440E" box="[271,385,1322,1348]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8921243D753FCB1768A6C375FE0E44E5" blockId="11.[225,1437,1322,1455]" pageId="11" pageNumber="12">
<collectingCountry id="F18964AD753FCB1768A6C375FEB24422" box="[225,379,1358,1384]" name="Tanzania" pageId="11" pageNumber="12">TANZANIA</collectingCountry>
: ♂,
<collectingRegion id="4B5AEADF753FCB1769E2C376FD81442D" box="[421,584,1357,1383]" country="Tanzania" name="Iringa" pageId="11" pageNumber="12">Iringa Region</collectingRegion>
,
<collectingRegion id="4B5AEADF753FCB176A17C376FD3F442D" box="[592,758,1357,1383]" country="Tanzania" name="Iringa" pageId="11" pageNumber="12">Iringa District</collectingRegion>
,
<collectingMunicipality id="6945BE47753FCB176AB8C376FC7A4422" box="[767,947,1357,1384]" pageId="11" pageNumber="12">Udzungwa Mts</collectingMunicipality>
,
<collectorName id="246B41EB753FCB176BFBC375FB254422" box="[956,1260,1357,1384]" pageId="11" pageNumber="12">
<location id="8C4172E6753FCB176BFBC375FB084422" LSID="urn:lsid:plazi:treatment:0137952B753FCB116A02C4D8FB0740A8:8C4172E6753FCB176BFBC375FB084422" box="[956,1217,1357,1384]" country="Tanzania" latitude="-7.88875" longLatPrecision="1" longitude="36.38656" municipality="Udzungwa Mts" name="West Kilombero Scarp" pageId="11" pageNumber="12" stateProvince="Iringa">West Kilombero Scarp</location>
FR
</collectorName>
,
<geoCoordinate id="ECAA42FA753FCB176CB2C376FA5E4422" box="[1269,1431,1357,1384]" degrees="07" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="12" precision="1" seconds="19.5" value="-7.88875">07°53'19.5&quot; S</geoCoordinate>
,
<geoCoordinate id="ECAA42FA753FCB1768A6C34BFE4F44C1" box="[225,390,1392,1419]" degrees="36" direction="east" minutes="23" orientation="longitude" pageId="11" pageNumber="12" precision="1" seconds="11.6" value="36.38656">36°23'11.6&quot; E</geoCoordinate>
, submontane forest,
<elevation id="02B3C30E753FCB176A30C34BFD3E44C1" box="[631,759,1392,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="11" pageNumber="12" unit="m" value="1145.0">
<quantity id="4E6689D8753FCB176A30C34BFD0744C1" box="[631,718,1392,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="11" pageNumber="12" unit="m" value="1145.0">1145 m</quantity>
asl
</elevation>
, trapsite
<collectorName id="246B41EB753FCB176B27C34AFC7A44C1" box="[864,947,1393,1419]" pageId="11" pageNumber="12">Ukami</collectorName>
, casual,
<date id="FD2002FD753FCB176C52C34AFB5B44C1" box="[1045,1170,1393,1419]" pageId="11" pageNumber="12" value="2000-11">
<collectingDate id="ED64FB15753FCB176C52C34AFB5B44C1" box="[1045,1170,1393,1419]" pageId="11" pageNumber="12" value="2000-11">Nov. 2000</collectingDate>
</date>
,
<collectorName id="246B41EB753FCB176CDBC34AFAA344C1" box="[1180,1386,1393,1419]" pageId="11" pageNumber="12">Frontier Tanzania</collectorName>
leg. (
<specimenCode id="D9388C46753FCB1768AEC3AFFE7244E5" box="[233,443,1428,1455]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="12">ZMUC00046992</specimenCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C18477B6753FCB1768A6C3EEFBFC470B" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8921243D753FCB1768A6C3EEFE4844A5" blockId="11.[225,385,1493,1519]" box="[225,385,1493,1519]" pageId="11" pageNumber="12">
<heading id="D2699351753FCB1768A6C3EEFE4844A5" bold="true" box="[225,385,1493,1519]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="BBEAF82F753FCB1768A6C3EEFE4844A5" bold="true" box="[225,385,1493,1519]" pageId="11" pageNumber="12">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8921243D753FCB1768A6C03FFBFC470B" blockId="11.[225,1437,1540,1602]" pageId="11" pageNumber="12">
<collectingCountry id="F18964AD753FCB1768A6C03FFEB04754" box="[225,377,1540,1566]" name="Tanzania" pageId="11" pageNumber="12">TANZANIA</collectingCountry>
,
<collectingRegion id="4B5AEADF753FCB1769C2C03FFDE24754" box="[389,555,1540,1566]" country="Tanzania" name="Iringa" pageId="11" pageNumber="12">Iringa Region</collectingRegion>
,
<collectingRegion id="4B5AEADF753FCB176A71C03FFD164754" box="[566,735,1540,1566]" country="Tanzania" name="Iringa" pageId="11" pageNumber="12">Iringa District</collectingRegion>
, Udzungwa Mts, West Kilombero Scarp FR,
<geoCoordinate id="ECAA42FA753FCB176CB5C03FFA5E4754" box="[1266,1431,1540,1567]" degrees="07" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="12" precision="1" seconds="19.5" value="-7.88875">07°53'19.5&quot; S</geoCoordinate>
,
<geoCoordinate id="ECAA42FA753FCB1768A6C01CFE4F4708" box="[225,390,1575,1602]" degrees="36" direction="east" minutes="23" orientation="longitude" pageId="11" pageNumber="12" precision="1" seconds="11.6" value="36.38656">36°23'11.6&quot; E</geoCoordinate>
, submontane forest,
<quantity id="4E6689D8753FCB176A3DC01CFD184708" box="[634,721,1575,1602]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="11" pageNumber="12" unit="m" value="1145.0">1145 m</quantity>
asl, cf.
<bibRefCitation id="ED0F59CC753FCB176B6AC01CFBF8470B" author="Marshall A. R. &amp; Brink H. &amp; Topp-Jorgensen J. E." box="[813,1073,1575,1602]" pageId="11" pageNumber="12" pagination="124 - 132" refId="ref8586" refString="Marshall A. R., Brink H. &amp; Topp-Jorgensen J. E. 2001. Millipede Diversity and Distribution - West Kilombero Scarp Forest Reserve. In: Doody K. Z., Howell K. M., &amp; Fanning E. (eds) West Kilombero Scarp Forest Reserve - Zoological Report: 124 - 132. Report for the Udzungwa Mountains Forest Management and Biodiversity Conservation Project, MEMA, Iringa, Tanzania." type="book chapter" year="2001">
Marshall
<emphasis id="BBEAF82F753FCB176BDAC013FC19470B" box="[925,976,1575,1601]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. (2001)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C18477B6753FCB1768A6C053FB2446CA" pageId="11" pageNumber="12" type="description">
<paragraph id="8921243D753FCB1768A6C053FEBB47C8" blockId="11.[225,370,1640,1666]" box="[225,370,1640,1666]" pageId="11" pageNumber="12">
<heading id="D2699351753FCB1768A6C053FEBB47C8" bold="true" box="[225,370,1640,1666]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="BBEAF82F753FCB1768A6C053FEBB47C8" bold="true" box="[225,370,1640,1666]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8921243D753FCB1768A6C0ACFAE647FB" blockId="11.[225,1327,1687,1713]" box="[225,1327,1687,1713]" pageId="11" pageNumber="12">
SIZE. Length ca.
<quantity id="4E6689D8753FCB1769E4C0ACFE2947FB" box="[419,480,1687,1713]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="12" unit="cm" value="7.0">7 cm</quantity>
. Diameter
<quantity id="4E6689D8753FCB176A27C0ACFD7447FB" box="[608,701,1687,1713]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="3.8">3.8 mm</quantity>
. 60 podous rings, no apodous rings in front of telson.
</paragraph>
<paragraph id="8921243D753FCB1768A6C0E3FB2C47B8" blockId="11.[225,1261,1752,1920]" box="[225,1253,1752,1779]" pageId="11" pageNumber="12">
OTHER CHARACTERS. As in
<taxonomicName id="4E9E5FBE753FCB176A49C0E2FD7047B8" authorityName="Enghoff" authorityYear="2016" box="[526,697,1752,1779]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="millemanus" status="gen. et sp. nov.">
<emphasis id="BBEAF82F753FCB176A49C0E2FD7047B8" box="[526,697,1752,1779]" italics="true" pageId="11" pageNumber="12">G. millemanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D94554753FCB176A86C0E2FCBA47B9" box="[705,883,1753,1779]" pageId="11" pageNumber="12" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, with the following exceptions:
</paragraph>
<paragraph id="8921243D753FCB1768A6C0C7FD26465F" blockId="11.[225,1261,1752,1920]" box="[225,751,1787,1814]" pageId="11" pageNumber="12">- telson (after 15 years in alcohol) yellowish,</paragraph>
<paragraph id="8921243D753FCB1768A6C124FCF94673" blockId="11.[225,1261,1752,1920]" box="[225,816,1823,1849]" pageId="11" pageNumber="12">- postfemoral and tibial pads on male legs smaller,</paragraph>
<paragraph id="8921243D753FCB1768A6C179FB4C4617" blockId="11.[225,1261,1752,1920]" box="[225,1157,1858,1885]" pageId="11" pageNumber="12">
- metaplical lamella (
<emphasis id="BBEAF82F753FCB17699CC179FDCE4616" box="[475,519,1858,1884]" italics="true" pageId="11" pageNumber="12">mla</emphasis>
) produced mesad, with two sharp angles (
<figureCitation id="11A538B8753FCB176BB0C179FBF54617" box="[1015,1084,1858,1885]" captionStart="Fig" captionStartId="12.[227,270,1862,1888]" captionTargetBox="[272,1377,302,1825]" captionTargetId="figure@12.[272,1377,302,1825]" captionTargetPageId="12" captionText="Fig. 5. Geotypodon submontanus gen. et sp. nov., holotype. A–E. Left gonopod. A. Anterior view. B. Posterior view. C. Apical part of coxa and proximal part of telopodite, anterior view. D. (Anterior-) mesal view. E. Posterior distal lobe of telomere; insertion highlights spine-like process. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mla = metaplical lamella; msp = metaplical spine-like process; ptl = posterior distal" figureDoi="http://doi.org/10.5281/zenodo.3832780" httpUri="https://zenodo.org/record/3832780/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
A–B),
</paragraph>
<paragraph id="8921243D753FCB1768A6C15DFB2446CA" blockId="11.[225,1261,1752,1920]" box="[225,1261,1893,1920]" pageId="11" pageNumber="12">
- posterior distal lobe of telomere (
<emphasis id="BBEAF82F753FCB176A3DC15EFD504635" box="[634,665,1893,1919]" italics="true" pageId="11" pageNumber="12">ptl</emphasis>
) apically with a dark, spine-like process (
<figureCitation id="11A538B8753FCB176CC3C15EFB1646CA" box="[1156,1247,1893,1920]" captionStart="Fig" captionStartId="12.[227,270,1862,1888]" captionTargetBox="[272,1377,302,1825]" captionTargetId="figure@12.[272,1377,302,1825]" captionTargetPageId="12" captionText="Fig. 5. Geotypodon submontanus gen. et sp. nov., holotype. A–E. Left gonopod. A. Anterior view. B. Posterior view. C. Apical part of coxa and proximal part of telopodite, anterior view. D. (Anterior-) mesal view. E. Posterior distal lobe of telomere; insertion highlights spine-like process. F. Limbus. Abbreviations: atl = anterior distal lobe of telomere; itl = intermediate distal lamella of telomere; ll = longitudinal lamella; mla = metaplical lamella; msp = metaplical spine-like process; ptl = posterior distal" figureDoi="http://doi.org/10.5281/zenodo.3832780" httpUri="https://zenodo.org/record/3832780/files/figure.png" pageId="11" pageNumber="12">Fig. 5E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C18477B6753FCB1168A6C19DFB0740A8" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8921243D753FCB1768A6C19DFDDB468A" blockId="11.[225,530,1958,1985]" box="[225,530,1958,1985]" pageId="11" pageNumber="12">
<heading id="D2699351753FCB1768A6C19DFDDB468A" bold="true" box="[225,530,1958,1985]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="BBEAF82F753FCB1768A6C19DFDDB468A" bold="true" box="[225,530,1958,1985]" pageId="11" pageNumber="12">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8921243D753FCB1068A6C1EDFA40495B" blockId="11.[225,1270,2005,2032]" lastBlockId="12.[1305,1417,2038,2065]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Known only from West Kilombero FR. Altitude:
<quantity id="4E6689D8753FCB176B65C1EEFCB346BA" box="[802,890,2005,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.145" pageId="11" pageNumber="12" unit="m" value="1145.0">1145 m</quantity>
asl. Habitat: submontane forest. lobe
<quantity id="4E6689D87538CB106D5EC1CDFA4D495B" box="[1305,1412,2038,2065]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="mm" value="0.01">0.01 mm</quantity>
.
</paragraph>
<caption id="DDE174B57538CB1068A4C17DFA5446A7" ID-DOI="http://doi.org/10.5281/zenodo.3832780" ID-Zenodo-Dep="3832780" httpUri="https://zenodo.org/record/3832780/files/figure.png" pageId="12" pageNumber="13" startId="12.[227,270,1862,1888]" targetBox="[272,1377,302,1825]" targetPageId="12">
<paragraph id="8921243D7538CB1068A4C17DFA5446A7" blockId="12.[227,1437,1862,2030]" pageId="12" pageNumber="13">
<emphasis id="BBEAF82F7538CB1068A4C17DFEFD462B" bold="true" box="[227,308,1862,1889]" pageId="12" pageNumber="13">Fig. 5.</emphasis>
<taxonomicName id="4E9E5FBE7538CB106907C17DFDBC462A" authorityName="Enghoff" authorityYear="2016" box="[320,629,1862,1888]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="submontanus" status="gen. et sp. nov.">
<emphasis id="BBEAF82F7538CB106907C17DFDBC462A" box="[320,629,1862,1888]" italics="true" pageId="12" pageNumber="13">Geotypodon submontanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D945547538CB106AC6C17CFC89462B" box="[641,832,1863,1889]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="BBEAF82F7538CB106B8EC17DFBC9462A" bold="true" box="[969,1024,1862,1888]" pageId="12" pageNumber="13">A–E</emphasis>
. Left gonopod.
<emphasis id="BBEAF82F7538CB106C8EC17DFB17462A" bold="true" box="[1225,1246,1862,1888]" pageId="12" pageNumber="13">A</emphasis>
. Anterior view.
<emphasis id="BBEAF82F7538CB1068A4C151FF3E46CE" bold="true" box="[227,247,1898,1924]" pageId="12" pageNumber="13">B</emphasis>
. Posterior view.
<emphasis id="BBEAF82F7538CB1069F8C152FE1D46C9" bold="true" box="[447,468,1897,1923]" pageId="12" pageNumber="13">C</emphasis>
. Apical part of coxa and proximal part of telopodite, anterior view.
<emphasis id="BBEAF82F7538CB106CB0C151FAC546CE" bold="true" box="[1271,1292,1898,1924]" pageId="12" pageNumber="13">D</emphasis>
. (Anterior-) mesal view.
<emphasis id="BBEAF82F7538CB10693BC1B6FE5946ED" bold="true" box="[380,400,1933,1959]" pageId="12" pageNumber="13">E</emphasis>
. Posterior distal lobe of telomere; insertion highlights spine-like process.
<emphasis id="BBEAF82F7538CB106D50C1B6FAE046ED" bold="true" box="[1303,1321,1933,1959]" pageId="12" pageNumber="13">F</emphasis>
. Limbus. Abbreviations:
<emphasis id="BBEAF82F7538CB1069DDC18BFE704680" box="[410,441,1968,1994]" italics="true" pageId="12" pageNumber="13">atl</emphasis>
= anterior distal lobe of telomere;
<emphasis id="BBEAF82F7538CB106B1BC18BFCBD4680" box="[860,884,1968,1994]" italics="true" pageId="12" pageNumber="13">itl</emphasis>
= intermediate distal lamella of telomere;
<emphasis id="BBEAF82F7538CB106D34C18BFA4A4680" box="[1395,1411,1968,1994]" italics="true" pageId="12" pageNumber="13">ll</emphasis>
= longitudinal lamella;
<emphasis id="BBEAF82F7538CB10699CC1E8FDCE46A7" box="[475,519,2003,2029]" italics="true" pageId="12" pageNumber="13">mla</emphasis>
= metaplical lamella;
<emphasis id="BBEAF82F7538CB106B4FC1EFFCFE46A7" box="[776,823,2004,2029]" italics="true" pageId="12" pageNumber="13">msp</emphasis>
= metaplical spine-like process;
<emphasis id="BBEAF82F7538CB106CF4C1E8FB1B46A7" box="[1203,1234,2003,2029]" italics="true" pageId="12" pageNumber="13">ptl</emphasis>
= posterior distal
</paragraph>
</caption>
<paragraph id="8921243D7538CB10695AC1CDFADB495B" blockId="12.[285,1298,2038,2065]" box="[285,1298,2038,2065]" pageId="12" pageNumber="13">
of telomere;
<emphasis id="BBEAF82F7538CB1069F5C1CCFE1D495A" box="[434,468,2039,2064]" italics="true" pageId="12" pageNumber="13">pts</emphasis>
= proximal telomeral spine. Scales: A–D =
<quantity id="4E6689D87538CB106B9EC1CDFBFF495B" box="[985,1078,2038,2065]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="mm" value="0.1">0.1 mm</quantity>
, E =
<quantity id="4E6689D87538CB106C35C1CDFB17495B" box="[1138,1246,2038,2065]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="0.05">0.05 mm</quantity>
, F =
</paragraph>
<paragraph id="8921243D7539CB1168A6C714FE0A4003" blockId="13.[225,451,303,329]" box="[225,451,303,329]" pageId="13" pageNumber="14">
<heading id="D26993517539CB1168A6C714FE0A4003" bold="true" box="[225,451,303,329]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="BBEAF82F7539CB1168A6C714FE0A4003" bold="true" box="[225,451,303,329]" pageId="13" pageNumber="14">Coexisting species</emphasis>
</heading>
</paragraph>
<paragraph id="8921243D7539CB1168A6C765FB0740A8" blockId="13.[225,1436,350,482]" pageId="13" pageNumber="14">
No other odontopygids were found in the same sample as
<taxonomicName id="4E9E5FBE7539CB116B8CC765FB584032" authorityName="Enghoff" authorityYear="2016" box="[971,1169,350,376]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="submontanus" status="gen. et sp. nov.">
<emphasis id="BBEAF82F7539CB116B8CC765FB584032" box="[971,1169,350,376]" italics="true" pageId="13" pageNumber="14">G. submontanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D945547539CB116CD9C765FAAD4032" box="[1182,1380,350,376]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, but
<taxonomicName id="4E9E5FBE7539CB1168A6C7B9FE4F40D1" box="[225,390,385,412]" class="Diplopoda" family="Odontopygidae" genus="Geotypodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="millimanus" status="gen. et sp. nov.">
<emphasis id="BBEAF82F7539CB1168A6C7B9FE4F40D1" box="[225,390,385,412]" italics="true" pageId="13" pageNumber="14">G. millimanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A0D945547539CB1169CAC7B9FDF440D6" box="[397,573,386,412]" pageId="13" pageNumber="14" rank="species">gen. et sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4E9E5FBE7539CB116A0FC7BAFBC440D1" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[584,1037,385,411]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="basiliscus">
<emphasis id="BBEAF82F7539CB116A0FC7BAFC9540D1" box="[584,860,385,411]" italics="true" pageId="13" pageNumber="14">Chaleponcus basiliscus</emphasis>
<bibRefCitation id="ED0F59CC7539CB116B25C7BAFBC440D1" author="Enghoff H." box="[866,1037,385,411]" pageId="13" pageNumber="14" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4E9E5FBE7539CB116C5FC7B9FA5140D1" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[1048,1432,385,412]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="circumvallatus">
<emphasis id="BBEAF82F7539CB116C5FC7B9FB2F40D1" box="[1048,1254,385,412]" italics="true" pageId="13" pageNumber="14">C. circumvallatus</emphasis>
<bibRefCitation id="ED0F59CC7539CB116CAAC7BAFA5140D1" author="Enghoff H." box="[1261,1432,385,411]" pageId="13" pageNumber="14" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4E9E5FBE7539CB1168A6C79EFEA340F4" authorityName="Enghoff" authorityYear="2014" box="[225,362,420,447]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gracilior">
<emphasis id="BBEAF82F7539CB1168A6C79EFEA340F4" box="[225,362,420,447]" italics="true" pageId="13" pageNumber="14">C. gracilior</emphasis>
</taxonomicName>
,
<taxonomicName id="4E9E5FBE7539CB116930C79EFDBA40F4" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[375,627,420,447]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ibis">
<emphasis id="BBEAF82F7539CB116930C79EFE0B40F4" box="[375,450,420,447]" italics="true" pageId="13" pageNumber="14">C. ibis</emphasis>
<bibRefCitation id="ED0F59CC7539CB11698FC79FFDBA40F4" author="Enghoff H." box="[456,627,420,446]" pageId="13" pageNumber="14" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4E9E5FBE7539CB116A3AC79EFC4540F4" authority="Enghoff, 2014" authorityName="Enghoff" authorityYear="2014" box="[637,908,420,447]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="netus">
<emphasis id="BBEAF82F7539CB116A3AC79EFD1540F4" box="[637,732,421,447]" italics="true" pageId="13" pageNumber="14">C. netus</emphasis>
<bibRefCitation id="ED0F59CC7539CB116AA6C79FFC4540F4" author="Enghoff H." box="[737,908,420,446]" pageId="13" pageNumber="14" pagination="1 - 75" refId="ref7990" refString="Enghoff H. 2014. A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 100: 1 - 75. http: // dx. doi. org / 10.5852 / ejt. 2014.100" type="journal article" year="2014">Enghoff, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4E9E5FBE7539CB116BD1C79EFB6C40F5" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" box="[918,1189,420,447]" class="Diplopoda" family="Odontopygidae" genus="Chaleponcus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tintin">
<emphasis id="BBEAF82F7539CB116BD1C79EFC3C40F4" box="[918,1013,421,447]" italics="true" pageId="13" pageNumber="14">C. tintin</emphasis>
<bibRefCitation id="ED0F59CC7539CB116BBDC79FFB6C40F5" author="Enghoff H. &amp; Frederiksen S. B." box="[1018,1189,420,447]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref8050" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4E9E5FBE7539CB116CE8C79EFE4340A8" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipala">
<emphasis id="BBEAF82F7539CB116CE8C79EFA5540F4" box="[1199,1436,420,447]" italics="true" pageId="13" pageNumber="14">Aquattuor longipala</emphasis>
<bibRefCitation id="ED0F59CC7539CB1168A6C7FCFE4340A8" author="Enghoff H. &amp; Frederiksen S. B." box="[225,394,455,482]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref8050" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4E9E5FBE7539CB116984C7F3FCFE40A8" authority="Enghoff, 2015" authorityName="Enghoff" authorityYear="2015" box="[451,823,455,482]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="udzungwensis">
<emphasis id="BBEAF82F7539CB116984C7F3FD4E40AB" box="[451,647,455,482]" italics="true" pageId="13" pageNumber="14">A. udzungwensis</emphasis>
<bibRefCitation id="ED0F59CC7539CB116AC9C7FCFCFE40A8" author="Enghoff H. &amp; Frederiksen S. B." box="[654,823,455,482]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref8050" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" type="journal article" year="2015">Enghoff, 2015</bibRefCitation>
</taxonomicName>
also occur in West Kilombero FR.
</paragraph>
</subSubSection>
</treatment>
</document>
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