treatments-xml/data/F8/75/1A/F8751A58FFE5FF93FF1DFDA1FC48EFF3.xml

<document id="4A3767BD82CD1478C4BDF692258CDA00" ID-CLB-Dataset="24960" ID-DOI="10.11646/zootaxa.4695.1.2" ID-GBIF-Dataset="929605d5-af60-4733-b17d-6379d750ae04" ID-ISSN="1175-5326" ID-Zenodo-Dep="3530087" ID-ZooBank="3007F762-721A-477B-94C9-AC0A24E2ACB8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573024085816" checkinUser="plazi" docAuthor="Kim, Hyun Kyong, Chan, Benny K. K., Corbari, Laure, Rodriguez Moreno, Paula A., Achituv, Yair &amp; Kim, Won" docDate="2019" docId="F8751A58FFE5FF93FF1DFDA1FC48EFF3" docLanguage="en" docName="zootaxa.4695.1.2.pdf" docOrigin="Zootaxa 4695 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyrgoma spurtruncata Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim 2019, sp. nov." docType="treatment" docUuid="9EA455E4-3168-4274-8B41-66AA11643C35" docUuidSource="ZooBank" docVersion="6" lastPageNumber="44" masterDocId="044C6220FFE1FF9DFF8AFFA3FFD8ED2A" masterDocTitle="A new species of the coral associated barnacle (Thoracica: Pyrgomatidae: Pyrgoma) from a deep-sea oculinid coral in New Caledonian waters" masterLastPageNumber="44" masterPageNumber="26" pageNumber="26" updateTime="1698763510647" updateUser="ExternalLinkService">
<mods:mods id="872667AF29D9DB8D658796F27ECBA6D8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B1BF1F573F3A5D6C0060EE8DFA962A1">
<mods:title id="F4752CA5F89CB6C41CFE58FDBC0DF2FF">A new species of the coral associated barnacle (Thoracica: Pyrgomatidae: Pyrgoma) from a deep-sea oculinid coral in New Caledonian waters</mods:title>
</mods:titleInfo>
<mods:name id="3FC9C30620EC38F258CC643B9FAE6D01" type="personal">
<mods:role id="4DD7ACE35AC2ED0EB761026082BD980B">
<mods:roleTerm id="896269932F954BCF3E21B7EB8C315965">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B42BAC03C6C735A5768AAE6C1E54EB1C">Kim, Hyun Kyong</mods:namePart>
</mods:name>
<mods:name id="33B90D58AEAF3CF521F1E5CD513F974B" type="personal">
<mods:role id="6FCBBBFB42BA921573C9E582EE710EFE">
<mods:roleTerm id="757D451FA85A9A5EDD259ADA97A65A3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B898714279845FA517DC5DC0299CCEA">Chan, Benny K. K.</mods:namePart>
</mods:name>
<mods:name id="553D92F9978E4A4E31E552611A6962B4" type="personal">
<mods:role id="BAF4C4A2B21B8125B9393F6566AA61F1">
<mods:roleTerm id="903D9FBA6585CF02D8EC42BA4F752C53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8F91183A5FD1D27F3822B8FB501B1FE">Corbari, Laure</mods:namePart>
</mods:name>
<mods:name id="201B8EB7D4162653C6E72283D8801200" type="personal">
<mods:role id="0BD5256488B30F3B261B4A2EC35E918C">
<mods:roleTerm id="BC5202D07A070632D6F5EF5A311F3956">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB5D1DC438936F9A05EE6C587F6B16F3">Rodriguez Moreno, Paula A.</mods:namePart>
</mods:name>
<mods:name id="F484ADB6B652770847341F234F115625" type="personal">
<mods:role id="4699D7C3E576372E097F564950072461">
<mods:roleTerm id="C0A582743F610D01B45EE4C929BD9718">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A5B091ACAFCB14AC68F9214C903B9B71">Achituv, Yair</mods:namePart>
</mods:name>
<mods:name id="A3EEE446333641398184718A5DE64C88" type="personal">
<mods:role id="E52E4D4A731FFA1278D226A3F8E7EB6D">
<mods:roleTerm id="BFAE6AAACD93ABC2D8D75BC5628E1A64">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="979BABFF811743862BF9BC780E24EB2C">Kim, Won</mods:namePart>
</mods:name>
<mods:typeOfResource id="72EB13B729091FA3B283944A0414C649">text</mods:typeOfResource>
<mods:relatedItem id="9A613D9C51E1989CCAA04DD2E5562F9A" type="host">
<mods:titleInfo id="63D3FDBCB005E9A68EA593301A80F2B9">
<mods:title id="EB127B046F14E9002A9A968BD8F54FE4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1009012222CEB315CF4BA530B0055879">
<mods:date id="89F3E5881E0C2580F6470C4DFF09C842">2019</mods:date>
<mods:detail id="40BEC38E5771C95DDF95827F0CBA4F0F" type="pubDate">
<mods:number id="66EC04168EB29683CF1ABD91775C6C6C">2019-11-06</mods:number>
</mods:detail>
<mods:detail id="FFA61D4818AFCB8E94DD98355C80B567" type="volume">
<mods:number id="95B9371C93BCF0D758B22F9FE65E9AB0">4695</mods:number>
</mods:detail>
<mods:detail id="E9B9E9EBC8CCFEC076FCAD2EE4258B99" type="issue">
<mods:number id="46819C39296089EF7983049DB1BAA0A9">1</mods:number>
</mods:detail>
<mods:extent id="FCCE9D57AD66CA63DC9CCB74FAB4CED8" unit="page">
<mods:start id="3FB27AB24F7D4C7D5E177294D3A602A2">26</mods:start>
<mods:end id="3D0735DE09301D6908AA972F3BF96883">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D63DF1A6FC344E770BE3D48259E39694">journal article</mods:classification>
<mods:identifier id="CF1364C568AABD5FB5C87C16A9414153" type="CLB-Dataset">24960</mods:identifier>
<mods:identifier id="A120E1092BFA297890ACDE6FE56EA3CD" type="DOI">10.11646/zootaxa.4695.1.2</mods:identifier>
<mods:identifier id="644105FCC6C9C05CBCEA5A62AD44C9B9" type="GBIF-Dataset">929605d5-af60-4733-b17d-6379d750ae04</mods:identifier>
<mods:identifier id="65424663541876891B883D81BFEE783B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="156940206E95B2E6C3F59C4B38468859" type="Zenodo-Dep">3530087</mods:identifier>
<mods:identifier id="F6A88652BCA50E548C66AD1B5BCA245B" type="ZooBank">3007F762-721A-477B-94C9-AC0A24E2ACB8</mods:identifier>
</mods:mods>
<treatment id="F8751A58FFE5FF93FF1DFDA1FC48EFF3" ID-GBIF-Taxon="161761773" LSID="urn:lsid:zoobank.org:act:9EA455E4-3168-4274-8B41-66AA11643C35" httpUri="http://treatment.plazi.org/id/F8751A58FFE5FF93FF1DFDA1FC48EFF3" lastPageId="14" pageId="4">
<subSubSection id="38C6F8C5FFE5FF99FF1DFDA1FE24EF36" box="[151,508,514,540]" pageId="4" type="nomenclature">
<paragraph id="7063AB4EFFE5FF99FF1DFDA1FE24EF36" blockId="4.[151,508,514,575]" box="[151,508,514,540]" pageId="4">
<heading id="2B2B1C22FFE5FF99FF1DFDA1FE24EF36" bold="true" box="[151,508,514,540]" fontSize="11" level="1" pageId="4" reason="1">
<emphasis id="42A8775CFFE5FF99FF1DFDA1FE24EF36" bold="true" box="[151,508,514,540]" pageId="4">
<taxonomicName id="B7DCD0CDFFE5FF99FF1DFDA1FE43EF36" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[151,411,514,540]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="4" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE5FF99FF1DFDA1FE43EF36" bold="true" box="[151,411,514,540]" italics="true" pageId="4">Pyrgoma spurtruncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFE5FF99FE29FDA1FE24EF36" box="[419,508,514,540]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="38C6F8C5FFE5FF99FF1DFD86FEC3EF15" box="[151,283,549,575]" pageId="4" type="description">
<paragraph id="7063AB4EFFE5FF99FF1DFD86FEC3EF15" blockId="4.[151,508,514,575]" box="[151,283,549,575]" pageId="4">
(
<figureCitation id="E8E7B7CBFFE5FF99FF15FD86FECBEF15" box="[159,275,549,575]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStartId-0="4.[151,250,1975,2000]" captionStartId-1="5.[151,250,1945,1970]" captionStartId-2="6.[151,250,1459,1484]" captionStartId-3="7.[151,250,1910,1935]" captionStartId-4="8.[151,250,1516,1541]" captionStartId-5="10.[151,250,1832,1857]" captionStartId-6="11.[151,250,1911,1936]" captionStartId-7="12.[151,250,1946,1971]" captionStartId-8="13.[151,250,1832,1857]" captionTargetBox-0="[189,1398,597,1949]" captionTargetBox-1="[344,1244,181,1920]" captionTargetBox-2="[164,1423,181,1434]" captionTargetBox-3="[191,1395,182,1884]" captionTargetBox-4="[151,1436,181,1491]" captionTargetBox-5="[280,1308,237,1808]" captionTargetBox-6="[177,1411,181,1885]" captionTargetBox-7="[157,1431,181,1920]" captionTargetBox-8="[177,1411,525,1807]" captionTargetId-0="figure@4.[189,1398,597,1949]" captionTargetId-1="figure@5.[344,1244,181,1920]" captionTargetId-2="figure@6.[164,1423,181,1434]" captionTargetId-3="figure@7.[191,1396,181,1885]" captionTargetId-4="figure@8.[151,1436,181,1491]" captionTargetId-5="figure@10.[280,1308,237,1808]" captionTargetId-6="figure@11.[177,1411,181,1885]" captionTargetId-7="figure@12.[157,1431,181,1920]" captionTargetId-8="figure@13.[177,1411,525,1807]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionText-0="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." captionText-1="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." captionText-2="FIGURE 4. Base view of Pyrgoma spurtuncata sp. nov.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A, top view of base without longitudinal ladder-like perforations; B, top view of base with short sections of ladder-like perforations; C, D, top view of base with some ladder-like perforations; E, F, enlarged view of basis; G, H, enlarged view of basis; I, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm" captionText-3="FIGURE 5. Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-5597): A, Light microscopy view showing the shell and basis together; B, Scanning Electron Microcopy (SEM) of shell and basis, showing the longitudinal ladder-like perforations; C−F, SEM views showing side view (C,D) and top view (E, F) of basis and the ladder-like longitudinal perforations. Scale bars in mm." captionText-4="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." captionText-5="FIGURE 7. Mouthparts of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, maxilla; B, mandibular palp; C, maxillule; D, mandible; E, lower margin and inferior angle of mandible; F, inferior margin of the mandible; G, labrum; H, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm." captionText-6="FIGURE 8. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus I; B, C, distal serrulate setae on posterior and anterior rami of cirrus I; D, cirrus II; E, distal serrulate setae on posterior and anterior rami of cirrus II; F, cirrus III; G. magnified view of cirrus II; H, I, distal serrulate setae on posterior and anterior rami of cirrus III. Scale bars: A−I = 100µm." captionText-7="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." captionText-8="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.3530091" figureDoi-1="http://doi.org/10.5281/zenodo.3530093" figureDoi-2="http://doi.org/10.5281/zenodo.3530095" figureDoi-3="http://doi.org/10.5281/zenodo.3530097" figureDoi-4="http://doi.org/10.5281/zenodo.3530099" figureDoi-5="http://doi.org/10.5281/zenodo.3530101" figureDoi-6="http://doi.org/10.5281/zenodo.3530103" figureDoi-7="http://doi.org/10.5281/zenodo.3530105" figureDoi-8="http://doi.org/10.5281/zenodo.3530107" httpUri-0="https://zenodo.org/record/3530091/files/figure.png" httpUri-1="https://zenodo.org/record/3530093/files/figure.png" httpUri-2="https://zenodo.org/record/3530095/files/figure.png" httpUri-3="https://zenodo.org/record/3530097/files/figure.png" httpUri-4="https://zenodo.org/record/3530099/files/figure.png" httpUri-5="https://zenodo.org/record/3530101/files/figure.png" httpUri-6="https://zenodo.org/record/3530103/files/figure.png" httpUri-7="https://zenodo.org/record/3530105/files/figure.png" httpUri-8="https://zenodo.org/record/3530107/files/figure.png" pageId="4">Figs 2−10</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="24A3FBC6FFE5FF99FF1DF814FA9DEADE" ID-DOI="http://doi.org/10.5281/zenodo.3530091" ID-Zenodo-Dep="3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="4" startId="4.[151,250,1975,2000]" targetBox="[189,1398,597,1949]" targetPageId="4">
<paragraph id="7063AB4EFFE5FF99FF1DF814FA9DEADE" blockId="4.[151,1436,1975,2036]" pageId="4">
<emphasis id="42A8775CFFE5FF99FF1DF814FDB6EAFA" bold="true" box="[151,622,1975,2000]" pageId="4">
FIGURE 2. 
<taxonomicName id="B7DCD0CDFFE5FF99FE92F81BFE21EAE5" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[280,505,1976,1999]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="4" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFE5FF99FE92F81BFE21EAE5" bold="true" box="[280,505,1976,1999]" italics="true" pageId="4">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFE5FF99FE75F81BFD8BEAFA" box="[511,595,1976,2000]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
: A
</emphasis>
(upper-lateral view), 
<emphasis id="42A8775CFFE5FF99FCC3F81BFC82EAFA" bold="true" box="[841,858,1976,2000]" pageId="4">B</emphasis>
(lateral view) MNHN-IU-2014-23572 (holotype); 
<emphasis id="42A8775CFFE5FF99FAD1F814FAB5EAFA" bold="true" box="[1371,1389,1975,2000]" pageId="4">C</emphasis>
(top view), 
<emphasis id="42A8775CFFE5FF99FF54F87FFF28EADE" bold="true" box="[222,240,2012,2036]" pageId="4">D</emphasis>
(lateral view) (paratype) MNHN-IU-2014-5597; 
<emphasis id="42A8775CFFE5FF99FD6DF87FFD20EADE" bold="true" box="[743,760,2012,2036]" pageId="4">E</emphasis>
, 
<emphasis id="42A8775CFFE5FF99FC8EF87FFCCBEADE" bold="true" box="[772,787,2012,2036]" pageId="4">F</emphasis>
(upper lateral view) (paratype) MNHN-IU-2014-5597.
</paragraph>
</caption>
<caption id="24A3FBC6FFE4FF98FF1DF83AFE26EAD0" ID-DOI="http://doi.org/10.5281/zenodo.3530093" ID-Zenodo-Dep="3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="5" startId="5.[151,250,1945,1970]" targetBox="[344,1244,181,1920]" targetPageId="5">
<paragraph id="7063AB4EFFE4FF98FF1DF83AFE26EAD0" blockId="5.[151,1437,1945,2042]" pageId="5">
<emphasis id="42A8775CFFE4FF98FF1DF83AFDB4EA98" bold="true" box="[151,620,1945,1970]" pageId="5">
FIGURE 3. 
<taxonomicName id="B7DCD0CDFFE4FF98FE9DF839FE2FEA9B" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[279,503,1946,1969]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="5" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFE4FF98FE9DF839FE2FEA9B" bold="true" box="[279,503,1946,1969]" italics="true" pageId="5">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFE4FF98FE77F839FD88EA98" box="[509,592,1946,1970]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
: A
</emphasis>
, external and internal view of shell (MNHN-IU-2014-23572: holotype); 
<emphasis id="42A8775CFFE4FF98FACFF839FA8EEA98" bold="true" box="[1349,1366,1946,1970]" pageId="5">B</emphasis>
, external and internal view of shell (MNHN-IU-2014-23573, paratype); 
<emphasis id="42A8775CFFE4FF98FCBAF81EFC9AEAFC" bold="true" box="[816,834,1981,2006]" pageId="5">C</emphasis>
, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm.
</paragraph>
</caption>
<caption id="24A3FBC6FFE7FF9BFF1DFA10FE85EBAA" ID-DOI="http://doi.org/10.5281/zenodo.3530095" ID-Zenodo-Dep="3530095" httpUri="https://zenodo.org/record/3530095/files/figure.png" pageId="6" startId="6.[151,250,1459,1484]" targetBox="[164,1423,181,1434]" targetPageId="6">
<paragraph id="7063AB4EFFE7FF9BFF1DFA10FE85EBAA" blockId="6.[151,1437,1459,1664]" pageId="6">
<emphasis id="42A8775CFFE7FF9BFF1DFA10FF71EB3E" bold="true" pageId="6">
FIGURE 4. Base view of 
<taxonomicName id="B7DCD0CDFFE7FF9BFE23FA17FD54E8E1" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[425,652,1460,1483]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="6" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFE7FF9BFE23FA17FD54E8E1" bold="true" box="[425,652,1460,1483]" italics="true" pageId="6">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFE7FF9BFD19FA17FD39E8E6" box="[659,737,1460,1484]" pageId="6" rank="species">sp. nov</taxonomicNameLabel>
.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A
</emphasis>
, top view of base without longitudinal ladder-like perforations; 
<emphasis id="42A8775CFFE7FF9BFCBAFA5FFC99EB3E" bold="true" box="[816,833,1532,1556]" pageId="6">B</emphasis>
, top view of base with short sections of ladder-like perforations; 
<emphasis id="42A8775CFFE7FF9BFF5CF9BCFF30EB12" bold="true" box="[214,232,1567,1592]" pageId="6">C</emphasis>
, 
<emphasis id="42A8775CFFE7FF9BFF7FF983FEDFEB12" bold="true" box="[245,263,1568,1592]" pageId="6">D</emphasis>
, top view of base with some ladder-like perforations; 
<emphasis id="42A8775CFFE7FF9BFCA4F983FCE7EB12" bold="true" box="[814,831,1568,1592]" pageId="6">E</emphasis>
, 
<emphasis id="42A8775CFFE7FF9BFCC6F983FC83EB12" bold="true" box="[844,859,1568,1592]" pageId="6">F</emphasis>
, enlarged view of basis; 
<emphasis id="42A8775CFFE7FF9BFBD0F9BCFBB6EB12" bold="true" box="[1114,1134,1567,1592]" pageId="6">G</emphasis>
, 
<emphasis id="42A8775CFFE7FF9BFBF0F983FB56EB12" bold="true" box="[1146,1166,1568,1592]" pageId="6">H</emphasis>
, enlarged view of basis; 
<emphasis id="42A8775CFFE7FF9BFA06F983FA4EEB12" bold="true" box="[1420,1430,1568,1592]" pageId="6">I</emphasis>
, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm
</paragraph>
</caption>
<subSubSection id="38C6F8C5FFE7FF9BFF1DF90BFC23EAC8" pageId="6" type="materials_examined">
<paragraph id="7063AB4EFFE7FF9BFF1DF90BFAA6EBCC" blockId="6.[151,1437,1703,2018]" pageId="6">
<emphasis id="42A8775CFFE7FF9BFF1DF90BFEE5EBE8" bold="true" box="[151,317,1703,1730]" pageId="6">Type material.</emphasis>
<materialsCitation id="C0B4A113FFE7FF9BFEC8F904FAA6EBCC" ID-GBIF-Occurrence="2550868461" collectingDate="1987-05-22" collectionCode="SMIB" collectorName="Stn DW" country="New Caledonia" latitude="-23.675" location="Norfolk Ridge" longLatPrecision="21" longitude="167.96167" municipality="Jumeau Ouest" pageId="6" specimenCode="SMIB 3" specimenCount="1" typeStatus="holotype">
<typeStatus id="AF6715ECFFE7FF9BFEC8F904FE72EBEB" box="[322,426,1703,1729]" pageId="6" type="holotype">Holotype</typeStatus>
: MNHN-IU-2014-23572, 
<collectingCountry id="08CBEBDEFFE7FF9BFD49F90BFCB3EBEB" box="[707,875,1703,1730]" name="New Caledonia" pageId="6">New Caledonia</collectingCountry>
, 
<location id="7503FD95FFE7FF9BFCFFF904FBCAEBEB" LSID="urn:lsid:plazi:treatment:F8751A58FFE5FF93FF1DFDA1FC48EFF3:7503FD95FFE7FF9BFCFFF904FBCAEBEB" box="[885,1042,1703,1729]" country="New Caledonia" latitude="-23.675" longLatPrecision="21" longitude="167.96167" municipality="Jumeau Ouest" name="Norfolk Ridge" pageId="6">Norfolk Ridge</location>
, 
<collectingMunicipality id="90073134FFE7FF9BFB96F90BFB6BEBE8" box="[1052,1203,1704,1730]" pageId="6">Jumeau Ouest</collectingMunicipality>
seamount, 
<collectedFrom id="C3C0E8B3FFE7FF9BFAA1F90BFE17EBCC" pageId="6">
on 
<taxonomicName id="B7DCD0CDFFE7FF9BFAC6F904FE17EBCC" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Anthozoa" family="Oculinidae" genus="Madrepora" kingdom="Animalia" order="Scleractinia" pageId="6" phylum="Cnidaria" rank="species" species="oculata">
<emphasis id="42A8775CFFE7FF9BFAC6F904FEF9EBCF" italics="true" pageId="6">Madrepora oculata</emphasis>
<bibRefCitation id="144DD6BFFFE7FF9BFEADF968FE17EBCC" author="Linnaeus, C." box="[295,463,1739,1766]" pageId="6" refId="ref8359" refString="Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Vol. 1. 10 th Revised Edition. Laurentius Salvius, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
</collectedFrom>
; 
<specimenCode id="207A0335FFE7FF9BFE53F96FFDF7EBCC" box="[473,559,1740,1766]" collectionCode="SMIB" pageId="6">SMIB 3</specimenCode>
cruise (
<collectorName id="DD29CE98FFE7FF9BFD09F96FFD38EBCC" box="[643,736,1740,1766]" pageId="6">Stn DW</collectorName>
14, 
<geoCoordinate id="15E8CD89FFE7FF9BFC8EF96FFC4FEBCC" box="[772,919,1740,1766]" degrees="23" direction="south" minutes="40" orientation="latitude" pageId="6" precision="15" seconds="30" value="-23.675">23° 40’ 30’’ S</geoCoordinate>
, 
<geoCoordinate id="15E8CD89FFE7FF9BFC28F968FB99EBCC" box="[930,1089,1739,1766]" degrees="167" direction="east" minutes="57" orientation="longitude" pageId="6" precision="15" seconds="42" value="167.96167">167° 57’42’’ E</geoCoordinate>
, 
<date id="04628D8EFFE7FF9BFBC6F96FFB05EBCC" box="[1100,1245,1740,1766]" pageId="6" value="1987-05-22">
<collectingDate id="14267466FFE7FF9BFBC6F96FFB05EBCC" box="[1100,1245,1740,1766]" pageId="6" value="1987-05-22">
22 May 
<quantity id="B72406ABFFE7FF9BFB2FF96FFB05EBCC" box="[1189,1245,1740,1766]" metricMagnitude="6" metricUnit="m" metricValue="1.9872459999999996" pageId="6" unit="m" value="1987246.0">1987</quantity>
</collectingDate>
</date>
, 246 m depth).
</materialsCitation>
</paragraph>
<paragraph id="7063AB4EFFE7FF9BFF4DF953FC5CEA23" blockId="6.[151,1437,1703,2018]" box="[199,900,1775,1802]" pageId="6">
<typeStatus id="AF6715ECFFE7FF9BFF4DF953FEE0EA20" box="[199,312,1776,1802]" pageId="6" type="paratype">Paratypes</typeStatus>
: MNHN-IU-2014-23573, data same as for holotype.
</paragraph>
<paragraph id="7063AB4EFFE7FF9BFF4DF8B0FB3EEA78" blockId="6.[151,1437,1703,2018]" pageId="6">
MNHN-IU-2014-23574, 
<collectingCountry id="08CBEBDEFFE7FF9BFE6EF8B7FD4DEA07" box="[484,661,1811,1838]" name="New Caledonia" pageId="6">New Caledonia</collectingCountry>
, Norfolk Ridge, Banc Kaimon, Maru seamount, on 
<taxonomicName id="B7DCD0CDFFE7FF9BFB6EF8B0FF1BEA7B" authorityName="Linnaeus" authorityYear="1758" class="Anthozoa" family="Oculinidae" genus="Madrepora" kingdom="Animalia" order="Scleractinia" pageId="6" phylum="Cnidaria" rank="species" species="oculata">
<emphasis id="42A8775CFFE7FF9BFB6EF8B0FF1BEA7B" italics="true" pageId="6">Madrepora oculata</emphasis>
</taxonomicName>
; 
<collectionCode id="16CD338BFFE7FF9BFF58F89BFECEEA78" box="[210,278,1848,1874]" pageId="6">SMIB</collectionCode>
4 cruise (Stn DW41, 
<geoCoordinate id="15E8CD89FFE7FF9BFD80F89BFD7DEA78" box="[522,677,1848,1874]" degrees="24" direction="south" minutes="44" orientation="latitude" pageId="6" precision="15" seconds="18" value="-24.738333">24° 44’ 18’’ S</geoCoordinate>
, 
<geoCoordinate id="15E8CD89FFE7FF9BFD38F894FC86EA78" box="[690,862,1847,1874]" degrees="168" direction="east" minutes="08" orientation="longitude" pageId="6" precision="15" seconds="42" value="168.145">168° 08’ 42’’ E</geoCoordinate>
; 
<date id="04628D8EFFE7FF9BFCE0F89BFBD2EA78" box="[874,1034,1847,1874]" pageId="6" value="1989-03-08">8 March 1989</date>
; 
<quantity id="B72406ABFFE7FF9BFB9FF894FB48EA78" box="[1045,1168,1847,1874]" metricMagnitude="2" metricUnit="m" metricValue="2.325" metricValueMax="2.35" metricValueMin="2.3" pageId="6" unit="m" value="232.5" valueMax="235.0" valueMin="230.0">230-235 m</quantity>
depth).
</paragraph>
<paragraph id="7063AB4EFFE7FF9BFF4DF8F8FC23EAC8" blockId="6.[151,1437,1703,2018]" pageId="6">
MNHN-IU-2014-5596, MNHN-IU-2014-5597, MNHN-IU-2016-9936, 
<collectingCountry id="08CBEBDEFFE7FF9BFC78F8FFFB7BEA5F" box="[1010,1187,1883,1910]" name="New Caledonia" pageId="6">New Caledonia</collectingCountry>
, Norfolk Ridge, Banc Antigonia, on 
<taxonomicName id="B7DCD0CDFFE7FF9BFEB0F8DCFDCCEAB3" authorityName="Linnaeus" authorityYear="1758" box="[314,532,1919,1945]" class="Anthozoa" family="Oculinidae" genus="Madrepora" kingdom="Animalia" order="Scleractinia" pageId="6" phylum="Cnidaria" rank="species" species="oculata">
<emphasis id="42A8775CFFE7FF9BFEB0F8DCFDCCEAB3" box="[314,532,1919,1945]" italics="true" pageId="6">Madrepora oculata</emphasis>
</taxonomicName>
; 
<collectionCode id="16CD338BFFE7FF9BFDAEF823FDB0EAB0" box="[548,616,1920,1946]" pageId="6">SMIB</collectionCode>
4 cruise (Stn DW55, 
<geoCoordinate id="15E8CD89FFE7FF9BFCEBF823FC27EAB0" box="[865,1023,1920,1946]" degrees="23" direction="south" minutes="20" orientation="latitude" pageId="6" precision="15" seconds="48" value="-23.346668">23° 20’ 48’’ S</geoCoordinate>
, 
<geoCoordinate id="15E8CD89FFE7FF9BFB87F8DCFB75EAB0" box="[1037,1197,1919,1946]" degrees="168" direction="east" minutes="4" orientation="longitude" pageId="6" precision="15" seconds="48" value="168.08">168° 4’ 48’’ E</geoCoordinate>
; 
<date id="04628D8EFFE7FF9BFB30F823FA84EAB0" box="[1210,1372,1919,1946]" pageId="6" value="1985-03-09">9 March 1985</date>
; 
<quantity id="B72406ABFFE7FF9BFAE3F8DCFF05EA94" metricMagnitude="2" metricUnit="m" metricValue="2.375" metricValueMax="2.6" metricValueMin="2.15" pageId="6" unit="m" value="237.5" valueMax="260.0" valueMin="215.0">215- 260 m</quantity>
depth). MNHN-IU-2014-23575, 
<collectingCountry id="08CBEBDEFFE7FF9BFDDBF807FD27EA97" box="[593,767,1955,1982]" name="New Caledonia" pageId="6">New Caledonia</collectingCountry>
, Norfolk Ridge, on 
<taxonomicName id="B7DCD0CDFFE7FF9BFC53F800FB76EA97" authorityName="Linnaeus" authorityYear="1758" box="[985,1198,1955,1981]" class="Anthozoa" family="Oculinidae" genus="Madrepora" kingdom="Animalia" order="Scleractinia" pageId="6" phylum="Cnidaria" rank="species" species="oculata">Madrepora oculata</taxonomicName>
; BIOCAL cruise (Stn CP110, 
<geoCoordinate id="15E8CD89FFE7FF9BFF7AF86BFE53EAC8" box="[240,395,1992,2018]" degrees="22" direction="south" minutes="13" orientation="latitude" pageId="6" precision="15" seconds="19" value="-22.221945">22° 13’ 19’’ S</geoCoordinate>
, 
<geoCoordinate id="15E8CD89FFE7FF9BFE12F864FD9CEAC8" box="[408,580,1991,2018]" degrees="167" direction="east" minutes="09" orientation="longitude" pageId="6" precision="15" seconds="16" value="167.15443">167° 09’ 16’’ E</geoCoordinate>
; 
<date id="04628D8EFFE7FF9BFDDAF86BFCC7EAC8" box="[592,799,1991,2018]" pageId="6" value="1985-09-09">9 September 1985</date>
; 
<quantity id="B72406ABFFE7FF9BFCA0F864FC7DEAC8" box="[810,933,1991,2018]" metricMagnitude="2" metricUnit="m" metricValue="2.975" metricValueMax="3.2" metricValueMin="2.75" pageId="6" unit="m" value="297.5" valueMax="320.0" valueMin="275.0">275-320 m</quantity>
depth).
</paragraph>
</subSubSection>
<caption id="24A3FBC6FFE6FF9AFF1DF8D5FAA5EAFC" ID-DOI="http://doi.org/10.5281/zenodo.3530097" ID-Zenodo-Dep="3530097" httpUri="https://zenodo.org/record/3530097/files/figure.png" pageId="7" startId="7.[151,250,1910,1935]" targetBox="[191,1395,182,1884]" targetPageId="7">
<paragraph id="7063AB4EFFE6FF9AFF1DF8D5FAA5EAFC" blockId="7.[151,1436,1910,2007]" pageId="7">
<emphasis id="42A8775CFFE6FF9AFF1DF8D5FCADEAA5" bold="true" box="[151,885,1910,1935]" pageId="7">
FIGURE 5. 
<taxonomicName id="B7DCD0CDFFE6FF9AFE9DF8D4FE20EAA4" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[279,504,1911,1934]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="7" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFE6FF9AFE9DF8D4FE20EAA4" bold="true" box="[279,504,1911,1934]" italics="true" pageId="7">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFE6FF9AFE74F8D4FD95EAA5" box="[510,589,1911,1935]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
(MNHN-IU-2014-5597): A,
</emphasis>
Light microscopy view showing the shell and basis together; 
<emphasis id="42A8775CFFE6FF9AFF69F838FF2CEA99" bold="true" box="[227,244,1947,1971]" pageId="7">B</emphasis>
, Scanning Electron Microcopy (SEM) of shell and basis, showing the longitudinal ladder-like perforations; 
<emphasis id="42A8775CFFE6FF9AFAA6F839FA84EA99" bold="true" box="[1324,1372,1946,1971]" pageId="7">C −F</emphasis>
, SEM views showing side view (C,D) and top view (E, F) of basis and the ladder-like longitudinal perforations. Scale bars in mm.
</paragraph>
</caption>
<caption id="24A3FBC6FFE9FF95FF1DFA4FFB84EBBF" ID-DOI="http://doi.org/10.5281/zenodo.3530099" ID-Zenodo-Dep="3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="8" startId="8.[151,250,1516,1541]" targetBox="[151,1436,181,1491]" targetPageId="8">
<paragraph id="7063AB4EFFE9FF95FF1DFA4FFB84EBBF" blockId="8.[151,1436,1516,1685]" pageId="8">
<emphasis id="42A8775CFFE9FF95FF1DFA4FFB75EB2F" bold="true" box="[151,1197,1516,1541]" pageId="8">
FIGURE 6. Scanning electron micrograph of scuta and terga of 
<taxonomicName id="B7DCD0CDFFE9FF95FCD8FA4EFBECEB2E" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[850,1076,1517,1540]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="8" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFE9FF95FCD8FA4EFBECEB2E" bold="true" box="[850,1076,1517,1540]" italics="true" pageId="8">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFE9FF95FBB1FA4EFB49EB2F" box="[1083,1169,1517,1541]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
: A
</emphasis>
, external view of scuta and terga (MNHN-IU-2014-23572: holotype); 
<emphasis id="42A8775CFFE9FF95FDFCF9B2FD5FEB03" bold="true" box="[630,647,1553,1577]" pageId="8">B</emphasis>
, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); 
<emphasis id="42A8775CFFE9FF95FA0EF9B3FA4EEB03" bold="true" box="[1412,1430,1552,1577]" pageId="8">C</emphasis>
, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); 
<emphasis id="42A8775CFFE9FF95FC0DF996FC41EB67" bold="true" box="[903,921,1589,1613]" pageId="8">D</emphasis>
, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); 
<emphasis id="42A8775CFFE9FF95FE04F9FAFE47EB5B" bold="true" box="[398,415,1625,1649]" pageId="8">E</emphasis>
, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); 
<emphasis id="42A8775CFFE9FF95FB70F9FAFAD1EB5B" bold="true" box="[1274,1289,1625,1649]" pageId="8">F</emphasis>
, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm.
</paragraph>
</caption>
<subSubSection id="38C6F8C5FFE9FF95FF4DF960FCF6EA0F" pageId="8" type="diagnosis">
<paragraph id="7063AB4EFFE9FF95FF4DF960FCF6EA0F" blockId="8.[151,1437,1731,2046]" pageId="8">
<emphasis id="42A8775CFFE9FF95FF4DF960FE9AEBF4" bold="true" box="[199,322,1731,1758]" pageId="8">Diagnosis.</emphasis>
Scutum triangular, transversally elongated, occludent and basal margins longer than rostral margin. Adductor plate thick; rostral tooth blunt, wide. Tergum upper part triangular, long narrow spur with moderately broad external furrow; without crests for depressor muscle.
</paragraph>
</subSubSection>
<subSubSection id="38C6F8C5FFE9FF94FF4DF88CFDB7E87D" lastPageId="9" pageId="8" type="description">
<paragraph id="7063AB4EFFE9FF94FF4DF88CFEB8EC4D" blockId="8.[151,1437,1731,2046]" lastBlockId="9.[151,1437,153,2015]" lastPageId="9" pageId="8">
<emphasis id="42A8775CFFE9FF95FF4DF88CFE89EA60" bold="true" box="[199,337,1839,1866]" pageId="8">Description</emphasis>
(mainly based on 
<typeStatus id="AF6715ECFFE9FF95FD97F88CFD5AEA63" box="[541,642,1839,1865]" pageId="8" type="holotype">holotype</typeStatus>
, except where stated). Shell ovate (
<figureCitation id="E8E7B7CBFFE9FF95FB83F88CFB89EA60" box="[1033,1105,1839,1866]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="8">Figs 2</figureCitation>
, 
<figureCitation id="E8E7B7CBFFE9FF95FBD6F893FBB2EA60" box="[1116,1130,1840,1866]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">3</figureCitation>
) plates totally fused, flat or low conical, white to pale yellow in preserved specimens (
<figureCitation id="E8E7B7CBFFE9FF95FC95F8F0FCBCEA44" box="[799,868,1875,1902]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="8">Fig. 2</figureCitation>
). External surface with 21 low, broad ridges radiating from orifice to shell circumference (
<figureCitation id="E8E7B7CBFFE9FF95FDD5F8D4FD71EAB8" box="[607,681,1911,1938]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="8">Figs 2</figureCitation>
, 
<figureCitation id="E8E7B7CBFFE9FF95FD3DF8DBFD1FEAB8" box="[695,711,1912,1938]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">3</figureCitation>
). 
<typeStatus id="AF6715ECFFE9FF95FD57F8DBFC92EAB8" box="[733,842,1912,1938]" pageId="8" type="paratype">Paratypes</typeStatus>
and other specimens with number of ridges ranging between 19−21 (
<figureCitation id="E8E7B7CBFFE9FF95FEDEF838FE75EA9C" box="[340,429,1947,1974]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="8">Fig. 2C</figureCitation>
). Internal surface of wall smooth, with 21 ribs at rim of shell (
<figureCitation id="E8E7B7CBFFE9FF95FBE6F838FB1EEA9C" box="[1132,1222,1947,1974]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">Fig. 3A</figureCitation>
); sheath 1/3 height of shell, measured at rostral end (
<figureCitation id="E8E7B7CBFFE9FF95FD9AF81CFD53EAF0" box="[528,651,1983,2010]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">Fig. 3A, C</figureCitation>
), surface with a number of growth rings (
<figureCitation id="E8E7B7CBFFE9FF95FBE8F81CFB63EAF0" box="[1122,1211,1983,2010]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">Fig. 3C</figureCitation>
). Orifice (
<figureCitation id="E8E7B7CBFFE9FF95FAA5F81CFAA0EAF0" box="[1327,1400,1983,2010]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="8">Figs 2</figureCitation>
, 
<figureCitation id="E8E7B7CBFFE9FF95FA0FF863FA4BEAF0" box="[1413,1427,1984,2010]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">3</figureCitation>
) ovate, ~ 1/3 of basal rostro-carinal diameter. Basis of shell (
<figureCitation id="E8E7B7CBFFE9FF95FCA2F840FCA7EAD4" box="[808,895,2019,2046]" captionStart="FIGURE 3" captionStartId="5.[151,250,1945,1970]" captionTargetBox="[344,1244,181,1920]" captionTargetId="figure@5.[344,1244,181,1920]" captionTargetPageId="5" captionText="FIGURE 3. Pyrgoma spurtuncata sp. nov.: A, external and internal view of shell (MNHN-IU-2014-23572: holotype); B, external and internal view of shell (MNHN-IU-2014-23573, paratype); C, external and internal view of shell (MNHN-IU-2014-5597, paratype). Scale bars: A−C = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530093" httpUri="https://zenodo.org/record/3530093/files/figure.png" pageId="8">Fig. 3A</figureCitation>
) moderately deep, cup shaped (
<figureCitation id="E8E7B7CBFFE9FF95FB57F840FA57EAD4" box="[1245,1423,2019,2046]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="8">Fig. 2B, D, E, F</figureCitation>
). Top region of basis, at junction between shell and basis, with simple sulci fitting short, longitudinal ribs developed from internal surface of shell (
<figureCitation id="E8E7B7CBFFE8FF94FE66FF1EFD5BEDFD" box="[492,643,189,215]" captionStart="FIGURE 4" captionStartId="6.[151,250,1459,1484]" captionTargetBox="[164,1423,181,1434]" captionTargetId="figure@6.[164,1423,181,1434]" captionTargetPageId="6" captionText="FIGURE 4. Base view of Pyrgoma spurtuncata sp. nov.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A, top view of base without longitudinal ladder-like perforations; B, top view of base with short sections of ladder-like perforations; C, D, top view of base with some ladder-like perforations; E, F, enlarged view of basis; G, H, enlarged view of basis; I, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm" figureDoi="http://doi.org/10.5281/zenodo.3530095" httpUri="https://zenodo.org/record/3530095/files/figure.png" pageId="9">Fig. 4A, E, F</figureCitation>
). In 
<typeStatus id="AF6715ECFFE8FF94FD3FFF1EFCF8EDFD" box="[693,800,189,215]" pageId="9" type="paratype">paratypes</typeStatus>
MNHN-IU-2014-23573 and MNHN-IU-2014-5597, top of basis with short sections of ladder-like, longitudinal perforations (
<figureCitation id="E8E7B7CBFFE8FF94FC07FF42FC16EDD1" box="[909,974,225,251]" captionStart="FIGURE 4" captionStartId="6.[151,250,1459,1484]" captionTargetBox="[164,1423,181,1434]" captionTargetId="figure@6.[164,1423,181,1434]" captionTargetPageId="6" captionText="FIGURE 4. Base view of Pyrgoma spurtuncata sp. nov.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A, top view of base without longitudinal ladder-like perforations; B, top view of base with short sections of ladder-like perforations; C, D, top view of base with some ladder-like perforations; E, F, enlarged view of basis; G, H, enlarged view of basis; I, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm" figureDoi="http://doi.org/10.5281/zenodo.3530095" httpUri="https://zenodo.org/record/3530095/files/figure.png" pageId="9">Fig. 4</figureCitation>
B-D, G-I). In cross-section view (
<figureCitation id="E8E7B7CBFFE8FF94FACBFF42FF76EC35" captionStart="FIGURE 4" captionStartId="6.[151,250,1459,1484]" captionTargetBox="[164,1423,181,1434]" captionTargetId="figure@6.[164,1423,181,1434]" captionTargetPageId="6" captionText="FIGURE 4. Base view of Pyrgoma spurtuncata sp. nov.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A, top view of base without longitudinal ladder-like perforations; B, top view of base with short sections of ladder-like perforations; C, D, top view of base with some ladder-like perforations; E, F, enlarged view of basis; G, H, enlarged view of basis; I, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm" figureDoi="http://doi.org/10.5281/zenodo.3530095" httpUri="https://zenodo.org/record/3530095/files/figure.png" pageId="9">Fig. 4C, D</figureCitation>
), basis developing into extensive ladder-like, longitudinal perforations (
<figureCitation id="E8E7B7CBFFE8FF94FC5CFEA6FBC2EC35" box="[982,1050,260,287]" captionStart="FIGURE 5" captionStartId="7.[151,250,1910,1935]" captionTargetBox="[191,1395,182,1884]" captionTargetId="figure@7.[191,1396,181,1885]" captionTargetPageId="7" captionText="FIGURE 5. Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-5597): A, Light microscopy view showing the shell and basis together; B, Scanning Electron Microcopy (SEM) of shell and basis, showing the longitudinal ladder-like perforations; C−F, SEM views showing side view (C,D) and top view (E, F) of basis and the ladder-like longitudinal perforations. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3530097" httpUri="https://zenodo.org/record/3530097/files/figure.png" pageId="9">Fig. 5</figureCitation>
); between ladder-like longitudinal perforations, single-row of deep grooves located between ladder-like longitudinal ridges, groove widening at shell rims (
<figureCitation id="E8E7B7CBFFE8FF94FF52FEEEFE8BEC4D" box="[216,339,332,359]" captionStart="FIGURE 5" captionStartId="7.[151,250,1910,1935]" captionTargetBox="[191,1395,182,1884]" captionTargetId="figure@7.[191,1396,181,1885]" captionTargetPageId="7" captionText="FIGURE 5. Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-5597): A, Light microscopy view showing the shell and basis together; B, Scanning Electron Microcopy (SEM) of shell and basis, showing the longitudinal ladder-like perforations; C−F, SEM views showing side view (C,D) and top view (E, F) of basis and the ladder-like longitudinal perforations. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.3530097" httpUri="https://zenodo.org/record/3530097/files/figure.png" pageId="9">Fig. 5C, D</figureCitation>
).
</paragraph>
<paragraph id="7063AB4EFFE8FF94FF4DFED2FA51EF15" blockId="9.[151,1437,153,2015]" pageId="9">
Scutum (
<figureCitation id="E8E7B7CBFFE8FF94FEA7FED2FEA8ECA6" box="[301,368,369,396]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6</figureCitation>
) triangular, height similar to width, several small teeth along ventral surface (
<figureCitation id="E8E7B7CBFFE8FF94FB54FED2FAADECA1" box="[1246,1397,369,395]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6B, D, F</figureCitation>
) of occludent margin. Tergal margin straight, less than 1/2 length of occludent margin. External view with visible adductor plate, deeply projecting below basal margin, reaching 1/3 height of scutum (
<figureCitation id="E8E7B7CBFFE8FF94FBCFFE1AFB39ECF9" box="[1093,1249,441,467]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6A, C, E</figureCitation>
). Adductor plate with blunt, thick, short rostral tooth (
<figureCitation id="E8E7B7CBFFE8FF94FDBFFE7EFD17ECDD" box="[565,719,477,503]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6A, C, E</figureCitation>
), height of tooth less than 1/3 height of adductor plate (
<figureCitation id="E8E7B7CBFFE8FF94FAB4FE7EFF74EF31" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6A, B</figureCitation>
). External surface with slightly sinuous striations, striations bearing rows of small pores. Internal surface with oval-shaped adductor muscle scar (
<figureCitation id="E8E7B7CBFFE8FF94FDABFD86FD61EF15" box="[545,697,549,575]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6B, D, F</figureCitation>
). Scutum of 
<typeStatus id="AF6715ECFFE8FF94FCC2FD86FC6EEF15" box="[840,950,549,575]" pageId="9" type="paratype">paratypes</typeStatus>
, consistent with 
<typeStatus id="AF6715ECFFE8FF94FBFBFD86FB0BEF15" box="[1137,1235,549,575]" pageId="9" type="holotype">holotype</typeStatus>
(
<figureCitation id="E8E7B7CBFFE8FF94FB69FD86FAA5EF15" box="[1251,1405,549,575]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6A, C, E</figureCitation>
).
</paragraph>
<paragraph id="7063AB4EFFE8FF94FF4DFDEAFB51EFE5" blockId="9.[151,1437,153,2015]" pageId="9">
Tergum (
<figureCitation id="E8E7B7CBFFE8FF94FEA1FDEAFEB5EF4E" box="[299,365,585,612]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6</figureCitation>
) moderately broad, upper part triangular, higher than wide, height to width ratio 3:1, apical angle 68°. Distinct spur with furrow reaching 3/5 height of tergum; medial furrow fused at mid length of tergum, distal end of spur truncated (
<figureCitation id="E8E7B7CBFFE8FF94FE1EFD32FE00EF86" box="[404,472,657,684]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6</figureCitation>
). External surface with slightly sinuous growth lines; basal margin not obvious due to spur (
<figureCitation id="E8E7B7CBFFE8FF94FF5DFD16FEA9EFE5" box="[215,369,693,719]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6A, C, E</figureCitation>
). Internal surface without crests for depressor muscle (
<figureCitation id="E8E7B7CBFFE8FF94FC5CFD16FBC3EFE5" box="[982,1051,693,719]" captionStart="FIGURE 6" captionStartId="8.[151,250,1516,1541]" captionTargetBox="[151,1436,181,1491]" captionTargetId="figure@8.[151,1436,181,1491]" captionTargetPageId="8" captionText="FIGURE 6. Scanning electron micrograph of scuta and terga of Pyrgoma spurtuncata sp. nov.: A, external view of scuta and terga (MNHN-IU-2014-23572: holotype); B, internal view of scutum and tergum (MNHN-IU-2014-23572: holotype); C, external view of scutum and tergum (MNHN-IU-2014-23573, Paratype); D, internal view of scutum and tergum (MNHN-IU- 2014-23573, Paratype); E, external view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype); F, internal view of scutum and tergum (MNHN-IU-2014-5597, additional Paratype). Scale bars: A−F = 500µm." figureDoi="http://doi.org/10.5281/zenodo.3530099" httpUri="https://zenodo.org/record/3530099/files/figure.png" pageId="9">Fig. 6</figureCitation>
, B, D, F).
</paragraph>
<paragraph id="7063AB4EFFE8FF94FF1DFD7AFE6AEEE1" blockId="9.[151,1437,153,2015]" pageId="9">
Maxilla (
<figureCitation id="E8E7B7CBFFE8FF94FF75FD7AFE80EFD9" box="[255,344,729,755]" captionStart="FIGURE 7" captionStartId="10.[151,250,1832,1857]" captionTargetBox="[280,1308,237,1808]" captionTargetId="figure@10.[280,1308,237,1808]" captionTargetPageId="10" captionText="FIGURE 7. Mouthparts of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, maxilla; B, mandibular palp; C, maxillule; D, mandible; E, lower margin and inferior angle of mandible; F, inferior margin of the mandible; G, labrum; H, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530101" httpUri="https://zenodo.org/record/3530101/files/figure.png" pageId="9">Fig. 7A</figureCitation>
) bilobed, with dense, serrulate setae on margins of both lobes. Mandibular palp (
<figureCitation id="E8E7B7CBFFE8FF94FB7BFD7AFA90EFD9" box="[1265,1352,729,755]" captionStart="FIGURE 7" captionStartId="10.[151,250,1832,1857]" captionTargetBox="[280,1308,237,1808]" captionTargetId="figure@10.[280,1308,237,1808]" captionTargetPageId="10" captionText="FIGURE 7. Mouthparts of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, maxilla; B, mandibular palp; C, maxillule; D, mandible; E, lower margin and inferior angle of mandible; F, inferior margin of the mandible; G, labrum; H, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530101" httpUri="https://zenodo.org/record/3530101/files/figure.png" pageId="9">Fig. 7B</figureCitation>
) ovate, bearing simple setae on distal end and outer and inner margins. Maxillule (
<figureCitation id="E8E7B7CBFFE8FF94FC6BFD5EFBEFEE3D" box="[993,1079,765,791]" captionStart="FIGURE 7" captionStartId="10.[151,250,1832,1857]" captionTargetBox="[280,1308,237,1808]" captionTargetId="figure@10.[280,1308,237,1808]" captionTargetPageId="10" captionText="FIGURE 7. Mouthparts of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, maxilla; B, mandibular palp; C, maxillule; D, mandible; E, lower margin and inferior angle of mandible; F, inferior margin of the mandible; G, labrum; H, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530101" httpUri="https://zenodo.org/record/3530101/files/figure.png" pageId="9">Fig. 7C</figureCitation>
) with small notch, 2 large setae above notch, row of 8 developed setae below notch; outer and inner margins with long, simple setae. Mandible (
<figureCitation id="E8E7B7CBFFE8FF94FAE5FC82FF7CEE75" captionStart="FIGURE 7" captionStartId="10.[151,250,1832,1857]" captionTargetBox="[280,1308,237,1808]" captionTargetId="figure@10.[280,1308,237,1808]" captionTargetPageId="10" captionText="FIGURE 7. Mouthparts of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, maxilla; B, mandibular palp; C, maxillule; D, mandible; E, lower margin and inferior angle of mandible; F, inferior margin of the mandible; G, labrum; H, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530101" httpUri="https://zenodo.org/record/3530101/files/figure.png" pageId="9">Fig. 7</figureCitation>
D−F) with 5 teeth excluding inferior angle; second to fourth teeth bifid; first 3 teeth occupying 3/4 length of cutting edge; fifth tooth blunt, close to inferior angle. Inferior angle with few large setae. Lateral side, lower margin and cutting edge of mandible bearing simple setae. Labrum (
<figureCitation id="E8E7B7CBFFE8FF94FC84FC2EFC5FEE8D" box="[782,903,909,935]" captionStart="FIGURE 7" captionStartId="10.[151,250,1832,1857]" captionTargetBox="[280,1308,237,1808]" captionTargetId="figure@10.[280,1308,237,1808]" captionTargetPageId="10" captionText="FIGURE 7. Mouthparts of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, maxilla; B, mandibular palp; C, maxillule; D, mandible; E, lower margin and inferior angle of mandible; F, inferior margin of the mandible; G, labrum; H, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530101" httpUri="https://zenodo.org/record/3530101/files/figure.png" pageId="9">Fig. 7G, H</figureCitation>
) with deep, V-shaped notch separating crests; 2 small teeth on each crest.
</paragraph>
<paragraph id="7063AB4EFFE8FF94FF4DFC76FCE5E825" blockId="9.[151,1437,153,2015]" pageId="9">
Cirrus I (
<figureCitation id="E8E7B7CBFFE8FF94FEBAFC76FEAAEEC5" box="[304,370,981,1007]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1936]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure@11.[177,1411,181,1885]" captionTargetPageId="11" captionText="FIGURE 8. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus I; B, C, distal serrulate setae on posterior and anterior rami of cirrus I; D, cirrus II; E, distal serrulate setae on posterior and anterior rami of cirrus II; F, cirrus III; G. magnified view of cirrus II; H, I, distal serrulate setae on posterior and anterior rami of cirrus III. Scale bars: A−I = 100µm." figureDoi="http://doi.org/10.5281/zenodo.3530103" httpUri="https://zenodo.org/record/3530103/files/figure.png" pageId="9">Fig. 8</figureCitation>
A−C) with unequal rami; anterior ramus long, slender, 12 segmented, bearing serrulate setae; posterior ramus short, stout, 6 segmented, bearing plumose setae on first segment (
<figureCitation id="E8E7B7CBFFE8FF94FBB5FC5AFB40E939" box="[1087,1176,1017,1043]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1936]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure@11.[177,1411,181,1885]" captionTargetPageId="11" captionText="FIGURE 8. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus I; B, C, distal serrulate setae on posterior and anterior rami of cirrus I; D, cirrus II; E, distal serrulate setae on posterior and anterior rami of cirrus II; F, cirrus III; G. magnified view of cirrus II; H, I, distal serrulate setae on posterior and anterior rami of cirrus III. Scale bars: A−I = 100µm." figureDoi="http://doi.org/10.5281/zenodo.3530103" httpUri="https://zenodo.org/record/3530103/files/figure.png" pageId="9">Fig. 8B</figureCitation>
). Cirrus II (
<figureCitation id="E8E7B7CBFFE8FF94FA94FC5AFF74E91D" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1936]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure@11.[177,1411,181,1885]" captionTargetPageId="11" captionText="FIGURE 8. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus I; B, C, distal serrulate setae on posterior and anterior rami of cirrus I; D, cirrus II; E, distal serrulate setae on posterior and anterior rami of cirrus II; F, cirrus III; G. magnified view of cirrus II; H, I, distal serrulate setae on posterior and anterior rami of cirrus III. Scale bars: A−I = 100µm." figureDoi="http://doi.org/10.5281/zenodo.3530103" httpUri="https://zenodo.org/record/3530103/files/figure.png" pageId="9">Fig. 8D, E, G</figureCitation>
) with un-equal rami; anterior ramus 7 segmented, bearing simple setae, posterior ramus 5 segmented, first segment bearing plumose setae. Cirrus III (
<figureCitation id="E8E7B7CBFFE8FF94FDE8FBE2FD2BE971" box="[610,755,1089,1115]" captionStart="FIGURE 8" captionStartId="11.[151,250,1911,1936]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure@11.[177,1411,181,1885]" captionTargetPageId="11" captionText="FIGURE 8. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus I; B, C, distal serrulate setae on posterior and anterior rami of cirrus I; D, cirrus II; E, distal serrulate setae on posterior and anterior rami of cirrus II; F, cirrus III; G. magnified view of cirrus II; H, I, distal serrulate setae on posterior and anterior rami of cirrus III. Scale bars: A−I = 100µm." figureDoi="http://doi.org/10.5281/zenodo.3530103" httpUri="https://zenodo.org/record/3530103/files/figure.png" pageId="9">Fig. 8F, H, I</figureCitation>
) anterior ramus 9 segmented, slightly longer than posterior ramus (7 segmented), first segment of posterior ramus bearing plumose setae. Cirri IV−VI (
<figureCitation id="E8E7B7CBFFE8FF94FB24FBC6FB2BE955" box="[1198,1267,1125,1151]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">Fig. 9</figureCitation>
) long, slender, rami of equal size. Cirrus IV (
<figureCitation id="E8E7B7CBFFE8FF94FE60FB2AFD9AE989" box="[490,578,1161,1187]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">Fig. 9A</figureCitation>
) with anterior ramus 13 segmented, posterior ramus 15 segmented; cirrus V (
<figureCitation id="E8E7B7CBFFE8FF94FF2AFB0EFF2EE9ED" box="[160,246,1197,1223]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">Fig. 9C</figureCitation>
) with anterior ramus 16 segmented, posterior ramus 17 segmented, cirrus VI (
<figureCitation id="E8E7B7CBFFE8FF94FBD4FB0EFB6BE9ED" box="[1118,1203,1197,1223]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">Fig. 9E</figureCitation>
) with anterior ramus 16 segmented, posterior ramus 14 segmented. Intermediate segments of cirri IV−VI (
<figureCitation id="E8E7B7CBFFE8FF94FBD9FB72FB32E9C1" box="[1107,1258,1233,1259]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">Fig. 9B, D, F</figureCitation>
) with 3 pairs of simple setae, distal two pairs longest, proximal pair shortest.
</paragraph>
<paragraph id="7063AB4EFFE8FF94FF4DFABAFDB7E87D" blockId="9.[151,1437,153,2015]" pageId="9">
Penis (
<figureCitation id="E8E7B7CBFFE8FF94FE98FABAFE52E819" box="[274,394,1305,1331]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">Fig. 9G, H</figureCitation>
) long (~1.2 times length of cirrus VI), annulated, basidorsal point acute, setae simple; apex of penis bearing short, simple setae (
<figureCitation id="E8E7B7CBFFE8FF94FD99FA9EFDBAE87D" box="[531,610,1341,1367]" captionStart="FIGURE 9" captionStartId="12.[151,250,1946,1971]" captionTargetBox="[157,1431,181,1920]" captionTargetId="figure@12.[157,1431,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Cirri of Pyrgoma spurtuncata sp. nov. (MNHN-IU-2014-23572: holotype): A, cirrus IV; B, intermediate segments of cirrus IV; C, cirrus V; D, intermediate segments of cirrus V; E, cirrus VI; F, intermediate segments of cirrus VI; G, penis; H, basidorsal point of penis; I, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm." figureDoi="http://doi.org/10.5281/zenodo.3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="9">
Fig. 
<date id="04628D8EFFE8FF94FDC2FA9EFDBAE87D" box="[584,610,1341,1367]" pageId="9">9I</date>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="38C6F8C5FFE8FF94FF4DFAC2FD47E8EE" pageId="9" type="etymology">
<paragraph id="7063AB4EFFE8FF94FF4DFAC2FD47E8EE" blockId="9.[151,1437,153,2015]" pageId="9">
<emphasis id="42A8775CFFE8FF94FF4DFAC2FE96E851" bold="true" box="[199,334,1377,1403]" pageId="9">Etymology.</emphasis>
Compared to 
<emphasis id="42A8775CFFE8FF94FE7AFAC1FBD5E851" box="[496,1037,1377,1403]" italics="true" pageId="9">
<taxonomicName id="B7DCD0CDFFE8FF94FE7AFAC1FDB7E851" authorityName="Weltner" authorityYear="1897" box="[496,623,1378,1403]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="japonica">P. japonica</taxonomicName>
and 
<taxonomicName id="B7DCD0CDFFE8FF94FD23FAC1FC8BE851" box="[681,851,1377,1403]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="cancallatum">P. cancallatum</taxonomicName>
, 
<taxonomicName id="B7DCD0CDFFE8FF94FCD5FAC1FBD5E851" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[863,1037,1378,1403]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">P. spurtruncata</taxonomicName>
</emphasis>
<emphasis id="42A8775CFFE8FF94FB9FFAC2FBB7E851" bold="true" box="[1045,1135,1377,1403]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FB9FFAC2FBB7E851" box="[1045,1135,1377,1403]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a wider, shorter tergal spur, and a blunt rostral tooth of the scutum. The specific name 
<taxonomicName id="B7DCD0CDFFE8FF94FCF8FA25FBDBE8B5" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[882,1027,1414,1439]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata">
<emphasis id="42A8775CFFE8FF94FCF8FA25FBDBE8B5" box="[882,1027,1414,1439]" italics="true" pageId="9">spurtruncata</emphasis>
</taxonomicName>
refers to the wide, short spur of the tergum (the Latin word 
<emphasis id="42A8775CFFE8FF94FE28FA09FDD9E8E9" box="[418,513,1450,1475]" italics="true" pageId="9">truncata</emphasis>
means short).
</paragraph>
</subSubSection>
<subSubSection id="38C6F8C5FFE8FF94FF4DFA6FFD9FEB21" pageId="9" type="distribution">
<paragraph id="7063AB4EFFE8FF94FF4DFA6FFD9FEB21" blockId="9.[151,1437,153,2015]" pageId="9">
<emphasis id="42A8775CFFE8FF94FF4DFA6FFEB9E8CD" bold="true" box="[199,353,1484,1511]" pageId="9">Distribution.</emphasis>
Presently recorded only from the deep-waters of 
<collectingCountry id="08CBEBDEFFE8FF94FC03FA6EFBEFE8CD" box="[905,1079,1485,1511]" name="New Caledonia" pageId="9">New Caledonia</collectingCountry>
and only from the coral 
<taxonomicName id="B7DCD0CDFFE8FF94FAC3FA6EFEFFEB21" authorityName="Linnaeus" authorityYear="1758" class="Anthozoa" family="Oculinidae" genus="Madrepora" kingdom="Animalia" order="Scleractinia" pageId="9" phylum="Cnidaria" rank="species" species="oculata">
<emphasis id="42A8775CFFE8FF94FAC3FA6EFEFFEB21" italics="true" pageId="9">Madrepora oculata</emphasis>
</taxonomicName>
in the family 
<taxonomicName id="B7DCD0CDFFE8FF94FE4EFA52FD9BEB21" box="[452,579,1521,1547]" class="Anthozoa" family="Oculinidae" kingdom="Animalia" order="Scleractinia" pageId="9" phylum="Cnidaria" rank="family">Oculinidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="38C6F8C5FFE8FF93FF4DF9B6FC48EFF3" lastPageId="14" pageId="9" type="discussion">
<paragraph id="7063AB4EFFE8FF97FF4DF9B6FF31EDFD" blockId="9.[151,1437,153,2015]" lastBlockId="10.[151,1436,153,215]" lastPageId="10" pageId="9">
<emphasis id="42A8775CFFE8FF94FF4DF9B6FEE3EB05" bold="true" box="[199,315,1557,1583]" pageId="9">Remarks.</emphasis>
In their revision of 
<taxonomicName id="B7DCD0CDFFE8FF94FD93F9B5FC77EB05" authority=", Ross and Newman (2002)" authorityName=", Ross and Newman" authorityYear="2002" box="[537,943,1557,1583]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFE8FF94FD93F9B5FDA7EB05" box="[537,639,1558,1583]" italics="true" pageId="9">Pyrgoma</emphasis>
, 
<bibRefCitation id="144DD6BFFFE8FF94FD06F9B6FC77EB05" author="Ross, A. &amp; Newman, W. A." box="[652,943,1557,1583]" pageId="9" pagination="407 - 421" refId="ref8876" refString="Ross, A. &amp; Newman, W. A. (2002) A review of Pyrgoma cancellatum species complex (Cirripedia: Pyrgomatidae). Journal of Natural History, 36, 407 - 421. https: // doi. org / 10.1080 / 00222930010009336" type="journal article" year="2002">Ross and Newman (2002)</bibRefCitation>
</taxonomicName>
divided 
<taxonomicName id="B7DCD0CDFFE8FF94FB9AF9B5FBAEEB05" authorityName=", Ross and Newman" authorityYear="2002" box="[1040,1142,1558,1583]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFE8FF94FB9AF9B5FBAEEB05" box="[1040,1142,1558,1583]" italics="true" pageId="9">Pyrgoma</emphasis>
</taxonomicName>
into two groups, based on the presence/absence of extended muscles crests of the tergum. 
<taxonomicName id="B7DCD0CDFFE8FF94FCD7F999FB8AEB79" authorityName="Leach" authorityYear="1818" box="[861,1106,1593,1619]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFE8FF94FCD7F999FB8AEB79" box="[861,1106,1593,1619]" italics="true" pageId="9">Pyrgoma cancellatum</emphasis>
</taxonomicName>
(
<figureCitation id="E8E7B7CBFFE8FF94FBEBF99AFADFEB79" box="[1121,1287,1593,1619]" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10A, B, E</figureCitation>
), 
<taxonomicName id="B7DCD0CDFFE8FF94FA91F999FA44EB79" authorityName="Weltner" authorityYear="1897" box="[1307,1436,1593,1619]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFE8FF94FA91F999FAF4EB79" box="[1307,1324,1594,1619]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FAB0F999FA44EB79" box="[1338,1436,1594,1619]" italics="true" pageId="9">japonica</emphasis>
</taxonomicName>
(
<figureCitation id="E8E7B7CBFFE8FF94FF2AF9FEFEF0EB5D" box="[160,296,1629,1655]" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10C, D</figureCitation>
) and 
<taxonomicName id="B7DCD0CDFFE8FF94FEE0F9FDFE60EB5D" authorityName="Hoek" authorityYear="1913" box="[362,440,1629,1655]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="kuri">
<emphasis id="42A8775CFFE8FF94FEE0F9FDFEA3EB5D" box="[362,379,1630,1655]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FE01F9FEFE60EB5D" box="[395,440,1629,1655]" italics="true" pageId="9">kuri</emphasis>
</taxonomicName>
(see 
<figureCitation id="E8E7B7CBFFE8FF94FE7CF9FEFDE1EB5D" box="[502,569,1629,1655]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="9">Fig. 2</figureCitation>
for SEM plate of scutum and tergum in 
<bibRefCitation id="144DD6BFFFE8FF94FB80F9FEFACEEB5D" author="Ross, A. &amp; Newman, W. A." box="[1034,1302,1629,1655]" pageId="9" pagination="245 - 260" refId="ref8813" refString="Ross, A. &amp; Newman, W. A. (2000) Pyrgoma kuri Hoek, 1913: A case study in morphology and systematics of a symbiotic coral barnacle (Cirripedia: Balanomorpha). Contributions to Zoology, 68 (4), 245 - 260. https: // doi. org / 10.1163 / 18759866 - 06804002" type="journal article" year="2000">Ross &amp; Newman, 2000</bibRefCitation>
) lack tergal muscle crests. In contrast, 
<taxonomicName id="B7DCD0CDFFE8FF94FE4EF921FD8DEBB1" authorityName="Nilsson-Cantell" authorityYear="1938" box="[452,597,1665,1691]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="projectum">
<emphasis id="42A8775CFFE8FF94FE4EF921FE0DEBB1" box="[452,469,1666,1691]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FE6EF921FD8DEBB1" box="[484,597,1666,1691]" italics="true" pageId="9">projectum</emphasis>
</taxonomicName>
(
<figureCitation id="E8E7B7CBFFE8FF94FDEEF922FD1EEBB1" box="[612,710,1665,1691]" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10F</figureCitation>
) and 
<taxonomicName id="B7DCD0CDFFE8FF94FC8DF921FCB2EBB1" baseAuthorityName="Ren" baseAuthorityYear="1986" box="[775,874,1665,1691]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="42A8775CFFE8FF94FC8DF921FCC0EBB1" box="[775,792,1666,1691]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FCADF921FCB2EBB1" box="[807,874,1666,1691]" italics="true" pageId="9">sinica</emphasis>
</taxonomicName>
(
<figureCitation id="E8E7B7CBFFE8FF94FCF0F922FC39EBB1" box="[890,993,1665,1691]" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10G</figureCitation>
) have extended tergal muscle crests on the tergum. 
<taxonomicName id="B7DCD0CDFFE8FF94FEAAF905FDC7EB95" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[288,543,1702,1727]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE8FF94FEAAF905FDC7EB95" box="[288,543,1702,1727]" italics="true" pageId="9">Pyrgoma spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFE8FF94FDADF906FD5AEB95" bold="true" box="[551,642,1701,1727]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FDADF906FD5AEB95" box="[551,642,1701,1727]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically similar to 
<taxonomicName id="B7DCD0CDFFE8FF94FC51F905FB5CEB95" authorityName="Leach" authorityYear="1818" box="[987,1156,1701,1727]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFE8FF94FC51F905FC34EB95" box="[987,1004,1702,1727]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FC71F906FB5CEB95" box="[1019,1156,1701,1727]" italics="true" pageId="9">cancellatum</emphasis>
</taxonomicName>
, 
<taxonomicName id="B7DCD0CDFFE8FF94FB19F905FACEEB95" authorityName="Weltner" authorityYear="1897" box="[1171,1302,1701,1727]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFE8FF94FB19F905FB7CEB95" box="[1171,1188,1702,1727]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FB3EF905FACEEB95" box="[1204,1302,1702,1727]" italics="true" pageId="9">japonica</emphasis>
</taxonomicName>
and 
<taxonomicName id="B7DCD0CDFFE8FF94FAC5F905FA44EB95" authorityName="Hoek" authorityYear="1913" box="[1359,1436,1701,1727]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="kuri">
<emphasis id="42A8775CFFE8FF94FAC5F905FAB8EB95" box="[1359,1376,1702,1727]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FAE5F906FA44EB95" box="[1391,1436,1701,1727]" italics="true" pageId="9">kuri</emphasis>
</taxonomicName>
in sharing an absence of tergal muscle crests. 
<taxonomicName id="B7DCD0CDFFE8FF94FD16F969FC42EBC9" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[668,922,1738,1763]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE8FF94FD16F969FC42EBC9" box="[668,922,1738,1763]" italics="true" pageId="9">Pyrgoma spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFE8FF94FC28F96AFC24EBC9" bold="true" box="[930,1020,1737,1763]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FC28F96AFC24EBC9" box="[930,1020,1737,1763]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however, can be distinguished from 
<taxonomicName id="B7DCD0CDFFE8FF94FF1DF94DFE9AEA2D" authorityName="Leach" authorityYear="1818" box="[151,322,1773,1799]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFE8FF94FF1DF94DFF70EA2D" box="[151,168,1774,1799]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FF33F94EFE9AEA2D" box="[185,322,1773,1799]" italics="true" pageId="9">cancellatum</emphasis>
</taxonomicName>
by the shape of the medial furrow on the external surface of the tergum (
<figureCitation id="E8E7B7CBFFE8FF94FB17F94EFA95EA2D" box="[1181,1357,1773,1799]" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10A, B, E</figureCitation>
). In 
<taxonomicName id="B7DCD0CDFFE8FF94FA0EF94DFEF8EA01" authorityName="Leach" authorityYear="1818" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFE8FF94FA0EF94DFA4DEA2D" box="[1412,1429,1774,1799]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FF1DF8B2FEF8EA01" box="[151,288,1809,1835]" italics="true" pageId="9">cancellatum</emphasis>
</taxonomicName>
(
<figureCitation id="E8E7B7CBFFE8FF94FEA5F8B2FE4EEA01" box="[303,406,1809,1835]" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10B</figureCitation>
), the medial furrow of the tergum is shallow and fully open. In 
<taxonomicName id="B7DCD0CDFFE8FF94FBD5F8B1FAD3EA01" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[1119,1291,1810,1835]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE8FF94FBD5F8B1FAD3EA01" box="[1119,1291,1810,1835]" italics="true" pageId="9">P. spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFE8FF94FA98F8B2FAB4EA01" bold="true" box="[1298,1388,1809,1835]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FA98F8B2FAB4EA01" box="[1298,1388,1809,1835]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the medial furrow is deeper and closed along half the length of the tergum. 
<taxonomicName id="B7DCD0CDFFE8FF94FC3FF895FB6AEA65" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[949,1202,1846,1871]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE8FF94FC3FF895FB6AEA65" box="[949,1202,1846,1871]" italics="true" pageId="9">Pyrgoma spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFE8FF94FB33F896FACAEA65" bold="true" box="[1209,1298,1845,1871]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FB33F896FACAEA65" box="[1209,1298,1845,1871]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="B7DCD0CDFFE8FF94FF1DF8F9FEC1EA59" authorityName="Weltner" authorityYear="1897" box="[151,281,1881,1907]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFE8FF94FF1DF8F9FF70EA59" box="[151,168,1882,1907]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FF3DF8F9FEC1EA59" box="[183,281,1882,1907]" italics="true" pageId="9">japonica</emphasis>
</taxonomicName>
in that the rostral teeth of both species are distinct and blunt, and the external furrow of the tergum is closed. Compared to 
<taxonomicName id="B7DCD0CDFFE8FF94FE0DF8DDFDDDEABD" authorityName="Weltner" authorityYear="1897" box="[391,517,1918,1943]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFE8FF94FE0DF8DDFDDDEABD" box="[391,517,1918,1943]" italics="true" pageId="9">P. japonica</emphasis>
</taxonomicName>
, the tergal spur of 
<taxonomicName id="B7DCD0CDFFE8FF94FD51F8DDFC54EABD" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[731,908,1917,1943]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE8FF94FD51F8DDFD34EABD" box="[731,748,1918,1943]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FD71F8DDFC54EABD" box="[763,908,1918,1943]" italics="true" pageId="9">spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFE8FF94FC19F8DEFC35EABD" bold="true" box="[915,1005,1917,1943]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FC19F8DEFC35EABD" box="[915,1005,1917,1943]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is much wider, and the ratio of length to width of the spur is 5.5:1 (compared to the slender and narrow tergal spur in 
<taxonomicName id="B7DCD0CDFFE8FF94FB83F801FB51EA91" authorityName="Weltner" authorityYear="1897" box="[1033,1161,1953,1979]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFE8FF94FB83F801FBC2EA91" box="[1033,1050,1954,1979]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FBADF801FB51EA91" box="[1063,1161,1954,1979]" italics="true" pageId="9">japonica</emphasis>
</taxonomicName>
with a ratio of 9:1) (
<figureCitation id="E8E7B7CBFFE8FF94FAE5F802FF14EAF5" captionStart="FIGURE 10" captionStartId="13.[151,250,1832,1857]" captionTargetBox="[177,1411,525,1807]" captionTargetId="figure@13.[177,1411,525,1807]" captionTargetPageId="13" captionText="FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of Pyrgoma: A, top and basal view of P. cancellatum (Taiwan, Chan et al., 2013b); B, scutum and tergum of P. cancellatum (Taiwan, Chan et al., 2013b); C, top and basal view of P. japonica (Taiwan, present study); D, scutum and tergum of P. japonica (Taiwan, present study); E. redrawing of opercular plates of P. cancellatum from Hiro (1935); F, redrawing of opercular plates of P. projectum from Nilsson-Cantell (1938); G, redrawing of opercular plates of P. sinica from Ren (1986). Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm." figureDoi="http://doi.org/10.5281/zenodo.3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="9">Fig. 10D</figureCitation>
). Similar to 
<taxonomicName id="B7DCD0CDFFE8FF94FEDFF865FDDAEAF5" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[341,514,1990,2015]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFE8FF94FEDFF865FDDAEAF5" box="[341,514,1990,2015]" italics="true" pageId="9">P. spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFE8FF94FD82F866FDBAEAF5" bold="true" box="[520,610,1989,2015]" pageId="9">
<taxonomicNameLabel id="599BCA27FFE8FF94FD82F866FDBAEAF5" box="[520,610,1989,2015]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="B7DCD0CDFFE8FF94FDFAF865FD64EAF5" authorityName="Hoek" authorityYear="1913" box="[624,700,1989,2015]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="kuri">
<emphasis id="42A8775CFFE8FF94FDFAF865FD59EAF5" box="[624,641,1990,2015]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FD05F866FD64EAF5" box="[655,700,1989,2015]" italics="true" pageId="9">kuri</emphasis>
</taxonomicName>
has a truncated distal end of the spur of the tergum, but in 
<taxonomicName id="B7DCD0CDFFE8FF94FADAF865FA44EAF5" authorityName="Hoek" authorityYear="1913" box="[1360,1436,1989,2015]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="9" phylum="Arthropoda" rank="species" species="kuri">
<emphasis id="42A8775CFFE8FF94FADAF865FAB9EAF5" box="[1360,1377,1990,2015]" italics="true" pageId="9">P</emphasis>
. 
<emphasis id="42A8775CFFE8FF94FAE5F866FA44EAF5" box="[1391,1436,1989,2015]" italics="true" pageId="9">kuri</emphasis>
</taxonomicName>
the tergal furrow is open (see 
<figureCitation id="E8E7B7CBFFEBFF97FE55FF3AFDE4ED99" box="[479,572,153,179]" captionStart="FIGURE 2" captionStartId="4.[151,250,1975,2000]" captionTargetBox="[189,1398,597,1949]" captionTargetId="figure@4.[189,1398,597,1949]" captionTargetPageId="4" captionText="FIGURE 2. Pyrgoma spurtuncata sp. nov.: A (upper-lateral view), B (lateral view) MNHN-IU-2014-23572 (holotype); C (top view), D (lateral view) (paratype) MNHN-IU-2014-5597; E, F (upper lateral view) (paratype) MNHN-IU-2014-5597." figureDoi="http://doi.org/10.5281/zenodo.3530091" httpUri="https://zenodo.org/record/3530091/files/figure.png" pageId="10">
Figure 
<quantity id="B72406ABFFEBFF97FDA4FF3AFDE4ED99" box="[558,572,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" unit="in" value="2.0">2</quantity>
</figureCitation>
in 
<bibRefCitation id="144DD6BFFFEBFF97FDD4FF3AFCBDED99" author="Ross, A. &amp; Newman, W. A." box="[606,869,153,179]" pageId="10" pagination="245 - 260" refId="ref8813" refString="Ross, A. &amp; Newman, W. A. (2000) Pyrgoma kuri Hoek, 1913: A case study in morphology and systematics of a symbiotic coral barnacle (Cirripedia: Balanomorpha). Contributions to Zoology, 68 (4), 245 - 260. https: // doi. org / 10.1163 / 18759866 - 06804002" type="journal article" year="2000">Ross &amp; Newman, 2000</bibRefCitation>
), whilst 
<taxonomicName id="B7DCD0CDFFEBFF97FC48FF39FBB5ED99" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[962,1133,154,179]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="10" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFEBFF97FC48FF39FBB5ED99" box="[962,1133,154,179]" italics="true" pageId="10">P. spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFEBFF97FBF9FF3AFB14ED99" bold="true" box="[1139,1228,153,179]" pageId="10">
<taxonomicNameLabel id="599BCA27FFEBFF97FBF9FF3AFB14ED99" box="[1139,1228,153,179]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a closed tergal furrow.
</paragraph>
<caption id="24A3FBC6FFEBFF97FF1DF88BFC44EAA3" ID-DOI="http://doi.org/10.5281/zenodo.3530101" ID-Zenodo-Dep="3530101" httpUri="https://zenodo.org/record/3530101/files/figure.png" pageId="10" startId="10.[151,250,1832,1857]" targetBox="[280,1308,237,1808]" targetPageId="10">
<paragraph id="7063AB4EFFEBFF97FF1DF88BFC44EAA3" blockId="10.[151,1436,1832,1929]" pageId="10">
<emphasis id="42A8775CFFEBFF97FF1DF88BFB78EA6B" bold="true" box="[151,1184,1832,1857]" pageId="10">
FIGURE 7. Mouthparts of 
<taxonomicName id="B7DCD0CDFFEBFF97FE49F889FD7EEA6B" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[451,678,1834,1857]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="10" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFEBFF97FE49F889FD7EEA6B" bold="true" box="[451,678,1834,1857]" italics="true" pageId="10">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFEBFF97FD25F88AFCD9EA6B" box="[687,769,1833,1857]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
(MNHN-IU-2014-23572: holotype): A
</emphasis>
, maxilla; 
<emphasis id="42A8775CFFEBFF97FA80F88AFAC3EA6B" bold="true" box="[1290,1307,1833,1857]" pageId="10">B</emphasis>
, mandibular palp; 
<emphasis id="42A8775CFFEBFF97FF45F8EFFF39EA4F" bold="true" box="[207,225,1868,1893]" pageId="10">C</emphasis>
, maxillule; 
<emphasis id="42A8775CFFEBFF97FED0F8EEFEB4EA4F" bold="true" box="[346,364,1869,1893]" pageId="10">D</emphasis>
, mandible; 
<emphasis id="42A8775CFFEBFF97FE69F8EEFE2CEA4F" bold="true" box="[483,500,1869,1893]" pageId="10">E</emphasis>
, lower margin and inferior angle of mandible; 
<emphasis id="42A8775CFFEBFF97FC40F8EEFC01EA4F" bold="true" box="[970,985,1869,1893]" pageId="10">F</emphasis>
, inferior margin of the mandible; 
<emphasis id="42A8775CFFEBFF97FAA4F8EFFA9AEA4F" bold="true" box="[1326,1346,1868,1893]" pageId="10">G</emphasis>
, labrum; 
<emphasis id="42A8775CFFEBFF97FF1DF8D2FF73EAA3" bold="true" box="[151,171,1905,1929]" pageId="10">H</emphasis>
, teeth on the labrum. Scale bars: A, B, D, G = 100µm, C, E, F, H = 50µm.
</paragraph>
</caption>
<caption id="24A3FBC6FFEAFF96FF1DF8D4FEF3EAD6" ID-DOI="http://doi.org/10.5281/zenodo.3530103" ID-Zenodo-Dep="3530103" httpUri="https://zenodo.org/record/3530103/files/figure.png" pageId="11" startId="11.[151,250,1911,1936]" targetBox="[177,1411,181,1885]" targetPageId="11">
<paragraph id="7063AB4EFFEAFF96FF1DF8D4FEF3EAD6" blockId="11.[151,1437,1911,2044]" pageId="11">
<emphasis id="42A8775CFFEAFF96FF1DF8D4FB83EABA" bold="true" box="[151,1115,1911,1936]" pageId="11">
FIGURE 8. Cirri of 
<taxonomicName id="B7DCD0CDFFEAFF96FEF1F8DBFD87EAA5" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[379,607,1912,1935]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="11" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFEAFF96FEF1F8DBFD87EAA5" bold="true" box="[379,607,1912,1935]" italics="true" pageId="11">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFEAFF96FDE2F8DBFD62EABA" box="[616,698,1912,1936]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
(MNHN-IU-2014-23572: holotype): A
</emphasis>
, cirrus I; 
<emphasis id="42A8775CFFEAFF96FB4FF8DBFB0EEABA" bold="true" box="[1221,1238,1912,1936]" pageId="11">B</emphasis>
, 
<emphasis id="42A8775CFFEAFF96FB6EF8D4FB2EEABA" bold="true" box="[1252,1270,1911,1936]" pageId="11">C</emphasis>
, distal serrulate setae on posterior and anterior rami of cirrus I; 
<emphasis id="42A8775CFFEAFF96FDF6F83FFD56EA9E" bold="true" box="[636,654,1948,1972]" pageId="11">D</emphasis>
, cirrus II; 
<emphasis id="42A8775CFFEAFF96FD71F83FFCD4EA9E" bold="true" box="[763,780,1948,1972]" pageId="11">E</emphasis>
, distal serrulate setae on posterior and anterior rami of cirrus II; 
<emphasis id="42A8775CFFEAFF96FF1DF863FF7EEAF2" bold="true" box="[151,166,1984,2008]" pageId="11">F</emphasis>
, cirrus III; 
<emphasis id="42A8775CFFEAFF96FE97F81CFEE9EAF2" bold="true" box="[285,305,1983,2008]" pageId="11">G</emphasis>
. magnified view of cirrus II; 
<emphasis id="42A8775CFFEAFF96FDEBF863FDADEAF2" bold="true" box="[609,629,1984,2008]" pageId="11">H</emphasis>
, 
<emphasis id="42A8775CFFEAFF96FD08F863FD4AEAF2" bold="true" box="[642,658,1984,2008]" pageId="11">I,</emphasis>
distal serrulate setae on posterior and anterior rami of cirrus III. Scale bars: A−I = 100µm.
</paragraph>
</caption>
<caption id="24A3FBC6FFEDFF91FF1DF839FAF9EAD1" ID-DOI="http://doi.org/10.5281/zenodo.3530105" ID-Zenodo-Dep="3530105" httpUri="https://zenodo.org/record/3530105/files/figure.png" pageId="12" startId="12.[151,250,1946,1971]" targetBox="[157,1431,181,1920]" targetPageId="12">
<paragraph id="7063AB4EFFEDFF91FF1DF839FAF9EAD1" blockId="12.[151,1436,1946,2043]" pageId="12">
<emphasis id="42A8775CFFEDFF91FF1DF839FB8DEA99" bold="true" box="[151,1109,1946,1971]" pageId="12">
FIGURE 9. Cirri of 
<taxonomicName id="B7DCD0CDFFEDFF91FEF2F838FD83EA98" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[376,603,1947,1970]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="12" phylum="Arthropoda" rank="species" species="spurtuncata" status="sp. nov.">
<emphasis id="42A8775CFFEDFF91FEF2F838FD83EA98" bold="true" box="[376,603,1947,1970]" italics="true" pageId="12">Pyrgoma spurtuncata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="599BCA27FFEDFF91FDEEF839FD6EEA98" box="[612,694,1946,1970]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
(MNHN-IU-2014-23572: holotype): A
</emphasis>
, cirrus IV; 
<emphasis id="42A8775CFFEDFF91FB46F839FB05EA98" bold="true" box="[1228,1245,1946,1970]" pageId="12">B</emphasis>
, intermediate segments of cirrus IV; 
<emphasis id="42A8775CFFEDFF91FEEBF81DFEA2EAFD" bold="true" box="[353,378,1982,2007]" pageId="12">C,</emphasis>
cirrus V; 
<emphasis id="42A8775CFFEDFF91FE6BF81DFE2BEAFC" bold="true" box="[481,499,1982,2006]" pageId="12">D</emphasis>
, intermediate segments of cirrus V; 
<emphasis id="42A8775CFFEDFF91FCE3F81DFCA2EAFC" bold="true" box="[873,890,1982,2006]" pageId="12">E</emphasis>
, cirrus VI; 
<emphasis id="42A8775CFFEDFF91FC78F81DFBD9EAFC" bold="true" box="[1010,1025,1982,2006]" pageId="12">F</emphasis>
, intermediate segments of cirrus VI; 
<emphasis id="42A8775CFFEDFF91FA08F81DFA4EEAFD" bold="true" box="[1410,1430,1982,2007]" pageId="12">G</emphasis>
, penis; 
<emphasis id="42A8775CFFEDFF91FF50F841FF36EAD0" bold="true" box="[218,238,2018,2042]" pageId="12">H</emphasis>
, basidorsal point of penis; 
<emphasis id="42A8775CFFEDFF91FD8AF841FDD2EAD0" bold="true" box="[512,522,2018,2042]" pageId="12">I</emphasis>
, tip of penis. Scale bars: A, C, E, G = 200µm, B, D, F, H = 100µm, I = 50µm.
</paragraph>
</caption>
<paragraph id="7063AB4EFFECFF90FF4DFF3AFC42ECDD" blockId="13.[151,1437,152,503]" pageId="13">
Most of the coral-inhabiting barnacles of the family 
<taxonomicName id="B7DCD0CDFFECFF90FC80FF3AFBFEED99" authority="Gray, 1825" authorityName="Gray" authorityYear="1825" box="[778,1062,152,179]" class="Maxillopoda" family="Pyrgomatidae" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="family">
Pyrgomatidae 
<bibRefCitation id="144DD6BFFFECFF90FC21FF3AFBFEED99" author="Gray, J. E." box="[939,1062,152,179]" pageId="13" pagination="97 - 107" refId="ref7982" refString="Gray, J. E. (1825) A synopsis of the genera of Cirripedes arranged in natural families, with a description of some new species. Annals of Philosophy, 10, 97 - 107." type="journal article" year="1825">Gray, 1825</bibRefCitation>
</taxonomicName>
(e.g., 
<taxonomicName id="B7DCD0CDFFECFF90FBE1FF3AFB06ED99" authorityName="Ross &amp; Newman" authorityYear="1973" box="[1131,1246,153,179]" class="Maxillopoda" family="Pyrgomatidae" genus="Cantellius" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFECFF90FBE1FF3AFB06ED99" box="[1131,1246,153,179]" italics="true" pageId="13">Cantellius</emphasis>
</taxonomicName>
, 
<taxonomicName id="B7DCD0CDFFECFF90FB61FF3AFA80ED99" authorityName="Perreault" authorityYear="2014" box="[1259,1368,153,179]" class="Maxillopoda" family="Pyrgomatidae" genus="Galkinius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFECFF90FB61FF3AFA80ED99" box="[1259,1368,153,179]" italics="true" pageId="13">Galkinius</emphasis>
</taxonomicName>
) have a solid basis. In 
<taxonomicName id="B7DCD0CDFFECFF90FEC7FF1DFE6BEDFD" authorityName=", Ross and Newman" authorityYear="2002" box="[333,435,190,215]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFECFF90FEC7FF1DFE6BEDFD" box="[333,435,190,215]" italics="true" pageId="13">Pyrgoma</emphasis>
</taxonomicName>
, the basis has perforations or arches that are believed to provide chemical mediations between the coral hosts and the barnacles (
<bibRefCitation id="144DD6BFFFECFF90FDD7FF42FCB2EDD1" author="Ross, A. &amp; Newman, W. A." box="[605,874,225,251]" pageId="13" pagination="245 - 260" refId="ref8813" refString="Ross, A. &amp; Newman, W. A. (2000) Pyrgoma kuri Hoek, 1913: A case study in morphology and systematics of a symbiotic coral barnacle (Cirripedia: Balanomorpha). Contributions to Zoology, 68 (4), 245 - 260. https: // doi. org / 10.1163 / 18759866 - 06804002" type="journal article" year="2000">Ross &amp; Newman, 2000</bibRefCitation>
). 
<bibRefCitation id="144DD6BFFFECFF90FCF6FF42FB79EDD1" author="Ross, A. &amp; Newman, W. A." box="[892,1185,225,251]" pageId="13" pagination="245 - 260" refId="ref8813" refString="Ross, A. &amp; Newman, W. A. (2000) Pyrgoma kuri Hoek, 1913: A case study in morphology and systematics of a symbiotic coral barnacle (Cirripedia: Balanomorpha). Contributions to Zoology, 68 (4), 245 - 260. https: // doi. org / 10.1163 / 18759866 - 06804002" type="journal article" year="2000">Ross and Newman (2000)</bibRefCitation>
examined the basis of 
<taxonomicName id="B7DCD0CDFFECFF90FF1DFEA5FF3BEC35" authorityName="Hoek" authorityYear="1913" box="[151,227,261,287]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="kuri">
<emphasis id="42A8775CFFECFF90FF1DFEA5FF70EC35" box="[151,168,262,287]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FF3CFEA6FF3BEC35" box="[182,227,261,287]" italics="true" pageId="13">kuri</emphasis>
</taxonomicName>
and found that the initial stage is ladder-like perforations, arranged longitudinally, that turn into arches when approaching the top of the shells. Additionally, the basis of 
<taxonomicName id="B7DCD0CDFFECFF90FCA6FE89FC0CEC69" authorityName="Leach" authorityYear="1818" box="[812,980,297,323]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFECFF90FCA6FE89FCE5EC69" box="[812,829,298,323]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FCC1FE8AFC0CEC69" box="[843,980,297,323]" italics="true" pageId="13">cancellatum</emphasis>
</taxonomicName>
becomes arch-like as the basis develops. Conversely, 
<taxonomicName id="B7DCD0CDFFECFF90FEA8FEEDFE15EC4D" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[290,461,334,359]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFECFF90FEA8FEEDFE15EC4D" box="[290,461,334,359]" italics="true" pageId="13">P. spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFECFF90FE59FEEEFDF3EC4D" bold="true" box="[467,555,333,359]" pageId="13">
<taxonomicNameLabel id="599BCA27FFECFF90FE59FEEEFDF3EC4D" box="[467,555,333,359]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows some variations in the basis. The top of the basis (junction between shell and basis) in the 
<typeStatus id="AF6715ECFFECFF90FEDAFED2FE6AECA1" box="[336,434,369,395]" pageId="13" type="holotype">holotype</typeStatus>
does not contain extensive longitudinal perforations and is mostly solid in nature (
<figureCitation id="E8E7B7CBFFECFF90FACAFED2FF1FEC85" captionStart="FIGURE 4" captionStartId="6.[151,250,1459,1484]" captionTargetBox="[164,1423,181,1434]" captionTargetId="figure@6.[164,1423,181,1434]" captionTargetPageId="6" captionText="FIGURE 4. Base view of Pyrgoma spurtuncata sp. nov.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A, top view of base without longitudinal ladder-like perforations; B, top view of base with short sections of ladder-like perforations; C, D, top view of base with some ladder-like perforations; E, F, enlarged view of basis; G, H, enlarged view of basis; I, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm" figureDoi="http://doi.org/10.5281/zenodo.3530095" httpUri="https://zenodo.org/record/3530095/files/figure.png" pageId="13">Fig. 4A, E, F</figureCitation>
) but, in some specimens, there are some ladder-like, longitudinal perforations extending from the deeper to the upper part of the basis (
<figureCitation id="E8E7B7CBFFECFF90FE11FE1AFE3FECF9" box="[411,487,441,467]" captionStart="FIGURE 4" captionStartId="6.[151,250,1459,1484]" captionTargetBox="[164,1423,181,1434]" captionTargetId="figure@6.[164,1423,181,1434]" captionTargetPageId="6" captionText="FIGURE 4. Base view of Pyrgoma spurtuncata sp. nov.. A, E and F are specimen MNHN-IU-2014-23572 (Holotype); B, G and H are specimen MNHN-IU-2014-23573 (Paratype); C, D, and I are specimen MNHN-IU-2014-5597 (paratype): A, top view of base without longitudinal ladder-like perforations; B, top view of base with short sections of ladder-like perforations; C, D, top view of base with some ladder-like perforations; E, F, enlarged view of basis; G, H, enlarged view of basis; I, enlarged view of basis showing the perforations extending from deeper to upper part. Scale bars: A, B, D = 1mm, E, G, H, I = 500µm, F = 200µm" figureDoi="http://doi.org/10.5281/zenodo.3530095" httpUri="https://zenodo.org/record/3530095/files/figure.png" pageId="13">Figs. 4</figureCitation>
B-D, 4G-I, 5). However, no arches were seen on the joint with the basis. It is probable that 
<taxonomicName id="B7DCD0CDFFECFF90FF42FE7DFEADECDD" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[200,373,478,503]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFECFF90FF42FE7DFEADECDD" box="[200,373,478,503]" italics="true" pageId="13">P. spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFECFF90FEF1FE7EFE0DECDD" bold="true" box="[379,469,477,503]" pageId="13">
<taxonomicNameLabel id="599BCA27FFECFF90FEF1FE7EFE0DECDD" box="[379,469,477,503]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does not develop arch-like perforations.
</paragraph>
<caption id="24A3FBC6FFECFF90FF1DF88BFE2EEADF" ID-DOI="http://doi.org/10.5281/zenodo.3530107" ID-Zenodo-Dep="3530107" httpUri="https://zenodo.org/record/3530107/files/figure.png" pageId="13" startId="13.[151,250,1832,1857]" targetBox="[177,1411,525,1807]" targetPageId="13">
<paragraph id="7063AB4EFFECFF90FF1DF88BFE2EEADF" blockId="13.[151,1436,1832,2037]" pageId="13">
<emphasis id="42A8775CFFECFF90FF1DF88BFAB6EA6B" bold="true" box="[151,1390,1832,1857]" pageId="13">
FIGURE 10. Scanning electron micrographs and re-drawings from previous published illustrations of 
<taxonomicName id="B7DCD0CDFFECFF90FB7AF88AFA96EA6A" authorityName=", Ross and Newman" authorityYear="2002" box="[1264,1358,1833,1856]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFECFF90FB7AF88AFA96EA6A" bold="true" box="[1264,1358,1833,1856]" italics="true" pageId="13">Pyrgoma</emphasis>
</taxonomicName>
: A
</emphasis>
, top and basal view of 
<taxonomicName id="B7DCD0CDFFECFF90FED8F8EEFE32EA4E" authorityName="Leach" authorityYear="1818" box="[338,490,1868,1892]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFECFF90FED8F8EEFEB9EA4E" box="[338,353,1869,1892]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FEE4F8EFFE32EA4E" box="[366,490,1868,1892]" italics="true" pageId="13">cancellatum</emphasis>
</taxonomicName>
(Taiwan, 
<bibRefCitation id="144DD6BFFFECFF90FDDBF8EFFCC9EA4F" author="Chan, B. K. K. &amp; Chen, Y. &amp; Achituv Y." box="[593,785,1868,1893]" pageId="13" refId="ref7649" refString="Chan, B. K. K., Chen, Y. Y &amp; Achituv Y. (2013 b) Crustacean Fauna of Taiwan: Barnacles, Volume II - Cirripedia: Thoracica: Pyrgomatidae. Academia Sinica Press, Taipei, 364 pp." type="book" year="2013">
Chan 
<emphasis id="42A8775CFFECFF90FD07F8EEFD62EA4E" box="[653,698,1868,1892]" italics="true" pageId="13">et al</emphasis>
., 2013b
</bibRefCitation>
); 
<emphasis id="42A8775CFFECFF90FCA9F8EEFCECEA4F" bold="true" box="[803,820,1869,1893]" pageId="13">B</emphasis>
, scutum and tergum of 
<taxonomicName id="B7DCD0CDFFECFF90FBADF8EEFB18EA4E" authorityName="Leach" authorityYear="1818" box="[1063,1216,1868,1892]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFECFF90FBADF8EEFBEEEA4E" box="[1063,1078,1869,1892]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FBCEF8EFFB18EA4E" box="[1092,1216,1868,1892]" italics="true" pageId="13">cancellatum</emphasis>
</taxonomicName>
(Taiwan, 
<bibRefCitation id="144DD6BFFFECFF90FAACF8EFFF02EAA3" author="Chan, B. K. K. &amp; Chen, Y. &amp; Achituv Y." pageId="13" refId="ref7649" refString="Chan, B. K. K., Chen, Y. Y &amp; Achituv Y. (2013 b) Crustacean Fauna of Taiwan: Barnacles, Volume II - Cirripedia: Thoracica: Pyrgomatidae. Academia Sinica Press, Taipei, 364 pp." type="book" year="2013">
Chan 
<emphasis id="42A8775CFFECFF90FAE9F8EEFA48EA4E" box="[1379,1424,1868,1892]" italics="true" pageId="13">et al</emphasis>
., 2013b
</bibRefCitation>
); 
<emphasis id="42A8775CFFECFF90FF67F8D3FF27EAA3" bold="true" box="[237,255,1904,1929]" pageId="13">C</emphasis>
, top and basal view of 
<taxonomicName id="B7DCD0CDFFECFF90FE78F8D2FDB0EAA2" authorityName="Weltner" authorityYear="1897" box="[498,616,1905,1928]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFECFF90FE78F8D2FDD9EAA2" box="[498,513,1905,1928]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FD85F8D2FDB0EAA2" box="[527,616,1905,1928]" italics="true" pageId="13">japonica</emphasis>
</taxonomicName>
(Taiwan, present study); 
<emphasis id="42A8775CFFECFF90FCE4F8D2FC58EAA3" bold="true" box="[878,896,1905,1929]" pageId="13">D</emphasis>
, scutum and tergum of 
<taxonomicName id="B7DCD0CDFFECFF90FBFFF8D2FB34EAA2" authorityName="Weltner" authorityYear="1897" box="[1141,1260,1905,1928]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFECFF90FBFFF8D2FB5CEAA2" box="[1141,1156,1905,1928]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FB19F8D2FB34EAA2" box="[1171,1260,1905,1928]" italics="true" pageId="13">japonica</emphasis>
</taxonomicName>
(Taiwan, present study); 
<emphasis id="42A8775CFFECFF90FF6EF836FF2DEA87" bold="true" box="[228,245,1941,1965]" pageId="13">E</emphasis>
. redrawing of opercular plates of 
<taxonomicName id="B7DCD0CDFFECFF90FDDBF836FD31EA86" authorityName="Leach" authorityYear="1818" box="[593,745,1940,1964]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFECFF90FDDBF836FDB8EA86" box="[593,608,1941,1964]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FDE7F837FD31EA86" box="[621,745,1940,1964]" italics="true" pageId="13">cancellatum</emphasis>
</taxonomicName>
from 
<bibRefCitation id="144DD6BFFFECFF90FCA2F837FC7CEA87" author="Hiro, F." box="[808,932,1940,1965]" pageId="13" pagination="45 - 72" refId="ref8059" refString="Hiro, F. (1935) A study of cirripeds associated with corals occurring in Tanabe Bay. Records of Oceanographic Works in Japan, 7, 45 - 72." type="journal article" year="1935">Hiro (1935)</bibRefCitation>
; 
<emphasis id="42A8775CFFECFF90FC25F836FC66EA87" bold="true" box="[943,958,1941,1965]" pageId="13">F</emphasis>
, redrawing of opercular plates of 
<taxonomicName id="B7DCD0CDFFECFF90FA90F836FA44EA86" authorityName="Nilsson-Cantell" authorityYear="1938" box="[1306,1436,1941,1964]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="projectum">
<emphasis id="42A8775CFFECFF90FA90F836FAF1EA86" box="[1306,1321,1941,1964]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FABCF836FA44EA86" box="[1334,1436,1941,1964]" italics="true" pageId="13">projectum</emphasis>
</taxonomicName>
from 
<bibRefCitation id="144DD6BFFFECFF90FF5AF81BFE64EAFB" author="Nilsson-Cantell, C. A." box="[208,444,1976,2001]" pageId="13" pagination="1 - 81" refId="ref8469" refString="Nilsson-Cantell, C. A. (1938) Cirripedes from the Indian Ocean in the collection of the Indian Museum, Calcutta. Memoirs of the Indian Museum, 13, 1 - 81." type="journal article" year="1938">Nilsson-Cantell (1938)</bibRefCitation>
; 
<emphasis id="42A8775CFFECFF90FE4DF81BFE03EAFB" bold="true" box="[455,475,1976,2001]" pageId="13">G</emphasis>
, redrawing of opercular plates of 
<taxonomicName id="B7DCD0CDFFECFF90FCB2F81AFC4AEAFA" baseAuthorityName="Ren" baseAuthorityYear="1986" box="[824,914,1977,2000]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="13" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="42A8775CFFECFF90FCB2F81AFC9FEAFA" box="[824,839,1977,2000]" italics="true" pageId="13">P</emphasis>
. 
<emphasis id="42A8775CFFECFF90FCDFF81AFC4AEAFA" box="[853,914,1977,2000]" italics="true" pageId="13">sinica</emphasis>
</taxonomicName>
from 
<bibRefCitation id="144DD6BFFFECFF90FC5BF81AFB9FEAFB" author="Ren, X." box="[977,1095,1976,2001]" pageId="13" pagination="129 - 158" refId="ref8606" refString="Ren, X. (1986) Studies on Chinese Cirripedia (Crustacea), VII. Family Pyrgomatidae, Studia Marina Sinica, 26, 129 - 158." type="journal article" year="1986">Ren (1986)</bibRefCitation>
. Scale bars: A = 2000µm, B, D = 500µm, C = 1mm, E, F, G = 1mm.
</paragraph>
</caption>
<paragraph id="7063AB4EFFEFFF93FF4DFF07FBE7EC00" blockId="14.[151,1437,163,730]" pageId="14">
<emphasis id="42A8775CFFEFFF93FF4DFF07FE76ED94" bold="true" box="[199,430,163,190]" pageId="14">Molecular analysis.</emphasis>
A total of nine sequences of COI and 12s rDNA from nine specimens (including four 
<taxonomicName id="B7DCD0CDFFEFFF93FAE1FF07FEB7EDCB" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFEFFF93FAE1FF07FEB7EDCB" italics="true" pageId="14">Pyrgoma spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFEFFF93FEFDFF6BFE0AEDC8" bold="true" box="[375,466,200,226]" pageId="14">
<taxonomicNameLabel id="599BCA27FFEFFF93FEFDFF6BFE0AEDC8" box="[375,466,200,226]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, one 
<taxonomicName id="B7DCD0CDFFEFFF93FD98FF6BFD4CEDCB" authorityName="Weltner" authorityYear="1897" box="[530,660,200,226]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFEFFF93FD98FF6BFDFBEDCB" box="[530,547,200,225]" italics="true" pageId="14">P</emphasis>
. 
<emphasis id="42A8775CFFEFFF93FDB8FF6BFD4CEDCB" box="[562,660,200,225]" italics="true" pageId="14">japonica</emphasis>
</taxonomicName>
and four 
<taxonomicName id="B7DCD0CDFFEFFF93FC8EFF6BFC75EDCB" authorityName="Leach" authorityYear="1818" box="[772,941,199,226]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="cancellatum">
<emphasis id="42A8775CFFEFFF93FC8EFF6BFCCDEDCB" box="[772,789,200,225]" italics="true" pageId="14">P</emphasis>
. 
<emphasis id="42A8775CFFEFFF93FCAEFF64FC75EDCB" box="[804,941,199,225]" italics="true" pageId="14">cancellatum</emphasis>
</taxonomicName>
) were amplified and compared to 
<taxonomicName id="B7DCD0CDFFEFFF93FABDFF6BFA45EDCB" authorityName=", Ross and Newman" authorityYear="2002" box="[1335,1437,200,225]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFEFFF93FABDFF6BFA45EDCB" box="[1335,1437,200,225]" italics="true" pageId="14">Pyrgoma</emphasis>
</taxonomicName>
sequences retrieved from GenBank. The sequences from the present study are deposited in GenBank (
<tableCitation id="3D5E9EF5FFEFFF93FA8CFF48FA81EC2C" box="[1286,1369,235,262]" captionStart="TABLE 1" captionStartId="2.[151,244,1376,1403]" captionTargetBox="[163,1390,1454,1973]" captionText="TABLE 1. Host corals and GenBank numbers for the sequences used in the molecular analysis of the species of Pyrgoma. Used symbol: N/A—not available." httpUri="http://table.plazi.org/id/24A3FBC6FFE3FF9FFF1DFAC3FD48E8B5" pageId="14" tableUuid="24A3FBC6FFE3FF9FFF1DFAC3FD48E8B5">Table 1</tableCitation>
). The concatenated alignment consisted of 452 nucleotides of 12S rDNA and 563 of COI.
</paragraph>
<paragraph id="7063AB4EFFEFFF93FF4DFE90FCE5ECF4" blockId="14.[151,1437,163,730]" pageId="14">
Molecular identification using both mitochondrial 12S rDNA and COI gene sequences complemented the morpholoical identifications. Sequence similarity (K2P distance, 
<tableCitation id="3D5E9EF5FFEFFF93FCDEFEF4FC73EC58" box="[852,939,343,370]" captionStart="TABLE 2" captionStartId="15.[151,244,697,724]" captionTargetBox="[162,1944,740,1509]" captionText="TABLE 2. Mean K2P distance for the concatenated molecular dataset for 12SrDNA+COI genes between and within the studied species." httpUri="http://table.plazi.org/id/24A3FBC6FFEEFF92FF1DFD1AF949EFFE" pageId="14" tableUuid="24A3FBC6FFEEFF92FF1DFD1AF949EFFE">Table 2</tableCitation>
) and the phylogenetic analyses (Maximum Likelihood, 
<figureCitation id="E8E7B7CBFFEFFF93FE95FED8FEB4ECBC" box="[287,364,379,406]" captionStart="FIGURE 11" captionStartId="14.[151,250,1947,1972]" captionTargetBox="[193,1394,756,1915]" captionTargetId="figure@14.[189,1398,751,1922]" captionTargetPageId="14" captionText="FIGURE 11. Phylogenetic trees for Pyrgoma specimens based on maximum likelihood analysis of mitochondrial 12S rDNA and COI gene: A, 12S rDNA; B, COI; C, combined data sets of two mitochondrial genes (12S rDNA+COI). Bootstrap scores of&gt; 80% are presented at nodes." figureDoi="http://doi.org/10.5281/zenodo.3530109" httpUri="https://zenodo.org/record/3530109/files/figure.png" pageId="14">Fig. 11</figureCitation>
) confirmed the identity of four 
<taxonomicName id="B7DCD0CDFFEFFF93FD40FEDFFCA1ECBF" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[714,889,380,406]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFEFFF93FD40FEDFFD03ECBF" box="[714,731,380,405]" italics="true" pageId="14">P</emphasis>
. 
<emphasis id="42A8775CFFEFFF93FD62FEDFFCA1ECBF" box="[744,889,380,405]" italics="true" pageId="14">spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFEFFF93FCF5FEDFFC00ECBC" bold="true" box="[895,984,380,406]" pageId="14">
<taxonomicNameLabel id="599BCA27FFEFFF93FCF5FEDFFC00ECBC" box="[895,984,380,406]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens; the phylograms based on the two markers showed high similarities (
<figureCitation id="E8E7B7CBFFEFFF93FDC6FE3CFD0BEC90" box="[588,723,415,442]" captionStart="FIGURE 11" captionStartId="14.[151,250,1947,1972]" captionTargetBox="[193,1394,756,1915]" captionTargetId="figure@14.[189,1398,751,1922]" captionTargetPageId="14" captionText="FIGURE 11. Phylogenetic trees for Pyrgoma specimens based on maximum likelihood analysis of mitochondrial 12S rDNA and COI gene: A, 12S rDNA; B, COI; C, combined data sets of two mitochondrial genes (12S rDNA+COI). Bootstrap scores of&gt; 80% are presented at nodes." figureDoi="http://doi.org/10.5281/zenodo.3530109" httpUri="https://zenodo.org/record/3530109/files/figure.png" pageId="14">Fig. 11A, B</figureCitation>
). The phylogeny of the concatenated data set was similar to the phylogenetic trees derived from individual genes (
<figureCitation id="E8E7B7CBFFEFFF93FD41FE60FCE8ECF4" box="[715,816,451,478]" captionStart="FIGURE 11" captionStartId="14.[151,250,1947,1972]" captionTargetBox="[193,1394,756,1915]" captionTargetId="figure@14.[189,1398,751,1922]" captionTargetPageId="14" captionText="FIGURE 11. Phylogenetic trees for Pyrgoma specimens based on maximum likelihood analysis of mitochondrial 12S rDNA and COI gene: A, 12S rDNA; B, COI; C, combined data sets of two mitochondrial genes (12S rDNA+COI). Bootstrap scores of&gt; 80% are presented at nodes." figureDoi="http://doi.org/10.5281/zenodo.3530109" httpUri="https://zenodo.org/record/3530109/files/figure.png" pageId="14">Fig. 11C</figureCitation>
).
</paragraph>
<paragraph id="7063AB4EFFEFFF93FF4DFE4BFC48EFF3" blockId="14.[151,1437,163,730]" pageId="14">
In pairwise comparisons, the K2P distance among the sequences ranged from 0.1% to 18.8% (
<tableCitation id="3D5E9EF5FFEFFF93FA8BFE44FA83EF28" box="[1281,1371,487,514]" captionStart="TABLE 2" captionStartId="15.[151,244,697,724]" captionTargetBox="[162,1944,740,1509]" captionText="TABLE 2. Mean K2P distance for the concatenated molecular dataset for 12SrDNA+COI genes between and within the studied species." httpUri="http://table.plazi.org/id/24A3FBC6FFEEFF92FF1DFD1AF949EFFE" pageId="14" tableUuid="24A3FBC6FFEEFF92FF1DFD1AF949EFFE">Table 2</tableCitation>
). The mean intraspecific divergence of 
<taxonomicName id="B7DCD0CDFFEFFF93FD85FDAFFD18EF0F" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[527,704,524,550]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFEFFF93FD85FDAFFDF8EF0F" box="[527,544,524,549]" italics="true" pageId="14">P</emphasis>
. 
<emphasis id="42A8775CFFEFFF93FDA5FDAFFD18EF0F" box="[559,704,524,549]" italics="true" pageId="14">spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFEFFF93FD4DFDAFFCFAEF0C" bold="true" box="[711,802,524,550]" pageId="14">
<taxonomicNameLabel id="599BCA27FFEFFF93FD4DFDAFFCFAEF0C" box="[711,802,524,550]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was only 0.1%. In contrast, the interspecific divergence (based on different species name from GenBank) of other 
<taxonomicName id="B7DCD0CDFFEFFF93FCA3FD93FC57EF63" authorityName=", Ross and Newman" authorityYear="2002" box="[809,911,560,585]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFEFFF93FCA3FD93FC57EF63" box="[809,911,560,585]" italics="true" pageId="14">Pyrgoma</emphasis>
</taxonomicName>
species ranged from 2.4 % to 15.6%. The low divergence value of 2.4% is between 
<taxonomicName id="B7DCD0CDFFEFFF93FDBEFDF7FD42EF47" authorityName=", Ross and Newman" authorityYear="2002" box="[564,666,596,621]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFEFFF93FDBEFDF7FD42EF47" box="[564,666,596,621]" italics="true" pageId="14">Pyrgoma</emphasis>
</taxonomicName>
sp. 1 (
<bibRefCitation id="144DD6BFFFEFFF93FD6BFDF0FBF8EF44" author="Malay, M. C. D. &amp; Michonneau, F." box="[737,1056,595,622]" pageId="14" pagination="162 - 179" refId="ref8427" refString="Malay, M. C. D. &amp; Michonneau, F. (2014) Phylogenetics and morphological evolution of coral-dwelling barnacles (Balanomorpha: Pyrgomatidae). Biological Journal of the Linnaean Society, 113, 162 - 179." type="journal article" year="2014">Malay &amp; Michonneau, 2014</bibRefCitation>
) and 
<taxonomicName id="B7DCD0CDFFEFFF93FBD1FDF7FB00EF47" authorityName="Weltner" authorityYear="1897" box="[1115,1240,596,621]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFEFFF93FBD1FDF7FB00EF47" box="[1115,1240,596,621]" italics="true" pageId="14">P. japonica</emphasis>
</taxonomicName>
, which located in the same clade in the molecular phylogenetic tree (
<figureCitation id="E8E7B7CBFFEFFF93FD43FDD4FCC1EFB8" box="[713,793,631,658]" captionStart="FIGURE 11" captionStartId="14.[151,250,1947,1972]" captionTargetBox="[193,1394,756,1915]" captionTargetId="figure@14.[189,1398,751,1922]" captionTargetPageId="14" captionText="FIGURE 11. Phylogenetic trees for Pyrgoma specimens based on maximum likelihood analysis of mitochondrial 12S rDNA and COI gene: A, 12S rDNA; B, COI; C, combined data sets of two mitochondrial genes (12S rDNA+COI). Bootstrap scores of&gt; 80% are presented at nodes." figureDoi="http://doi.org/10.5281/zenodo.3530109" httpUri="https://zenodo.org/record/3530109/files/figure.png" pageId="14">Fig. 11</figureCitation>
), suggesting 
<taxonomicName id="B7DCD0CDFFEFFF93FC21FDDBFBC9EFBB" authorityName=", Ross and Newman" authorityYear="2002" box="[939,1041,632,657]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFEFFF93FC21FDDBFBC9EFBB" box="[939,1041,632,657]" italics="true" pageId="14">Pyrgoma</emphasis>
</taxonomicName>
sp. 1 from 
<bibRefCitation id="144DD6BFFFEFFF93FB07FDD4FF17EF9C" author="Malay, M. C. D. &amp; Michonneau, F." pageId="14" pagination="162 - 179" refId="ref8427" refString="Malay, M. C. D. &amp; Michonneau, F. (2014) Phylogenetics and morphological evolution of coral-dwelling barnacles (Balanomorpha: Pyrgomatidae). Biological Journal of the Linnaean Society, 113, 162 - 179." type="journal article" year="2014">Malay and Michonneau, 2014</bibRefCitation>
is 
<taxonomicName id="B7DCD0CDFFEFFF93FF65FD3FFEB3EF9F" authorityName="Weltner" authorityYear="1897" box="[239,363,668,693]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFEFFF93FF65FD3FFEB3EF9F" box="[239,363,668,693]" italics="true" pageId="14">P. japonica</emphasis>
</taxonomicName>
. Among the 
<taxonomicName id="B7DCD0CDFFEFFF93FE70FD3FFDB8EF9F" authorityName=", Ross and Newman" authorityYear="2002" box="[506,608,668,693]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="genus">
<emphasis id="42A8775CFFEFFF93FE70FD3FFDB8EF9F" box="[506,608,668,693]" italics="true" pageId="14">Pyrgoma</emphasis>
</taxonomicName>
species, 
<taxonomicName id="B7DCD0CDFFEFFF93FD4EFD3FFCACEF9F" authority="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim, 2019" authorityName="Kim &amp; Chan &amp; Corbari &amp; Rodriguez Moreno &amp; Achituv &amp; Kim" authorityYear="2019" box="[708,884,668,694]" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="spurtruncata" status="sp. nov.">
<emphasis id="42A8775CFFEFFF93FD4EFD3FFD0DEF9F" box="[708,725,668,693]" italics="true" pageId="14">P</emphasis>
. 
<emphasis id="42A8775CFFEFFF93FD69FD3FFCACEF9F" box="[739,884,668,693]" italics="true" pageId="14">spurtruncata</emphasis>
</taxonomicName>
<emphasis id="42A8775CFFEFFF93FCF0FD3FFC0BEF9C" bold="true" box="[890,979,668,694]" pageId="14">
<taxonomicNameLabel id="599BCA27FFEFFF93FCF0FD3FFC0BEF9C" box="[890,979,668,694]" pageId="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shows a sister group relationship with 
<taxonomicName id="B7DCD0CDFFEFFF93FA0EFD3FFF21EFF3" authorityName="Weltner" authorityYear="1897" class="Maxillopoda" family="Pyrgomatidae" genus="Pyrgoma" kingdom="Animalia" order="Sessilia" pageId="14" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="42A8775CFFEFFF93FA0EFD3FFA4DEF9F" box="[1412,1429,668,693]" italics="true" pageId="14">P</emphasis>
. 
<emphasis id="42A8775CFFEFFF93FF1DFD63FF21EFF3" box="[151,249,704,729]" italics="true" pageId="14">japonica</emphasis>
</taxonomicName>
and the pairwise distance was 11.9% between two species.
</paragraph>
</subSubSection>
</treatment>
</document>