treatments-xml/data/E5/72/98/E572986CFFF3C86001ACFAD2FBC4FAFF.xml
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<document id="73B9F1514ABCC533FA4EE802062E8695" ID-CLB-Dataset="28982" ID-DOI="10.11646/zootaxa.4461.2.3" ID-GBIF-Dataset="9ec1478c-2bdc-43d3-9317-6e38503f1d51" ID-ISSN="1175-5326" ID-Zenodo-Dep="1460028" ID-ZooBank="E8ABFFFA-3F81-4C87-9EBC-D1D984F235C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539336798418" checkinUser="plazi" docAuthor="Fernandes, Jose Antonio Marin, Nascimento, Agata Tyanne Silva &amp; Nunes, Benedito Mendes" docDate="2018" docId="E572986CFFF3C86001ACFAD2FBC4FAFF" docLanguage="en" docName="zootaxa.4461.2.3.pdf" docOrigin="Zootaxa 4461 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygoda variegata Fernandes &amp; Nascimento &amp; Nunes 2018, sp. nov." docType="treatment" docUuid="3B6867AE-2E1B-491E-8505-96FC485E16E4" docUuidSource="ZooBank" docVersion="10" lastPageNumber="212" masterDocId="194BE014FFF6C867013BFFC5FF98FFF3" masterDocTitle="Revision of Pygoda Amyot &amp; Serville, 1843 stat. rest. (Heteroptera: Pentatomidae: Edessinae) with description of four new species" masterLastPageNumber="232" masterPageNumber="205" pageNumber="210" updateTime="1698579730646" updateUser="ExternalLinkService">
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<mods:title id="6C1F147143264C65CD769A2265CC9A05">Revision of Pygoda Amyot &amp; Serville, 1843 stat. rest. (Heteroptera: Pentatomidae: Edessinae) with description of four new species</mods:title>
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<mods:namePart id="B87D475BB6F21B921019622D97E393C5">Fernandes, Jose Antonio Marin</mods:namePart>
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<mods:namePart id="07A105E5DA42679FB2F9800D86FA23B8">Nascimento, Agata Tyanne Silva</mods:namePart>
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<mods:namePart id="C2300821B8A3DD5EA893D91133B02E7C">Nunes, Benedito Mendes</mods:namePart>
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<emphasis id="5FAFF568FFF3C86201ACFAD2FE54FAC1" bold="true" box="[151,460,1303,1330]" pageId="5" pageNumber="210">
<taxonomicName id="AADB52F9FFF3C86201ACFAD2FEF0FAC2" authority="Fernandes &amp; Nascimento &amp; Nunes, 2018" authorityName="Fernandes &amp; Nascimento &amp; Nunes" authorityYear="2018" box="[151,360,1303,1329]" class="Insecta" genus="Pygoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="210" phylum="Arthropoda" rank="species" species="variegata" status="sp. nov.">
<emphasis id="5FAFF568FFF3C86201ACFAD2FEF0FAC2" bold="true" box="[151,360,1303,1329]" italics="true" pageId="5" pageNumber="210">Pygoda variegata</emphasis>
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<taxonomicNameLabel id="449C4813FFF3C8620054FADDFE54FAC1" box="[367,460,1304,1330]" pageId="5" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="6D64297AFFF3C86201ACFAFFFDFFFAA0" blockId="5.[151,615,1303,1363]" box="[151,615,1338,1363]" pageId="5" pageNumber="210">
(
<figureCitation id="F5E035FFFFF3C86201A5FAFFFEDBFAA0" box="[158,323,1338,1363]" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="5" pageNumber="210">Figures 2AD</figureCitation>
,
<figureCitation id="F5E035FFFFF3C8620076FAFEFE14FAA0" box="[333,396,1339,1363]" captionStart="FIGURE 10" captionStartId="22.[151,250,1749,1771]" captionTargetBox="[204,1388,317,1580]" captionTargetId="figure@22.[176,1410,295,1744]" captionTargetPageId="22" captionText="FIGURE 10. Phallus: A—Hypoxys sp.; B—Pygoda amianta sp. n.; C—Pygoda variegata sp. n.; D—Pygoda poecila sp. n.; E—Pygoda irrorata; F—Pygoda polita; G—Pygoda ramosa sp. n.; H—Pygoda expolita (ph—phallus; ves—vesica. Scale = 1mm)." httpUri="https://zenodo.org/record/1460048/files/figure.png" pageId="5" pageNumber="210">10C</figureCitation>
,
<figureCitation id="F5E035FFFFF3C86200A3FAFEFE4EFAA0" box="[408,470,1339,1363]" captionStart="FIGURE 11" captionStartId="23.[151,250,1864,1886]" captionText="FIGURE 11. Internal female: A—Hypoxys sp.; B—Pygoda amianta sp. n.; C—Pygoda variegata sp. n.; D—Pygoda poecila sp. n.; E—Pygoda irrorata; F—Pygoda polita; G—Pygoda ramosa sp. n.; H—Pygoda expolita (aaf—anterior annular flange; ch—chitinellipsen; cs—capsula seminalis; dr—ductus receptaculi; gc9—gonocoxite 9; la9—laterotergite 9; paf—posterior annular flange; pc—pars communis; pi—pars intermedialis; tdr—thickening of ductus receptaculi; tvi—thickening of vaginal intima; x—abdominal segment X. Scale = 1mm)." pageId="5" pageNumber="210">11C</figureCitation>
,
<figureCitation id="F5E035FFFFF3C86200D9FAFEFDADFAA0" box="[482,565,1339,1363]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="5" pageNumber="210">12CD</figureCitation>
,
<figureCitation id="F5E035FFFFF3C862037AFAFEFDC6FAA0" box="[577,606,1339,1363]" captionStart="FIGURE 14" captionStartId="25.[151,250,2000,2022]" captionText="FIGURE 14. Distributional map of the new species of Pygoda." pageId="5" pageNumber="210">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="25C17AF1FFF3C86201ACFA47FA1DFA68" box="[151,1413,1410,1435]" pageId="5" pageNumber="210" type="etymology">
<paragraph id="6D64297AFFF3C86201ACFA47FA1DFA68" blockId="5.[151,1437,1410,2011]" box="[151,1413,1410,1435]" pageId="5" pageNumber="210">
<emphasis id="5FAFF568FFF3C86201ACFA47FE86FA68" bold="true" box="[151,286,1410,1435]" pageId="5" pageNumber="210">Etymology.</emphasis>
Name refers to the mix of brown, green and yellow of the body (L. Variegatus—of different colors).
</paragraph>
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<subSubSection id="25C17AF1FFF3C86201FCFA63FCE1F8B8" pageId="5" pageNumber="210" type="materials_examined">
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<emphasis id="5FAFF568FFF3C86201FCFA63FE2CFA4C" bold="true" box="[199,436,1446,1471]" pageId="5" pageNumber="210">Material examined.</emphasis>
<typeStatus id="B26097D8FFF3C8620084FA63FDB3FA4C" box="[447,555,1446,1471]" pageId="5" pageNumber="210" type="holotype">Holotype</typeStatus>
. Male.
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<collectingCountry id="15CC69EAFFF3C86203BEFA63FCAFFA4C" box="[645,823,1446,1471]" name="Costa Rica" pageId="5" pageNumber="210">COSTA RICA</collectingCountry>
.
<collectingRegion id="AF1FE798FFF3C8620273FA63FC48FA4C" box="[840,976,1446,1471]" country="Costa Rica" name="Puntarenas" pageId="5" pageNumber="210">Puntarenas</collectingRegion>
</emphasis>
: Fca. Cafrosa, Est. Las Mellizas, P. N. Amistad,
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<date id="19650FBAFFF3C862005DFA0EFE40FA10" box="[358,472,1483,1507]" pageId="5" pageNumber="210" value="1989-10">Oct. 1989</date>
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).
</paragraph>
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.
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:
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Est. Biol. Las Alturas, Coto Brus,
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<date id="19650FBAFFF3C86205B6FA2AFA9CF9F4" box="[1165,1284,1519,1543]" pageId="5" pageNumber="210" value="1991-08">Ago. 1991</date>
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same data,
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&amp;
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000 308886/308905/308911 (
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);
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Finca Cafrosa, Embalce,
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</quantity>
NO de
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,
<elevation id="E6F6CE49FFF3C8620588F99EFA93F980" box="[1203,1291,1627,1651]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="210" unit="m" value="1.2">
<quantity id="AA23849FFFF3C8620588F99EFA93F980" box="[1203,1291,1627,1651]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="210" unit="m" value="1.2">1.200m</quantity>
</elevation>
,
<collectingDate id="0921F652FFF3C8620422F99EFF4AF964" pageId="5" pageNumber="210" value="1996-04-17" valueMax="1996-04-24" valueMin="1996-04-17">
<date id="19650FBAFFF3C8620422F99EFF4AF964" pageId="5" pageNumber="210" value="1996-04-17" valueMax="1996-04-24" valueMin="1996-04-17">1724-Apr-1996</date>
</collectingDate>
,
<collectorName id="C02E4CACFFF3C86201E6F9BAFEC4F965" box="[221,348,1663,1686]" pageId="5" pageNumber="210">E. Navarro</collectorName>
, L-S-317800, 596200
</materialsCitation>
. CRI002 439095 (UFRG); ♀ San Vito, Coto Brus, Las Cruces,
<date id="19650FBAFFF3C862041CF9BBFA0FF964" box="[1319,1431,1662,1687]" pageId="5" pageNumber="210" value="1995-11-07">7-xi-1995</date>
, J. A. Chacón,
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147769 (
<collectionCode id="0BCAB1BFFFF3C862033FF966FDCDF948" box="[516,597,1699,1723]" country="Brazil" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="5" pageNumber="210">UFRG</collectionCode>
);
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<location id="68047FA1FFF3C86203B2F967FCCBF948" LSID="urn:lsid:plazi:treatment:E572986CFFF3C86001ACFAD2FBC4FAFF:68047FA1FFF3C86203B2F967FCCBF948" box="[649,851,1698,1723]" country="Costa Rica" name="Finca Las Cruces" pageId="5" pageNumber="210" stateProvince="Java">Finca Las Cruces</location>
,
<quantity id="AA23849FFFF3C862025BF966FC0DF948" box="[864,917,1698,1723]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="5" pageNumber="210" unit="mi" value="4.0">4 mi</quantity>
S San Vito de
<collectingRegion id="AF1FE798FFF3C8620572F966FBE5F949" box="[1097,1149,1699,1722]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="210">Java</collectingRegion>
,
<collectingDate id="0921F652FFF3C86205B1F966FAC3F948" box="[1162,1371,1698,1723]" pageId="5" pageNumber="210" value="1969-08-11" valueMax="1969-08-19" valueMin="1969-08-11">
<date id="19650FBAFFF3C86205B1F966FAC3F948" box="[1162,1371,1698,1723]" pageId="5" pageNumber="210" value="1969-08-11" valueMax="1969-08-19" valueMin="1969-08-11">Aug. 1119, 1969</date>
</collectingDate>
, DHJ
<elevation id="E6F6CE49FFF3C86201ACF903FF7DF92C" box="[151,229,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="5" pageNumber="210" unit="m" value="1260.0">
<quantity id="AA23849FFFF3C86201ACF903FF7DF92C" box="[151,229,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="5" pageNumber="210" unit="m" value="1260.0">1260m</quantity>
</elevation>
elev. Collected by
<collectorName id="C02E4CACFFF3C86200F8F902FDCFF92D" box="[451,599,1735,1758]" pageId="5" pageNumber="210">D. H. Janzen</collectorName>
(
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)
</materialsCitation>
.
<materialsCitation id="DDB32327FFF3C86203E8F903FC93F8D4" ID-GBIF-Occurrence="1919493352" collectingDate="1968-06" collectionCode="MNHN" collectorName="Coll. Noualhier &amp; Museum Paris &amp; M. Lemaire Leg. &amp; Lino &amp; Los Naranjos" country="Panama" location="PANAMA" pageId="5" pageNumber="210" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="PANAMA" typeStatus="paratype">
<emphasis id="5FAFF568FFF3C86203E8F903FC5CF92C" bold="true" box="[723,964,1734,1759]" pageId="5" pageNumber="210">
<collectingRegion id="AF1FE798FFF3C86203E8F903FCC8F92C" box="[723,848,1734,1759]" country="Panama" name="Panama" pageId="5" pageNumber="210">
<collectingCountry id="15CC69EAFFF3C86203E8F903FCC8F92C" box="[723,848,1734,1759]" name="Panama" pageId="5" pageNumber="210">PANAMA</collectingCountry>
</collectingRegion>
.
<collectingRegion id="AF1FE798FFF3C8620266F903FC5CF92C" box="[861,964,1734,1759]" country="Panama" name="Chiriqui" pageId="5" pageNumber="210">Chiriquí</collectingRegion>
</emphasis>
:
<specimenCount id="7BDDE2F3FFF3C86202EEF902FC72F92D" box="[981,1002,1735,1758]" pageId="5" pageNumber="210" type="male"></specimenCount>
1898,
<collectorName id="C02E4CACFFF3C8620501F903FB75F92C" box="[1082,1261,1734,1759]" pageId="5" pageNumber="210">Coll. Noualhier</collectorName>
,
<collectorName id="C02E4CACFFF3C86205C2F902FA04F92C" box="[1273,1436,1734,1759]" pageId="5" pageNumber="210">Museum Paris</collectorName>
<collectionCode id="0BCAB1BFFFF3C86201ACF92EFF77F8F0" box="[151,239,1771,1795]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="210">MNHN</collectionCode>
(EH) 24131 (
<collectionCode id="0BCAB1BFFFF3C86200B2F92EFE7DF8F0" box="[393,485,1771,1795]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="210">MNHN</collectionCode>
);
<specimenCount id="7BDDE2F3FFF3C86200C7F92EFD87F8F0" box="[508,543,1771,1795]" pageId="5" pageNumber="210" type="male">2♂</specimenCount>
Chaine du
<collectingRegion id="AF1FE798FFF3C8620397F92FFC94F8F0" box="[684,780,1770,1795]" country="Panama" name="Chiriqui" pageId="5" pageNumber="210">Chiriqui</collectingRegion>
,
<collectingDate id="0921F652FFF3C8620227F92FFCEEF8F0" box="[796,886,1770,1795]" pageId="5" pageNumber="210" value="1968-06">
<date id="19650FBAFFF3C8620227F92FFCEEF8F0" box="[796,886,1770,1795]" pageId="5" pageNumber="210" value="1968-06">vi-1968</date>
</collectingDate>
,
<collectorName id="C02E4CACFFF3C86202BEF92EFBD6F8F0" box="[901,1102,1770,1795]" pageId="5" pageNumber="210">M. Lemaire Leg.</collectorName>
,
<collectorName id="C02E4CACFFF3C8620565F92EFA9BF8F0" box="[1118,1283,1770,1795]" pageId="5" pageNumber="210">Museum Paris</collectorName>
<collectionCode id="0BCAB1BFFFF3C8620434F92EFAFFF8F0" box="[1295,1383,1771,1795]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="210">MNHN</collectionCode>
(EH) 24132 and 24133 (
<collectionCode id="0BCAB1BFFFF3C8620043F8CAFE4CF8D4" box="[376,468,1807,1831]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="210">MNHN</collectionCode>
);
<specimenCount id="7BDDE2F3FFF3C86200D7F8CAFD99F8D5" box="[492,513,1807,1830]" pageId="5" pageNumber="210" type="female"></specimenCount>
<collectorName id="C02E4CACFFF3C8620336F8CBFDDAF8D4" box="[525,578,1806,1831]" pageId="5" pageNumber="210">Lino</collectorName>
[
<collectorName id="C02E4CACFFF3C862036EF8CAFD63F8D4" box="[597,763,1806,1831]" pageId="5" pageNumber="210">Los Naranjos</collectorName>
?]
</materialsCitation>
,
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. Coll. Fassl. Coll.
<materialsCitation id="DDB32327FFF3C8620579F8CAFA00F8D4" ID-GBIF-Occurrence="1919493371" box="[1090,1432,1806,1831]" collectionCode="NNM" country="Panama" location="Mac Gillavry" municipality="Dr." pageId="5" pageNumber="210" specimenCount="1" stateProvince="PANAMA" typeStatus="paratype">
<collectingMunicipality id="8D00B300FFF3C8620579F8CAFBFEF8D5" box="[1090,1126,1807,1830]" pageId="5" pageNumber="210">Dr.</collectingMunicipality>
D.
<location id="68047FA1FFF3C86205A2F8CAFAACF8D4" LSID="urn:lsid:plazi:treatment:E572986CFFF3C86001ACFAD2FBC4FAFF:68047FA1FFF3C86205A2F8CAFAACF8D4" box="[1177,1332,1806,1831]" country="Panama" municipality="Dr." name="Mac Gillavry" pageId="5" pageNumber="210" stateProvince="PANAMA">Mac Gillavry</location>
(
<collectionCode id="0BCAB1BFFFF3C8620473F8CAFA08F8D4" box="[1352,1424,1807,1831]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="5" pageNumber="210">NNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="DDB32327FFF3C86201ACF8F7FCE1F8B8" ID-GBIF-Occurrence="1919493374" box="[151,889,1842,1867]" collectingDate="1936-12-26" collectionCode="USNM" collectorName="R. Bliss" country="Panama" location="Corozal Canal Zone" pageId="5" pageNumber="210" specimenCount="1" specimenCount-male="1" stateProvince="Panama" typeStatus="paratype">
<collectingRegion id="AF1FE798FFF3C86201ACF8F7FF61F8B8" box="[151,249,1842,1867]" country="Panama" name="Panama" pageId="5" pageNumber="210">
<collectingCountry id="15CC69EAFFF3C86201ACF8F7FF61F8B8" box="[151,249,1842,1867]" name="Panama" pageId="5" pageNumber="210">
<emphasis id="5FAFF568FFF3C86201ACF8F7FF61F8B8" bold="true" box="[151,249,1842,1867]" pageId="5" pageNumber="210">Panama</emphasis>
</collectingCountry>
</collectingRegion>
:
<specimenCount id="7BDDE2F3FFF3C8620033F8F6FE85F8B9" box="[264,285,1843,1866]" pageId="5" pageNumber="210" type="male"></specimenCount>
<location id="68047FA1FFF3C862001FF8F7FD90F8B9" LSID="urn:lsid:plazi:treatment:E572986CFFF3C86001ACFAD2FBC4FAFF:68047FA1FFF3C862001FF8F7FD90F8B9" box="[292,520,1842,1867]" country="Panama" name="Corozal Canal Zone" pageId="5" pageNumber="210" stateProvince="Panama">Corozal Canal Zone</location>
,
<collectingDate id="0921F652FFF3C8620328F8F7FD3CF8B8" box="[531,676,1842,1867]" pageId="5" pageNumber="210" value="1936-12-26">
<date id="19650FBAFFF3C8620328F8F7FD3CF8B8" box="[531,676,1842,1867]" pageId="5" pageNumber="210" value="1936-12-26">XII-26-1936</date>
</collectingDate>
.
<collectorName id="C02E4CACFFF3C8620395F8F6FC9FF8B8" box="[686,775,1842,1867]" pageId="5" pageNumber="210">R. Bliss</collectorName>
(
<collectionCode id="0BCAB1BFFFF3C862022DF8F6FCF5F8B8" box="[790,877,1843,1867]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="210">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="25C17AF1FFF3C86201FCF893FB8FF844" pageId="5" pageNumber="210" type="description">
<paragraph id="6D64297AFFF3C86201FCF893FB8FF844" blockId="5.[151,1437,1410,2011]" pageId="5" pageNumber="210">
<emphasis id="5FAFF568FFF3C86201FCF893FEE5F89C" bold="true" box="[199,381,1878,1903]" pageId="5" pageNumber="210">Measurements.</emphasis>
Total length 21.023.7; total width 12.213.4; head length 1.81.9; head width 3.53.9; length of antennal articles I: 0.81.1; II: 2.02.5; III: 1.72.0; IV: 4.14.9; V: 4.34.8; pronotum length 4.14.8; pronotum width 14.215.1; scutellum length 9.611.0; scutellum width 7.48.3.
</paragraph>
</subSubSection>
<subSubSection id="25C17AF1FFF3C86001FCF807FABAFB8E" lastPageId="7" lastPageNumber="212" pageId="5" pageNumber="210" type="diagnosis">
<paragraph id="6D64297AFFF3C86101FCF807FAC7FEB3" blockId="5.[151,1437,1410,2011]" lastBlockId="6.[151,1437,151,429]" lastPageId="6" lastPageNumber="211" pageId="5" pageNumber="210">
<emphasis id="5FAFF568FFF3C86201FCF807FED9F828" bold="true" box="[199,321,1986,2011]" pageId="5" pageNumber="210">Diagnosis.</emphasis>
Species large (
<quantity id="AA23849FFFF3C86200CAF806FDE5F828" box="[497,637,1987,2011]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="5" pageNumber="210" unit="mm" value="22.5" valueMax="24.0" valueMin="21.0">21—24 mm</quantity>
) dorsally dark green or bluish green. Base of the head with a median black stain and two stripes that project anteriorly delimiting tylus. Pronotum and scutellum with punctuation black, sometimes on dark spots forming dark lines especially on scutellum (
<figureCitation id="F5E035FFFFF0C86102E8FF79FBA7FF27" box="[979,1087,188,213]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="6" pageNumber="211">Fig. 12C</figureCitation>
). Evaporatorium concolorous. Connexivum green with medial yellow callus (
<figureCitation id="F5E035FFFFF0C861038DFF1AFC87FF0B" box="[694,799,223,248]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="6" pageNumber="211">Fig. 12C</figureCitation>
); segment VII with basal third or less green remaining black, medial yellow callus present. Body ventrally variegated with green (bluish green), yellow and dark brown lines on the edge of sternites and pseudosutures. Antennae yellow to orange. Legs yellow and reddish brown.
</paragraph>
<paragraph id="6D64297AFFF0C86101FCFE89FD44FE5E" blockId="6.[151,1437,151,429]" pageId="6" pageNumber="211">
Head. Mandibular plates with dark lines on lateral margin and few dark punctures. Tylus with median greenish yellow to yellow stripe (
<figureCitation id="F5E035FFFFF0C8610080FEAAFDBFFE7B" box="[443,551,367,392]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="6" pageNumber="211">Fig. 12C</figureCitation>
). Ventral surface greenish yellow anteriorly and green distally. First rostral segment clearly surpassing bucculae in lateral view.
</paragraph>
<caption id="39A479F2FFF0C86101ACF8FBFD80F880" httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="6" pageNumber="211" startId="6.[151,250,1854,1876]" targetBox="[236,1369,516,1805]" targetPageId="6">
<paragraph id="6D64297AFFF0C86101ACF8FBFD80F880" blockId="6.[151,1436,1854,1907]" pageId="6" pageNumber="211">
<emphasis id="5FAFF568FFF0C86101ACF8FBFE8FF8A0" bold="true" box="[151,279,1854,1876]" pageId="6" pageNumber="211">FIGURE 2.</emphasis>
<taxonomicName id="AADB52F9FFF0C8610019F8FBFE43F8A7" authority="Fernandes &amp; Nascimento &amp; Nunes, 2018" authorityName="Fernandes &amp; Nascimento &amp; Nunes" authorityYear="2018" box="[290,475,1854,1876]" class="Insecta" genus="Pygoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="211" phylum="Arthropoda" rank="species" species="variegata" status="sp. nov.">
<emphasis id="5FAFF568FFF0C8610019F8FBFE43F8A7" box="[290,475,1854,1876]" italics="true" pageId="6" pageNumber="211">Pygoda variegata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="449C4813FFF0C86100DEF8FAFDBBF8A7" box="[485,547,1855,1876]" pageId="6" pageNumber="211" rank="species">
<emphasis id="5FAFF568FFF0C86100DEF8FAFDBBF8A7" bold="true" box="[485,547,1855,1876]" pageId="6" pageNumber="211">sp. n.</emphasis>
</taxonomicNameLabel>
AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm).
</paragraph>
</caption>
<paragraph id="6D64297AFFF0C86001FCF85AFDF3FE07" blockId="6.[151,1436,1951,2012]" lastBlockId="7.[151,1437,151,1293]" lastPageId="7" lastPageNumber="212" pageId="6" pageNumber="211">
Thorax. Pronotum densely punctured, punctures frequently on dark spots; cicatrices variegate with brown, green and greenish yellow, delimiting dark brown furrows surrounded by pale greenish yellow calloused stripes (
<figureCitation id="F5E035FFFFF1C86001A4FF52FE9FFF43" box="[159,263,151,176]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="7" pageNumber="212">Fig. 12C</figureCitation>
); anterior delimiting furrow punctured. Anterolateral margin pale yellow to yellow; line of black coarse punctures long, exceeding ½ of the margin. Scutellum with coarse, large, black punctures sparser on anterior half and median longitudinal line than on remaining parts, lateral and distal punctures small, fine and black; most of the punctures on dark spots. Corium with a few yellow veins not branched but forming a rhombus (
<figureCitation id="F5E035FFFFF1C8600587FEC1FA84FEEF" box="[1212,1308,260,285]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="7" pageNumber="212">Fig 12C</figureCitation>
). Propleura with dark stripe alongside concolorous to dark, coarse, dense punctures; lateral margin green. Evaporatorium concolorous with dark stains and rugose. Peritreme long reaching 2/3 of the distance between ostiole of the scent gland and margin of the body. Metasternal process ventrally green, but lateral part predominantly greenish yellow; arms broad and receiving fourth and most of third rostral segment (
<figureCitation id="F5E035FFFFF1C86002B9FE51FC73FE5F" box="[898,1003,404,429]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="7" pageNumber="212">Fig. 12D</figureCitation>
). Legs with part of trochanters, femora and proximal part of tibiae reddish brown; remaining of fore and mid legs dark yellow, tarsal segments suffused; remaining of hind legs yellow (
<figureCitation id="F5E035FFFFF1C86000CDFE19FDC7FE07" box="[502,607,476,501]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="7" pageNumber="212">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="6D64297AFFF1C86001FCFE3AFE8CFD77" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="212">
Abdomen. Connexivum with excavations dark and densely punctured, except close to distal margin; base of lateral spines yellow. Lateral part of segment VII, corresponding to ventral side of connexivum, light green with black margin. Large yellow spot surrounding spiracle. Small black rounded spot close to spiracle on each segment (
<figureCitation id="F5E035FFFFF1C86001A4FDA9FE90FD77" box="[159,264,620,645]" captionStart="FIGURE 12" captionStartId="24.[151,250,1969,1991]" captionTargetBox="[262,1368,199,1922]" captionTargetId="figure@24.[196,1390,193,1950]" captionTargetPageId="24" captionText="FIGURE 12. Dorsal and ventral view. Pygoda amianta sp. n. AB; Pygoda variegata sp. n. CD; Pygoda poecila sp. n. EF; Pygoda irrorata GH; Pygoda polita IJ; Pygoda ramosa sp. n. KL (Scale = 5mm)." httpUri="https://zenodo.org/record/1460050/files/figure.png" pageId="7" pageNumber="212">Fig. 12D</figureCitation>
).
</paragraph>
<paragraph id="6D64297AFFF1C86001FCFD4AFE8BFC57" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="212">
Male (
<figureCitation id="F5E035FFFFF1C860002BFD4AFE13FD5B" box="[272,395,655,680]" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Fig. 2AC</figureCitation>
). Pygophore with posterolateral angles dorsally flattened and well projected posteriorly (
<figureCitation id="F5E035FFFFF1C8600454FD4AFF25FD3F" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Fig. 2A</figureCitation>
). Superior process of the genital cup small, short and elliptical. Paramere with posterior process linguiform and laterally curved; dorsal process short and rounded; dorsal margin distinctly excavated (
<figureCitation id="F5E035FFFFF1C8600548FD12FB55FD03" box="[1139,1229,727,752]" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Fig. 2B</figureCitation>
). Proctiger almost entirely black, long, slightly exceeding the level of posterolateral angles (
<figureCitation id="F5E035FFFFF1C86002E6FD39FBFFFCE7" box="[989,1127,764,789]" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Figs 2 A, C</figureCitation>
); without dorsal crest (
<figureCitation id="F5E035FFFFF1C8600454FD39FF47FCCB" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Fig. 2AB</figureCitation>
); dorsal part of posterior face bitumid. Ventral rim medially with dark brown spot; expansions barely developed and not tumid. Ventral surface mostly yellow with green distal stripe and few suffused stains; punctures coarse and concolorous (
<figureCitation id="F5E035FFFFF1C860008FFCA2FD97FC73" box="[436,527,871,896]" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Fig. 2C</figureCitation>
). Phallus long, distal part remarkably developed in a ventral conical expansion (
<figureCitation id="F5E035FFFFF1C86001A4FC49FE9EFC57" box="[159,262,908,933]" captionStart="FIGURE 10" captionStartId="22.[151,250,1749,1771]" captionTargetBox="[204,1388,317,1580]" captionTargetId="figure@22.[176,1410,295,1744]" captionTargetPageId="22" captionText="FIGURE 10. Phallus: A—Hypoxys sp.; B—Pygoda amianta sp. n.; C—Pygoda variegata sp. n.; D—Pygoda poecila sp. n.; E—Pygoda irrorata; F—Pygoda polita; G—Pygoda ramosa sp. n.; H—Pygoda expolita (ph—phallus; ves—vesica. Scale = 1mm)." httpUri="https://zenodo.org/record/1460048/files/figure.png" pageId="7" pageNumber="212">Fig. 10C</figureCitation>
).
</paragraph>
<paragraph id="6D64297AFFF1C86001FCFC6AFABAFB8E" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="212">
Female (
<figureCitation id="F5E035FFFFF1C8600017FC6AFE13FC3B" box="[300,395,943,968]" captionStart="FIGURE 2" captionStartId="6.[151,250,1854,1876]" captionTargetBox="[236,1369,516,1805]" captionTargetId="figure@6.[200,1396,481,1850]" captionTargetPageId="6" captionText="FIGURE 2. Pygoda variegata sp. n. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D— Female, genital plates (Scale = 1mm)." httpUri="https://zenodo.org/record/1460032/files/figure.png" pageId="7" pageNumber="212">Fig. 2D</figureCitation>
). Gonocoxites 8 with lateral margin almost straight and posterior margin slightly curved; punctures concentrated on convex part; sutural border contiguous; gonapophysis 8 not visible. Gonocoxites 9 narrow, distal part straight; punctures black. Laterotergites 8 with black distal margin projected in a black spine that clearly exceeds apex of segment VII. Laterotergites 9 with small part of the base flat; black apices clearly below the level of laterotergites 8. Capsula seminalis globose with processes long, clearly exceeding anterior annular flange, and increasing in length (
<figureCitation id="F5E035FFFFF1C8600089FBA1FD8FFB8F" box="[434,535,1124,1149]" captionStart="FIGURE 11" captionStartId="23.[151,250,1864,1886]" captionText="FIGURE 11. Internal female: A—Hypoxys sp.; B—Pygoda amianta sp. n.; C—Pygoda variegata sp. n.; D—Pygoda poecila sp. n.; E—Pygoda irrorata; F—Pygoda polita; G—Pygoda ramosa sp. n.; H—Pygoda expolita (aaf—anterior annular flange; ch—chitinellipsen; cs—capsula seminalis; dr—ductus receptaculi; gc9—gonocoxite 9; la9—laterotergite 9; paf—posterior annular flange; pc—pars communis; pi—pars intermedialis; tdr—thickening of ductus receptaculi; tvi—thickening of vaginal intima; x—abdominal segment X. Scale = 1mm)." pageId="7" pageNumber="212">Fig. 11C</figureCitation>
). Ductus receptaculi (tdr) long, clearly longer than capsula seminalis.
</paragraph>
</subSubSection>
<subSubSection id="25C17AF1FFF1C86001FCFB42FB2DFB1B" pageId="7" pageNumber="212" type="discussion">
<paragraph id="6D64297AFFF1C86001FCFB42FB2DFB1B" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="212">
<emphasis id="5FAFF568FFF1C86001FCFB42FED7FB53" bold="true" box="[199,335,1159,1184]" pageId="7" pageNumber="212">Comments.</emphasis>
The specimens observed here has the green color (a kind of bluish green) more intense and better preserved than the other species of the genus. This is particularly remarkable on metasternal process that has a green surface, but laterals greenish yellow, as if the structure was painted only on ventral side.
</paragraph>
</subSubSection>
<subSubSection id="25C17AF1FFF1C86001FCFB31FBC4FAFF" box="[199,1116,1268,1293]" pageId="7" pageNumber="212" type="distribution">
<paragraph id="6D64297AFFF1C86001FCFB31FBC4FAFF" blockId="7.[151,1437,151,1293]" box="[199,1116,1268,1293]" pageId="7" pageNumber="212">
<emphasis id="5FAFF568FFF1C86001FCFB31FEC1FAFE" bold="true" box="[199,345,1268,1293]" pageId="7" pageNumber="212">Distribution</emphasis>
(
<figureCitation id="F5E035FFFFF1C8600053FB31FE24FAFF" box="[360,444,1268,1293]" captionStart="FIGURE 14" captionStartId="25.[151,250,2000,2022]" captionText="FIGURE 14. Distributional map of the new species of Pygoda." pageId="7" pageNumber="212">Fig. 14</figureCitation>
).
<collectingCountry id="15CC69EAFFF1C86000EBFB31FDE0FAFF" box="[464,632,1268,1292]" name="Costa Rica" pageId="7" pageNumber="212">COSTA RICA</collectingCountry>
:
<collectingRegion id="AF1FE798FFF1C86003B9FB30FC99FAFF" box="[642,769,1269,1292]" country="Costa Rica" name="Puntarenas" pageId="7" pageNumber="212">Puntarenas</collectingRegion>
.
<collectingRegion id="AF1FE798FFF1C8600230FB31FC1FFAFF" box="[779,903,1268,1292]" country="Panama" name="Panama" pageId="7" pageNumber="212">
<collectingCountry id="15CC69EAFFF1C8600230FB31FC1FFAFF" box="[779,903,1268,1292]" name="Panama" pageId="7" pageNumber="212">PANAMA</collectingCountry>
</collectingRegion>
:
<collectingRegion id="AF1FE798FFF1C86002AAFB31FC69FAFE" box="[913,1009,1268,1293]" country="Panama" name="Chiriqui" pageId="7" pageNumber="212">Chiriquí</collectingRegion>
,
<collectingRegion id="AF1FE798FFF1C86002C7FB30FBC0FAFF" box="[1020,1112,1269,1292]" country="Panama" name="Panama" pageId="7" pageNumber="212">
<collectingCountry id="15CC69EAFFF1C86002C7FB30FBC0FAFF" box="[1020,1112,1269,1292]" name="Panama" pageId="7" pageNumber="212">Panama</collectingCountry>
</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>