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<document id="FC3F5656A4B6CB39AB5E1F86DA9C6B9E" ID-DOI="10.5281/zenodo.208677" ID-GBIF-Dataset="75cbad61-d248-413c-9575-cf9de777319f" ID-ISSN="1175-5326" ID-Zenodo-Dep="208677" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460559972897" checkinUser="plazi" docAuthor="Nielsen, Jørgen G., Schwarzhans, Werner &amp; Cohen, Daniel M." docDate="2012" docId="EB64193EE23EA34286A8F977FB4208B0" docLanguage="en" docName="zt03579p036.pdf" docOrigin="Zootaxa 3579" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Saccogaster brayae Nielsen, Schwarzhans &amp; Cohen, 2012, n. sp." docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="175D6146E238A34A863FFE79FF8C0A7C" masterDocTitle="Revision of Hastatobythites and Saccogaster (Teleostei, Bythitidae) with three new species and a new genus" masterLastPageNumber="36" masterPageNumber="1" pageNumber="7" updateTime="1698320134232" updateUser="plazi">
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<mods:title id="3DC7A1B8ECBF8439E80C11D6FCBDAE97">Revision of Hastatobythites and Saccogaster (Teleostei, Bythitidae) with three new species and a new genus</mods:title>
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<mods:namePart id="572D4DB589661CEE1CA59D9823A02CA8">Nielsen, Jørgen G.</mods:namePart>
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<mods:namePart id="4D88248C9D2B7846A9E48CA35976D2F3">Schwarzhans, Werner</mods:namePart>
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<mods:namePart id="D57515C5D099C2D004DCD89A49F6BA62">Cohen, Daniel M.</mods:namePart>
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<mods:date id="4B2972D2BA0217B6E103CCBED8ED249C">2012</mods:date>
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<treatment id="EB64193EE23EA34286A8F977FB4208B0" ID-DOI="http://doi.org/10.5281/zenodo.6175246" ID-GBIF-Taxon="119556537" ID-Zenodo-Dep="6175246" LSID="urn:lsid:plazi:treatment:EB64193EE23EA34286A8F977FB4208B0" httpUri="http://treatment.plazi.org/id/EB64193EE23EA34286A8F977FB4208B0" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="2BD7FBA3E23EA34C86A8F977FE4E0D35" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="6372A828E23EA34C86A8F977FE5D0D54" blockId="6.[151,465,1806,1865]" box="[151,465,1806,1832]" pageId="6" pageNumber="7">
<heading id="383A1F44E23EA34C86A8F977FE5D0D54" bold="true" box="[151,465,1806,1832]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="51B9743AE23EA34C86A8F977FE5D0D54" bold="true" box="[151,465,1806,1832]" pageId="6" pageNumber="7">
<taxonomicName id="A4CDD3ABE23EA34C86A8F977FE0C0D54" ID-CoL="4TX2K" box="[151,384,1806,1832]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="brayae" status="sp. nov.">
<emphasis id="51B9743AE23EA34C86A8F977FE0C0D54" bold="true" box="[151,384,1806,1832]" italics="true" pageId="6" pageNumber="7">Saccogaster brayae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A8AC941E23EA34C87B0F977FE5D0D54" box="[399,465,1806,1832]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6372A828E23EA34C86A8F948FE4E0D35" blockId="6.[151,465,1806,1865]" box="[151,450,1841,1865]" pageId="6" pageNumber="7">
<figureCitation id="FBF6B4ADE23EA34C86A8F948FF680D35" box="[151,228,1841,1865]" captionStart="FIGURE 3" captionStartId="7.[151,250,681,704]" captionTargetBox="[165,1410,205,622]" captionTargetId="figure@7.[151,1436,183,660]" captionTargetPageId="7" captionText="FIGURE 3. Saccogaster brayae, holotype, NMV A 29655 - 002, SL 36 mm (photo M. Krag)." httpUri="https://zenodo.org/record/208680/files/figure.png" pageId="6" pageNumber="7">Figs. 3</figureCitation>
,
<figureCitation id="FBF6B4ADE23EA34C86CEF948FE8D0D35" box="[241,257,1841,1865]" captionStart="FIGURE 4" captionStartId="8.[151,250,1832,1855]" captionTargetBox="[168,1428,809,1785]" captionTargetId="figure@8.[151,1436,774,1806]" captionTargetPageId="8" captionText="FIGURE 4. Saccogaster brayae, holotype: A—Lateral view of head. B—Lateral radiograph of head. C—Dorsal view of head. D— Anterior right gill arch. E—Median view of right otolith. F—Dorsal view of right otolith. G—Intromittent organ." httpUri="https://zenodo.org/record/208681/files/figure.png" pageId="6" pageNumber="7">4</figureCitation>
,
<figureCitation id="FBF6B4ADE23EA34C8732F948FEA70D35" box="[269,299,1841,1865]" captionStart="FIGURE 25" captionStartId="32.[151,250,1767,1790]" captionTargetBox="[161,1424,1103,1720]" captionTargetId="figure@32.[151,1436,1073,1753]" captionTargetPageId="32" captionText="FIGURE 25. Geographic distribution of Hastatobythites and Saccogaster in the Indo-West Pacific. Each marking may represent more than one locality." httpUri="https://zenodo.org/record/208702/files/figure.png" pageId="6" pageNumber="7">25</figureCitation>
,
<tableCitation id="2E4F9D93E23EA34C8708F948FE060D35" box="[311,394,1841,1865]" captionStart="TABLE 1" captionStartId="2.[151,239,1157,1181]" captionTargetBox="[165,1351,1217,2019]" captionTargetPageId="2" captionText="TABLE 1. Meristic characters of Hastatobythites, Parasaccogaster n. gen. and Saccogaster" httpUri="http://table.plazi.org/id/37B2F8A0E23AA34886A8FAFCFBEA0EE1" pageId="6" pageNumber="7" tableUuid="37B2F8A0E23AA34886A8FAFCFBEA0EE1">Table 1</tableCitation>
,
<tableCitation id="2E4F9D93E23EA34C87A7F948FE240D35" box="[408,424,1841,1865]" captionStart="TABLE 2" captionStartId="3.[151,239,151,175]" captionText="TABLE 2. Meristic characters of Hastatobythites, Parasaccogaster n. gen. and Saccogaster." pageId="6" pageNumber="7">2</tableCitation>
,
<tableCitation id="2E4F9D93E23EA34C878BF948FE4E0D35" box="[436,450,1841,1865]" captionStart="TABLE 4" captionStartId="9.[151,239,151,175]" captionTargetBox="[165,1386,208,1519]" captionTargetPageId="9" captionText="TABLE 4. Meristic and morphometric characters of Saccogaster brayi, S. hawaii and S. horrida." httpUri="http://table.plazi.org/id/37B2F8A0E231A34386A8FEEEFB280AD3" pageId="6" pageNumber="7" tableUuid="37B2F8A0E231A34386A8FEEEFB280AD3">4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="2BD7FBA3E23EA34C86A8F900FD0C0DA5" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="6372A828E23EA34C86A8F900FD0C0DA5" blockId="6.[151,1436,1913,2009]" pageId="6" pageNumber="7">
<emphasis id="51B9743AE23EA34C86A8F900FEF60DEE" bold="true" box="[151,378,1913,1938]" pageId="6" pageNumber="7">Material examined</emphasis>
(
<specimenCount id="75CB63A1E23EA34C87B2F900FD820DED" box="[397,526,1913,1937]" pageId="6" pageNumber="7" type="generic">1 specimen</specimenCount>
,
<quantity id="A43505CDE23EA34C8425F900FDE70DED" box="[538,619,1913,1937]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="7" unit="mm" value="36.0">36 mm</quantity>
, SL).
<typeStatus id="BC76168AE23EA34C848EF900FC920DED" box="[689,798,1913,1937]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
:
<collectionCode id="05DC30EDE23EA34C8517F903FCE50DED" LSID="urn:lsid:biocol.org:col:34978" box="[808,873,1914,1937]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="6" pageNumber="7">NMV</collectionCode>
A 29655-002, male,
<quantity id="A43505CDE23EA34C8264F900FB270DED" box="[1115,1195,1913,1937]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="7" unit="mm" value="36.0">36 mm</quantity>
SL, off northwestern
<collectingCountry id="1BDAE8B8E23EA34C86A8F9E4FE8D0DC9" box="[151,257,1949,1973]" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
, Barrow L1 transect,
<geoCoordinate id="06F9CEEFE23EA34C87C8F9E4FD0E0DC9" box="[503,642,1949,1973]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="-21.01361">21°0049S</geoCoordinate>
,
<geoCoordinate id="06F9CEEFE23EA34C84B0F9E4FCAA0DC9" box="[655,806,1949,1973]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="114.652504">114°3909E</geoCoordinate>
to
<geoCoordinate id="06F9CEEFE23EA34C8572F9E4FC540DC9" box="[845,984,1949,1973]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="-21.013056">21°0047S</geoCoordinate>
,
<geoCoordinate id="06F9CEEFE23EA34C85DAF9E4FBF20DC9" box="[997,1150,1949,1973]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="114.64833">114°3854E</geoCoordinate>
, R/V Southern Surveyor, Sherman sledge,
<quantity id="A43505CDE23EA34C8768F9B8FE540DA5" box="[343,472,1985,2009]" metricMagnitude="2" metricUnit="m" metricValue="2.54" metricValueMax="2.58" metricValueMin="2.5" pageId="6" pageNumber="7" unit="m" value="254.0" valueMax="258.0" valueMin="250.0">250258 m</quantity>
,
<date id="17738EE8E23EA34C87DCF9B8FDF10DA5" box="[483,637,1985,2009]" pageId="6" pageNumber="7" value="2007-06-10">10 June 2007</date>
.
</paragraph>
</subSubSection>
<caption id="37B2F8A0E23FA34D86A8FCD0FBCA08C3" box="[151,1094,681,704]" httpUri="https://zenodo.org/record/208680/files/figure.png" pageId="7" pageNumber="8" targetBox="[165,1410,205,622]" targetPageId="7">
<paragraph id="6372A828E23FA34D86A8FCD0FBCA08C3" blockId="7.[151,1094,681,704]" box="[151,1094,681,704]" pageId="7" pageNumber="8">
<emphasis id="51B9743AE23FA34D86A8FCD0FE9E08C2" bold="true" box="[151,274,681,704]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
<taxonomicName id="A4CDD3ABE23FA34D8726FCD3FE6D08C3" box="[281,481,681,703]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="brayae">
<emphasis id="51B9743AE23FA34D8726FCD3FE6D08C3" box="[281,481,681,703]" italics="true" pageId="7" pageNumber="8">Saccogaster brayae</emphasis>
</taxonomicName>
, holotype, NMV A 29655-002, SL 36 mm (photo M. Krag).
</paragraph>
</caption>
<subSubSection id="2BD7FBA3E23FA34D86F8FC92FE9B0EA4" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="6372A828E23FA34D86F8FC92FE990E58" blockId="7.[151,1437,747,2033]" pageId="7" pageNumber="8">
<emphasis id="51B9743AE23FA34D86F8FC92FEC90978" bold="true" box="[199,325,747,772]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<taxonomicName id="A4CDD3ABE23FA34D8772FC94FDBE0978" box="[333,562,748,772]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="brayae">
<emphasis id="51B9743AE23FA34D8772FC94FDBE0978" box="[333,562,748,772]" italics="true" pageId="7" pageNumber="8">Saccogaster brayae</emphasis>
</taxonomicName>
differs from all other
<taxonomicName id="A4CDD3ABE23FA34D8509FC94FC4F0978" box="[822,963,749,772]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
<emphasis id="51B9743AE23FA34D8509FC94FC4F0978" box="[822,963,749,772]" italics="true" pageId="7" pageNumber="8">Saccogaster</emphasis>
</taxonomicName>
species in having a broken brown stripe between eye and hind margin of opercle (except for
<emphasis id="51B9743AE23FA34D84C2FD68FCFF0954" box="[765,883,784,808]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="A4CDD3ABE23FA34D84C2FD68FCE70954" box="[765,875,784,808]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hawaii">S. hawaii</taxonomicName>
)
</emphasis>
, precaudal vertebrae 21 and total vertebrae 53, and becoming ripe at a very small SL (
<quantity id="A43505CDE23FA34D8461FD4DFD3B0930" box="[606,695,820,844]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="8" unit="mm" value="36.0">36 mm</quantity>
). Also the following combination of characters is diagnostic: A pair of free spines on frontal above and behind eyes, a large median sub-dermal ethmoidal spine in front of eyes with forward pointed tip; antero-ventrally directed spine at lower angle of preopercle; thin transparent skin; scales present on sides of body; three long gill rakers on anterior arch 45 times the length of gill filaments; palatines with 12 rows of teeth; pectoral peduncle longer than high; fin rays in dorsal 85, anal 52 and pectoral 18; otolith length to height = 1.9, otolith length to colliculum length = 3.1, sulcus without anterior projection.
</paragraph>
<paragraph id="6372A828E23FA34D86F8FA56FE9B0EA4" blockId="7.[151,1437,747,2033]" pageId="7" pageNumber="8">
<emphasis id="51B9743AE23FA34D86F8FA56FEC80E34" bold="true" box="[199,324,1071,1096]" pageId="7" pageNumber="8">Similarity.</emphasis>
<taxonomicName id="A4CDD3ABE23FA34D8773FA48FDA00E34" box="[332,556,1072,1096]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="brayae">
<emphasis id="51B9743AE23FA34D8773FA48FDA00E34" box="[332,556,1072,1096]" italics="true" pageId="7" pageNumber="8">Saccogaster brayae</emphasis>
</taxonomicName>
seems most similar to
<taxonomicName id="A4CDD3ABE23FA34D850AFA48FC130E34" box="[821,927,1072,1096]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="hawaii">
<emphasis id="51B9743AE23FA34D850AFA48FC130E34" box="[821,927,1072,1096]" italics="true" pageId="7" pageNumber="8">S. hawaii</emphasis>
</taxonomicName>
by the brown stripe behind eye, the forward pointed tip of the ethmoidal spine and the scaled body. They differ by
<taxonomicName id="A4CDD3ABE23FA34D8589FA2CFBAD0E10" box="[950,1057,1108,1132]" class="Actinopterygii" family="Bythitidae" genus="Saccogaster" kingdom="Animalia" order="Ophidiiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="brayae">
<emphasis id="51B9743AE23FA34D8589FA2CFBAD0E10" box="[950,1057,1108,1132]" italics="true" pageId="7" pageNumber="8">S. brayae</emphasis>
</taxonomicName>
having opercular spine ending in one prong (
<emphasis id="51B9743AE23FA34D8722FA00FEB00EEC" box="[285,316,1145,1168]" italics="true" pageId="7" pageNumber="8">vs.</emphasis>
three prongs), by long rakers on anterior arch 45 times length of filaments (
<emphasis id="51B9743AE23FA34D82FDFA00FB6C0EEC" box="[1218,1248,1145,1168]" italics="true" pageId="7" pageNumber="8">vs.</emphasis>
long rakers and filaments almost equal in length), precaudal vertebrae 21 (
<emphasis id="51B9743AE23FA34D8513FAE4FCC70EC8" box="[812,843,1181,1204]" italics="true" pageId="7" pageNumber="8">vs.</emphasis>
18), pectoral fin rays 18 (vs. 22) and dorsal fin rays 85 (vs. 92).
</paragraph>
</subSubSection>
<subSubSection id="2BD7FBA3E23FA34286F8FA9AFBFF0840" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="6372A828E23FA34D86F8FA9AFE8B0C60" blockId="7.[151,1437,747,2033]" pageId="7" pageNumber="8">
<emphasis id="51B9743AE23FA34D86F8FA9AFED50E80" bold="true" box="[199,345,1251,1276]" pageId="7" pageNumber="8">Description.</emphasis>
The principal meristic and morphometric characters are shown in
<tableCitation id="2E4F9D93E23FA34D8244FA9DFB5A0E80" box="[1147,1238,1252,1276]" captionStart="TABLE 4" captionStartId="9.[151,239,151,175]" captionTargetBox="[165,1386,208,1519]" captionTargetPageId="9" captionText="TABLE 4. Meristic and morphometric characters of Saccogaster brayi, S. hawaii and S. horrida." httpUri="http://table.plazi.org/id/37B2F8A0E231A34386A8FEEEFB280AD3" pageId="7" pageNumber="8" tableUuid="37B2F8A0E231A34386A8FEEEFB280AD3">Table 4</tableCitation>
. Head and body elongate and compressed with tapering caudal part and blunt snout. Most scales rubbed off but scale-pockets and a few remaining scales (
<quantity id="A43505CDE23FA34D87AEFB55FE640F38" box="[401,488,1324,1348]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="mm" value="0.8">0.8 mm</quantity>
long) indicate squamation on sides of body. Lateral line not visible. Dorsal fin origin above proximal part of pectoral fin and anal fin origin well behind midpoint of fish. Pectoral fin ending less than halfway to anus with peduncle a little longer than broad and free of body. Anterior gill arch with three spiny knobs on upper branch, a developed raker in angle and lower branch with two developed rakers with a knob in between followed by eight knobs. Developed rakers 45 times length of gill filaments (
<figureCitation id="FBF6B4ADE23FA34D82B9FBC5FB580FA8" box="[1158,1236,1468,1492]" captionStart="FIGURE 4" captionStartId="8.[151,250,1832,1855]" captionTargetBox="[168,1428,809,1785]" captionTargetId="figure@8.[151,1436,774,1806]" captionTargetPageId="8" captionText="FIGURE 4. Saccogaster brayae, holotype: A—Lateral view of head. B—Lateral radiograph of head. C—Dorsal view of head. D— Anterior right gill arch. E—Median view of right otolith. F—Dorsal view of right otolith. G—Intromittent organ." httpUri="https://zenodo.org/record/208681/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
D). Two small pseudobranchial filaments. The well developed testes and intromittent organ (
<figureCitation id="FBF6B4ADE23FA34D85C3FB99FBB20F84" box="[1020,1086,1504,1528]" captionStart="FIGURE 4" captionStartId="8.[151,250,1832,1855]" captionTargetBox="[168,1428,809,1785]" captionTargetId="figure@8.[151,1436,774,1806]" captionTargetPageId="8" captionText="FIGURE 4. Saccogaster brayae, holotype: A—Lateral view of head. B—Lateral radiograph of head. C—Dorsal view of head. D— Anterior right gill arch. E—Median view of right otolith. F—Dorsal view of right otolith. G—Intromittent organ." httpUri="https://zenodo.org/record/208681/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
G) show that the specimen is a ripe male.
</paragraph>
<paragraph id="6372A828E23FA34D86F8F851FB340CD0" blockId="7.[151,1437,747,2033]" pageId="7" pageNumber="8">Axial skeleton (from radiographs): Number of precaudal vertebrae 21. Anterior neural spine very short. Neural spines 24 long, depressed and with pointed tips. Neural spines 520 short and with large basal parts, length of spines increasing posteriorad. Parapophyses developed on vertebrae 620. Pleural ribs distinct at vertebrae 18, but pleural ribs on remaining precaudal vertebrae and presence of epipleural ribs are questionable.</paragraph>
<paragraph id="6372A828E23FA34D86F8F8C1FE3A0D40" blockId="7.[151,1437,747,2033]" pageId="7" pageNumber="8">Dentition: Palatines with about 25 pointed teeth in 12 rows; larger teeth in inner row. Vomer boomerangshaped with about 25 pointed teeth in 23 irregular rows; six longer teeth in inner row. Premaxillaries with granular teeth except for five fangs at symphysis. Dentaries with 34 irregular tooth rows, outer rows with granular teeth and inner row with fangs.</paragraph>
<paragraph id="6372A828E23FA34D86F8F931FA110D8C" blockId="7.[151,1437,747,2033]" pageId="7" pageNumber="8">
Head morphology (
<figureCitation id="FBF6B4ADE23FA34D8793F931FE780D1C" box="[428,500,1864,1888]" captionStart="FIGURE 4" captionStartId="8.[151,250,1832,1855]" captionTargetBox="[168,1428,809,1785]" captionTargetId="figure@8.[151,1436,774,1806]" captionTargetPageId="8" captionText="FIGURE 4. Saccogaster brayae, holotype: A—Lateral view of head. B—Lateral radiograph of head. C—Dorsal view of head. D— Anterior right gill arch. E—Median view of right otolith. F—Dorsal view of right otolith. G—Intromittent organ." httpUri="https://zenodo.org/record/208681/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
AC): Head profile concave above eyes. Head with pair of free spines as an integrated part of a transverse rim on frontal above and slightly behind eyes and a large, median sub-dermal ethmoidal spine in front of eyes with forward pointed tip. Anterior nostril placed close to upper lip and small posterior nostril placed close to eye. Head pores: 1 supraorbital pore at tip of snout, 2 anterior infraorbital pores below and in front of eye, 1 anterior mandibular pore at tip of jaw. Head skin thin, tight. No neuromasts visible.
</paragraph>
<paragraph id="6372A828E230A34286A8FEEEFDCC0B60" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="9">Head with dusky background pigmentation, broken brown stripe between eye and hind margin of opercle and broken brown pigmentation patches below and behind eye. Upper jaw ends well behind eye with posterior margin vertically expanded. Opercular spine pointed, just reaching hind margin of operculum. Antero-ventrally directed spine at lower angle of preoperculum.</paragraph>
<paragraph id="6372A828E230A34286F8FF5EFF630BB3" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="9">
Otolith (
<figureCitation id="FBF6B4ADE230A342871AFF5EFEE40B43" box="[293,360,295,319]" captionStart="FIGURE 4" captionStartId="8.[151,250,1832,1855]" captionTargetBox="[168,1428,809,1785]" captionTargetId="figure@8.[151,1436,774,1806]" captionTargetPageId="8" captionText="FIGURE 4. Saccogaster brayae, holotype: A—Lateral view of head. B—Lateral radiograph of head. C—Dorsal view of head. D— Anterior right gill arch. E—Median view of right otolith. F—Dorsal view of right otolith. G—Intromittent organ." httpUri="https://zenodo.org/record/208681/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
EF): Otolith oval, moderately thick; otolith length to height = 1.9, otolith height to thickness = 1.9. Dorsal and ventral rims gently curved, dorsal rim broadly undulating. Anterior and posterior tips regularly rounded resulting in a regular oval appearance of the otolith. Inner face flat with centrally positioned, undivided, small, oval, shallow sulcus without anterior projection. Otolith length to colliculum length = 3.1. Outer face convex.
</paragraph>
<paragraph id="6372A828E230A34286F8FFA5FBFF0840" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="9">Coloration: After five years of preservation body uniformly brownish with faint dark pigmentation dorsally. Head with dusky, dense pigmentation and a dark brown stripe from eye to hind margin of opercle. Larger, dark pigment spots on frontal plate seen through skin on dorsum. Eyes blue with yellow lens.</paragraph>
</subSubSection>
<subSubSection id="2BD7FBA3E230A34286F8FC3EFDFB08DB" pageId="8" pageNumber="9" type="biology_ecology">
<paragraph id="6372A828E230A34286F8FC3EFDFB08DB" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="9">
<emphasis id="51B9743AE230A34286F8FC3EFE7C081C" bold="true" box="[199,496,583,608]" pageId="8" pageNumber="9">Biology and distribution</emphasis>
(
<figureCitation id="FBF6B4ADE230A342843EFC3EFDD40823" box="[513,600,583,608]" captionStart="FIGURE 25" captionStartId="32.[151,250,1767,1790]" captionTargetBox="[161,1424,1103,1720]" captionTargetId="figure@32.[151,1436,1073,1753]" captionTargetPageId="32" captionText="FIGURE 25. Geographic distribution of Hastatobythites and Saccogaster in the Indo-West Pacific. Each marking may represent more than one locality." httpUri="https://zenodo.org/record/208702/files/figure.png" pageId="8" pageNumber="9">Fig. 25</figureCitation>
). A viviparous species occurring on the upper continental slope. Known from a
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male with well developed testes and intromittent organ, caught off northwestern
<collectingCountry id="1BDAE8B8E230A34282D3FC15FADF08F8" box="[1260,1363,620,644]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
organ (
<figureCitation id="FBF6B4ADE230A34286A0FCF6FF6F08DB" box="[159,227,655,679]" captionStart="FIGURE 4" captionStartId="8.[151,250,1832,1855]" captionTargetBox="[168,1428,809,1785]" captionTargetId="figure@8.[151,1436,774,1806]" captionTargetPageId="8" captionText="FIGURE 4. Saccogaster brayae, holotype: A—Lateral view of head. B—Lateral radiograph of head. C—Dorsal view of head. D— Anterior right gill arch. E—Median view of right otolith. F—Dorsal view of right otolith. G—Intromittent organ." httpUri="https://zenodo.org/record/208681/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
G), caught at a depth of
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250
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.
</paragraph>
</subSubSection>
<subSubSection id="2BD7FBA3E230A34286F8FCCDFB4208B0" box="[199,1230,692,717]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="6372A828E230A34286F8FCCDFB4208B0" blockId="8.[151,1437,151,717]" box="[199,1230,692,717]" pageId="8" pageNumber="9">
<emphasis id="51B9743AE230A34286F8FCCDFEC208B0" bold="true" box="[199,334,692,716]" pageId="8" pageNumber="9">Etymology.</emphasis>
Named after Dianne Bray, MNV, for her support with material for this revision.
</paragraph>
</subSubSection>
</treatment>
</document>