215 lines
36 KiB
XML
215 lines
36 KiB
XML
<document id="CA2E0B45277DBDD8F89FD8034A497E1A" ID-DOI="10.11646/zootaxa.5041.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531880" ID-ZooBank="5E82BE60-609F-4287-AC67-D86536FB7686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632751225130" checkinUser="plazi" docAuthor="Adrain, Jonathan M. & Pérez-Peris, Francesc" docDate="2021" docId="D97A87D4FFAE6C64F9C63EAE1856F9E2" docLanguage="en" docName="zootaxa.5041.1.1.pdf" docOrigin="Zootaxa 5041 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5041.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Harebayaspis plurima Adrain & Pérez-Peris 2021, n. sp." docType="treatment" docUuid="72EA6652-6582-4A20-ACB1-A23158987AC3" docUuidSource="ZooBank" docVersion="3" lastPageNumber="37" masterDocId="2543FFACFF8C6C40F9513A281D2AFFEB" masterDocTitle="Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada" masterLastPageNumber="73" masterPageNumber="1" pageNumber="35" updateTime="1699415937727" updateUser="plazi">
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<mods:title id="2F302D65C9991E70A4A0ABFE72FE7C69">Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada</mods:title>
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<mods:name id="1F881E67CC482BC98D58DF16BA47EC62" type="personal">
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<mods:roleTerm id="9C0114CF700E71E442B8EC0547ABCDEE">Author</mods:roleTerm>
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<mods:namePart id="B2EAD111012A47761651B36F2546A0B8">Adrain, Jonathan M.</mods:namePart>
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<mods:nameIdentifier id="7CB9F034C3B24A63D5AC0EB4CF709366" type="ORCID">0000-0002-7000-1311</mods:nameIdentifier>
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<mods:nameIdentifier id="910F05CC7CB8585A2C38B24959834C2C" type="email">jonathan-adrain@uiowa.edu</mods:nameIdentifier>
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<mods:roleTerm id="DDFAC26790BC30DB1FBA152ED55471B2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4C2E1D9D95E4E2BE15B57DDEE88F3B88">Pérez-Peris, Francesc</mods:namePart>
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<mods:nameIdentifier id="B95A32AEFCB37653494FB0E3E90A4CDC" type="ORCID">0000-0002-7000-1311</mods:nameIdentifier>
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<mods:nameIdentifier id="115CEDD6A853889DDC9ED3E87B1B9661" type="email">jonathan-adrain@uiowa.edu</mods:nameIdentifier>
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<mods:typeOfResource id="E318A51C0A8DFBCD490089D708209A3F">text</mods:typeOfResource>
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<mods:title id="E0C71AA5D7D7F18CE726326BB516FAD7">Zootaxa</mods:title>
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<mods:part id="26B4E9EFBA8806ED07B837BC81CE1EBA">
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<mods:date id="50E070A28B69542A350C26ABA14BFBFD">2021</mods:date>
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<mods:number id="CEC2D1FA0E99879577C55834F7B240AF">2021-09-24</mods:number>
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<mods:detail id="5E852C13AB5A43AE8D424D5FA857E6CD" type="volume">
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<mods:number id="916F1973789B49C52E8861ECB6467E45">5041</mods:number>
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<mods:detail id="484B932C1888147FFA86C709A51CC31A" type="issue">
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<mods:number id="0A9A0A9B36CC37BACE7DEBDF298343DF">1</mods:number>
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<mods:start id="4A3C4B11C1DABAD8CFAC82043DCEA1FF">1</mods:start>
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<mods:url id="49A6C89FF21ED792572CBC842C0E951C">http://dx.doi.org/10.11646/zootaxa.5041.1.1</mods:url>
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<mods:classification id="B631AB166FD3D7C7E9C5D852EBA310FE">journal article</mods:classification>
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<mods:identifier id="92309A0E375AEB0E78D12FB3262AE966" type="DOI">10.11646/zootaxa.5041.1.1</mods:identifier>
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<mods:identifier id="771A29336D08B5C629A257D40F442745" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="102A384A5986A050F95F97D8BEF10040" type="Zenodo-Dep">5531880</mods:identifier>
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<mods:identifier id="A34EE71034E317A632A6114AFB8B4A8D" type="ZooBank">5E82BE60-609F-4287-AC67-D86536FB7686</mods:identifier>
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<treatment id="D97A87D4FFAE6C64F9C63EAE1856F9E2" ID-DOI="http://doi.org/10.5281/zenodo.5531923" ID-Zenodo-Dep="5531923" LSID="urn:lsid:zoobank.org:act:72EA6652-6582-4A20-ACB1-A23158987AC3" httpUri="http://treatment.plazi.org/id/D97A87D4FFAE6C64F9C63EAE1856F9E2" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
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<subSubSection id="19C96549FFAE6C62F9C63EAE1CC7FB4A" box="[151,493,1158,1185]" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="516C36C2FFAE6C62F9C63EAE1CC7FB4A" blockId="34.[151,493,1158,1221]" box="[151,493,1158,1185]" pageId="34" pageNumber="35">
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<heading id="0A2481AEFFAE6C62F9C63EAE1CC7FB4A" bold="true" box="[151,493,1158,1185]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
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<emphasis id="63A7EAD0FFAE6C62F9C63EAE1CC7FB4A" bold="true" box="[151,493,1158,1185]" pageId="34" pageNumber="35">
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<taxonomicName id="96D34D41FFAE6C62F9C63EAE1C89FB4A" authority="Adrain & Pérez-Peris, 2021" authorityName="Adrain & Pérez-Peris" authorityYear="2021" box="[151,419,1158,1185]" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="plurima" status="sp. nov.">
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<emphasis id="63A7EAD0FFAE6C62F9C63EAE1C89FB4A" bold="true" box="[151,419,1158,1185]" italics="true" pageId="34" pageNumber="35">Harebayaspis plurima</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="789457ABFFAE6C62F8FA3EAF1CC7FB4A" box="[427,493,1159,1185]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="19C96549FFAE6C62F9C63E831C02FB2E" box="[151,296,1195,1221]" pageId="34" pageNumber="35" type="description">
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<paragraph id="516C36C2FFAE6C62F9C63E831C02FB2E" blockId="34.[151,493,1158,1221]" box="[151,296,1195,1221]" pageId="34" pageNumber="35">
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<figureCitation id="C9E82A47FFAE6C62F9C63E831C02FB2E" box="[151,296,1195,1221]" captionStart-0="PLATE 13" captionStart-1="PLATE 14" captionStart-2="PLATE 15" captionStart-3="PLATE 16" captionStartId-0="66.[151,234,1944,1968]" captionStartId-1="68.[151,234,1944,1968]" captionStartId-2="70.[151,234,1944,1968]" captionStartId-3="72.[151,234,1944,1968]" captionTargetBox-0="[151,1435,181,1920]" captionTargetBox-1="[151,1435,181,1920]" captionTargetBox-2="[151,1435,181,1920]" captionTargetBox-3="[151,1435,181,1920]" captionTargetId-0="figure-27@66.[151,1435,181,1920]" captionTargetId-1="figure-27@68.[151,1435,181,1920]" captionTargetId-2="figure-27@70.[151,1435,181,1920]" captionTargetId-3="figure-27@72.[151,1435,181,1920]" captionTargetPageId-0="66" captionTargetPageId-1="68" captionTargetPageId-2="70" captionTargetPageId-3="72" captionText-0="PLATE 13" captionText-1="PLATE 14" captionText-2="PLATE 15" captionText-3="PLATE 16" figureDoi-0="http://doi.org/10.5281/zenodo.5531933" figureDoi-1="http://doi.org/10.5281/zenodo.5531935" figureDoi-2="http://doi.org/10.5281/zenodo.5531937" figureDoi-3="http://doi.org/10.5281/zenodo.5531939" httpUri-0="https://zenodo.org/record/5531933/files/figure.png" httpUri-1="https://zenodo.org/record/5531935/files/figure.png" httpUri-2="https://zenodo.org/record/5531937/files/figure.png" httpUri-3="https://zenodo.org/record/5531939/files/figure.png" pageId="34" pageNumber="35">Plates 13–16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="19C96549FFAE6C62F9C63EDA1ED8FA92" pageId="34" pageNumber="35" type="materials_examined">
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<paragraph id="516C36C2FFAE6C62F9C63EDA1ED8FA92" blockId="34.[151,1437,1266,2013]" pageId="34" pageNumber="35">
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<emphasis id="63A7EAD0FFAE6C62F9C63EDA1C2CFAE7" bold="true" box="[151,262,1266,1292]" pageId="34" pageNumber="35">Material.</emphasis>
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<materialsCitation id="E1BB3C9FFFAE6C62F8413EDB1ED8FA92" collectionCode="GSC" pageId="34" pageNumber="35" specimenCode="GSC 135277, GSC 135256, 135257, 135259, 135260, 135261, 135264-135268, 135274, 135275, GSC 135258, 135262, 135273, 135276, 135279, GSC 135255, 135263, 135269, 135271, 135272, 135278, GSC 135270" specimenCount="1" typeStatus="holotype">
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<typeStatus id="8E688860FFAE6C62F8413EDB1C51FAE6" box="[272,379,1267,1293]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
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, pygidium,
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<specimenCode id="01759EB9FFAE6C62FB523EDB1FBDFAE6" box="[515,663,1266,1293]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">GSC 135277</specimenCode>
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(Pl. 16, figs 8, 9, 13, 17, 18), from horizon TCM 24, and assigned specimens
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<specimenCode id="01759EB9FFAE6C62F8453F3F1C83FADA" box="[276,425,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">GSC 135256</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62F8E43F3E1F26FADA" box="[437,524,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135257</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FB493F3E1F45FADA" box="[536,623,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135259</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FB2A3F3E1FF8FADA" box="[635,722,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135260</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FB8F3F3E1E1FFADA" box="[734,821,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135261</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FA103F3E1ED0FADA" box="[833,1018,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135264–135268</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FD543F3E1976FADA" box="[1029,1116,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135274</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FD393F3E1996FADA" box="[1128,1212,1302,1329]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135275</specimenCode>
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from horizon TCM 24,
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<specimenCode id="01759EB9FFAE6C62F9913F131C7EFABE" box="[192,340,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">GSC 135258</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62F80F3F121C9FFABE" box="[350,437,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135262</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62F8913F121F3DFABE" box="[448,535,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135273</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FB703F121F52FABE" box="[545,632,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135276</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FBD33F121FFCFABE" box="[642,726,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135279</specimenCode>
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from horizon TCM 18,
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<specimenCode id="01759EB9FFAE6C62FAB03F13195EFABE" box="[993,1140,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">GSC 135255</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FD2E3F1219FCFABE" box="[1151,1238,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135263</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FDB13F12181DFABE" box="[1248,1335,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135269</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62FC103F1218B2FABE" box="[1345,1432,1338,1365]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135271</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62F9C63F761DC4FA92" box="[151,238,1374,1401]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135272</specimenCode>
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,
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<specimenCode id="01759EB9FFAE6C62F9A83F761C67FA92" box="[249,333,1374,1401]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">135278</specimenCode>
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from locality TCL 2, and
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<specimenCode id="01759EB9FFAE6C62FB233F771E2EFA92" box="[626,772,1374,1401]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="34" pageNumber="35">GSC 135270</specimenCode>
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from locality TCL 6.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="19C96549FFAE6C62F9963FAB198CFA76" box="[199,1190,1411,1437]" pageId="34" pageNumber="35" type="etymology">
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<paragraph id="516C36C2FFAE6C62F9963FAB198CFA76" blockId="34.[151,1437,1266,2013]" box="[199,1190,1411,1437]" pageId="34" pageNumber="35">
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<emphasis id="63A7EAD0FFAE6C62F9963FAB1C64FA76" bold="true" box="[199,334,1411,1437]" pageId="34" pageNumber="35">Etymology.</emphasis>
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Latin
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<emphasis id="63A7EAD0FFAE6C62F8C63FAB1CD1FA76" box="[407,507,1411,1437]" italics="true" pageId="34" pageNumber="35">plurimus</emphasis>
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: most, in reference to the high number of pygidial segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="19C96549FFAE6C62F9963F8E18A1FA0E" pageId="34" pageNumber="35" type="diagnosis">
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<paragraph id="516C36C2FFAE6C62F9963F8E18A1FA0E" blockId="34.[151,1437,1266,2013]" pageId="34" pageNumber="35">
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<emphasis id="63A7EAD0FFAE6C62F9963F8E1C68FA2B" bold="true" box="[199,322,1446,1472]" pageId="34" pageNumber="35">Diagnosis.</emphasis>
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Eye set close to glabella, eye ridge short; fixigena abaxial to palpebral lobe narrow and turned posteriorly; librigenal field with large caecal pits and robustly inflated caecal trunks; pygidium with seven segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="19C96549FFAE6C64F9963FC61E0DFB4A" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
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<paragraph id="516C36C2FFAE6C63F9963FC61FDCFA92" blockId="34.[151,1437,1266,2013]" lastBlockId="35.[151,1437,150,2013]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<emphasis id="63A7EAD0FFAE6C62F9963FC61C73F9E3" bold="true" box="[199,345,1518,1544]" pageId="34" pageNumber="35">Description.</emphasis>
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Cranidial measurements are based on the specimens of
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<figureCitation id="C9E82A47FFAE6C62FA8F3FC71917F9E2" box="[990,1085,1519,1545]" captionStart="PLATE 13" captionStartId="66.[151,234,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-27@66.[151,1435,181,1920]" captionTargetPageId="66" captionText="PLATE 13" figureDoi="http://doi.org/10.5281/zenodo.5531933" httpUri="https://zenodo.org/record/5531933/files/figure.png" pageId="34" pageNumber="35">Plate 13</figureCitation>
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||
, figures 2, 17. Cranidium subtrapezoidal in outline, broad (tr.) with maximum width across genal angles (excluding genal spines), tapering forward, slightly vaulted medially, in anterior view fixigenal field downturned abaxially, in lateral view subtly sloping downwards from the horizontal anteriorly; anterior border describing a broad arch directed forward with the medial portion opposite to the frontal lobe of the glabella almost transverse, medially and in most distal region shorter (sag., exsag.), maximum length (sag.) opposite intersection of S3 with axial furrow (e.g., Pl. 13, figs 2, 3), in anterior view medial part slightly vaulted dorsally (e.g., Pl. 13, fig. 5), gradually flattening distally, with sculpture of densely and evenly distributed tiny granules; anterior border furrow short (sag.), deeply incised, describing an anteriorly directed arch opposite glabella, with the median part almost transverse, distally beyond the glabella running slightly posteriorly, slightly anteriorly convex; glabella subrectangular in outline, with sagittal length 98.85% (96.2–101.5%) maximum glabellar width across L1, lateral margins subparallel, subtly tapering forward, anterolateral corners slightly rounded, posterolateral corners more squared off, anterior margin describing anterior arch with the medial part almost transverse, posterior margin with medial part transverse and distally slightly concave anteriorly across L1, anterior part slightly vaulted, posterior part flatter, sloped downward from the horizontal anterior to S1 (e.g., Pl. 13, fig. 5), with sculpture of dense medium-sized tubercles (larger than on the anterior border) arranged in arcs flanking the glabellar furrows (e.g., Pl. 13, figs 1, 2); S1–S3 well defined, deeply incised, relatively short (exsag.), extending about three quarters of the glabellar width; S1 slightly longer (sag.) than S2/S3, deeply incised distally and shallower proximally, directed posteromedially with the most proximal part curving more posteriorly, not connected with SO, proximal tip followed by a shallow dorsal depression that connects with SO and separates L1 from the median glabellar part; S2 parallel to S1, similar width and depth to S1; S3 shorter than S1/S2, strongly directed backwards; SO deeply incised, slightly shallower medially, long (sag.) behind median glabellar area, shorter abaxially; LO subrectangular in outline, relatively long in medial part behind median glabellar area, slightly shorter laterally, anterior margin transverse, slightly anteriorly concave medially, with distal part running slightly forward, posterior margin describing a broad posteriorly directed arch, corners squared off with the anterior corner slightly more extended outwardly than posterior corner, sculpture similar to that of the glabella; L1 wedge-shaped, slightly longer (exsag.) than L2, posterior margin subtly bowed posteriorly, dorsally inflated compared with the median area of the glabella (e.g., Pl. 13, fig. 11); L2 and L3 subrectangular in outline, similar in size, slightly shorter (exsag.) than L1; frontal lobe subtrapezoidal in outline, broader anteriorly, tapering posteriorly, slightly shorter than L1–L3; axial furrow deep and narrow, similar to the glabellar furrows; fixigenal field subtriangular in outline, with sculpture of dense irregular caecal pits of medium size and small tubercles distributed evenly across the surface; anterior fixigena shorter (exsag.) and narrower (tr.) than posterior part, in anterior view almost transverse to the horizontal, divided by eye ridge into anterior and posterior part (e.g., Pl. 13, figs 1, 2, 3, 10); anterior part very small with same width as the rest of anterior fixigena, very short (sag.), shorter laterally; posterior part of the anterior fixigena triangular in outline, anterior margin anterolaterally concave; posterior part of the fixigena subrectangular in outline, wider than long, anterior margin directed slightly posteriorly, in anterior view proximal part almost transverse to the horizontal and distal part downturned; posterior border furrow narrow deeply incised, short (sag., exsag.), transversely directed adaxial to genal angle, then abruptly curved anteriorly and slightly laterally; lateral border furrow with similar width and depth as the posterior border furrow, directed anterolaterally; posterior border shorter (exsag.) opposite L1, lengthening gradually abaxially until maximum length at genal angle, distally the genal angle is abruptly curved, anterior margin transverse, posterior margin slightly running posteriorly; genal spine conical in shape, directed posterolaterally, tapering distally, ending in a pointed tip (e.g., Pl. 15, figs 12, 13, 15, 16); lateral border short (exsag.) and narrow (tr.), directed anteriorly and slightly laterally, wider near the genal angle, narrowing forward; posterior border, genal spine and lateral border share a sculpture of densely distributed tiny tubercles; palpebral lobe small, posterior end set opposite L2, anterior end continuous with a well defined eye ridge (e.g., Pl. 13, figs 3, 6, 10), eye ridge running adaxially and anteriorly, connected proximally to axial furrow just posterior to the intersection of S3 with axial furrow; palpebral furrow running anteriorly from the posterior end of the palpebral lobe, merged anteriorly with a fine furrow running along the posterior side of the eye ridge; doublure beneath LO short (sag.). approximately half the sagittal length of LO; very short (exsag.), transverse doublure beneath the posterior border, slightly expanded under the genal angle.
|
||
</paragraph>
|
||
<paragraph id="516C36C2FFAF6C63F9963FAB181BF8EE" blockId="35.[151,1437,150,2013]" pageId="35" pageNumber="36">
|
||
Librigenal measurements were made on the specimens of
|
||
<figureCitation id="C9E82A47FFAF6C63FA053FAB1E98FA76" box="[852,946,1411,1437]" captionStart="PLATE 14" captionStartId="68.[151,234,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-27@68.[151,1435,181,1920]" captionTargetPageId="68" captionText="PLATE 14" figureDoi="http://doi.org/10.5281/zenodo.5531935" httpUri="https://zenodo.org/record/5531935/files/figure.png" pageId="35" pageNumber="36">Plate 14</figureCitation>
|
||
, figures 18, 19, 20. Librigena narrow, elongate; eye small, set in the middle of the librigena, separated from field by smooth, weakly inflated socle lacking sculpture; librigenal field triangular in outline, elongate and narrow, width of field 36.4% (31.6–40.7%) length along lateral border furrow; in lateral view field slopes downward from the base of the eye to border furrow; field with sculpture of small to medium caecal pits and anastomosing caecal trunks, with tiny tubercles scattered irregularly over trunks; border furrow deep, narrow, slightly bowed laterally; border narrow, with width 57.1% (50.0–66.7%) width of field, outer margin gently curved laterally; posterior projection slightly longer than anterior projection, slightly curved medially in external view; anterior projection curved medially; posterior facial suture long curving convexly laterally, changing slope opposite border furrow from slightly curved laterally through the border to adaxially (e.g., Pl. 14, fig. 18); anterior facial suture short, curved laterally, with a gentle slope opposite border furrow; lateral border doublure very narrow beneath the entire border (e.g., Pl. 14, figs 21, 22), lacking sculpture.
|
||
</paragraph>
|
||
<paragraph id="516C36C2FFAF6C63F9963D271FF0F8C2" blockId="35.[151,1437,150,2013]" box="[199,730,1807,1833]" pageId="35" pageNumber="36">Rostral plate, hypostome, and thorax unknown.</paragraph>
|
||
<paragraph id="516C36C2FFAF6C64F9963D1B1E0DFB4A" blockId="35.[151,1437,150,2013]" lastBlockId="36.[151,1437,151,1545]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
Pygidial measurements were made on the specimens of
|
||
<figureCitation id="C9E82A47FFAF6C63FA013D1B1E9AF8A6" box="[848,944,1842,1869]" captionStart="PLATE 15" captionStartId="70.[151,234,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-27@70.[151,1435,181,1920]" captionTargetPageId="70" captionText="PLATE 15" figureDoi="http://doi.org/10.5281/zenodo.5531937" httpUri="https://zenodo.org/record/5531937/files/figure.png" pageId="35" pageNumber="36">Plate 15</figureCitation>
|
||
, figure 1, and
|
||
<figureCitation id="C9E82A47FFAF6C63FD0D3D1B1990F8A6" box="[1116,1210,1843,1869]" captionStart="PLATE 16" captionStartId="72.[151,234,1944,1968]" captionTargetBox="[151,1435,181,1920]" captionTargetId="figure-27@72.[151,1435,181,1920]" captionTargetPageId="72" captionText="PLATE 16" figureDoi="http://doi.org/10.5281/zenodo.5531939" httpUri="https://zenodo.org/record/5531939/files/figure.png" pageId="35" pageNumber="36">Plate 16</figureCitation>
|
||
figure 9. Pygidium approximately semicircular in outline, broader anteriorly across the first pleural segment, narrowing posteriorly, sagittal length (excluding articulating half-ring and pleural spines) 45.5% (40.3–45.5%) maximum width (excluding pleural spines), anterior part slightly vaulted medially; fulcrum set far from the axis (e.g., Pl. 16, fig. 18), pleurae slightly downturned distal to the fulcrum, almost transverse; articulating half-ring short, longer sagittally with sagittal length 10.0% (8.9–11.1%) pygidial length, shorter exsagittally, anterior margin broadly bowed anteriorly, posterior almost transverse, vaulted dorsally, smooth; articulating furrow short (sag., exsag.), longer sagittally, slightly shorter exsagittally, shallow, deepening exsagittally; axis composed of seven axial rings, narrow and elongate, with sagittal length 147.6% (142.3–152.9%) maximum width across first axial ring, slightly tapering posteriorly, slightly vaulted anteriorly, gradually flattening posteriorly (e.g., Pl. 16, fig. 8); axial rings with sculpture of evenly distributed medium-sized tubercles; first six axial rings similar in shape, subrectangular in outline, gradually reducing in length (sag.) and width (tr.) from the first to the sixth segment; each of first six segments slightly shorter sagittally, longer exsagittally, anterior margin slightly bowed posteriorly, posterior margin slightly bowed anteriorly, corners squared off; seventh axial ring semicircular in outline, longer sagittally, much shorter exsagittally, anterior margin transverse, posterior posteriorly bowed; terminal piece absent; six inter-ring furrows, decreasing gradually in length (sag.) and width (tr.) from anterior to posterior, longer sagittally, shorter exsagittally, shallow sagittally, slightly deeper exsagittally; axial furrow with width similar to length of articulating furrow, forming wider areas in the intersections with inter-ring furrows, pleural furrows and interpleural furrows, deeply incised anteriorly, shallowing posteriorly; first pleural segment divided by a short (exsag.) and deep pleural furrow; pleural furrow with transverse course, distal part slightly curved posteriorly (e.g., Pl. 16, fig. 9), anterior pleural band shorter (exsag.) than posterior pleural band, transverse, distal tip slightly curved posteriorly, extended slightly beyond the pygidial margin; posterior pleural band long (sag.), running transversely, curving posteriorly before reaching pygidial margin, slightly longer (exsag.) across the pygidial margin, extended beyond pygidial margin by a pleural spine; pleural spine running posterolaterally, conical in outline, circular in cross-section (e.g., Pl. 16, fig. 17); second to fifth pleural segments with similar morphology, divided in anterior and posterior band by a pleural furrow as the first pleural segment, pleural spines more posteriorly directed in the more posterior segments; sixth and seventh pleural segments with pleural furrow and anterior pleural band obscure, segments strongly directed backwards; pleural segments separated by six interpleural furrows similar in lenght (sag.), width (tr.) and depth to pleural furrows, forming a wider area distally in the intersection with the pleural furrows (e.g., Pl. 16, figs 9, 14); posteriorly, interpleural furrows become gradually narrower (tr.), ending distally in the pygidial margin, first interpleural furrow running almost transversely, curving posteriorly slightly distally, posterior furrows directed more strongly posteriorly, shallowing distally across the pygidial margin; pleural segments with dense sculpture of medium-sized tubercles, similar to axis; pygidial margin forming continuous ventral rim (e.g., Pl. 15, fig. 2, Pl. 16, fig. 13); doublure very short, set vertically above margin (e.g., Pl. 15, fig. 4, pl. 16, fig. 18).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="19C96549FFA86C64F9963E821856F9E2" pageId="36" pageNumber="37" type="discussion">
|
||
<paragraph id="516C36C2FFA86C64F9963E821856F9E2" blockId="36.[151,1437,151,1545]" pageId="36" pageNumber="37">
|
||
<emphasis id="63A7EAD0FFA86C64F9963E821C66FB2F" bold="true" box="[199,332,1194,1220]" pageId="36" pageNumber="37">Discussion.</emphasis>
|
||
<taxonomicName id="96D34D41FFA86C64F8073E831F79FB2E" authorityName="Adrain & Pérez-Peris" authorityYear="2021" box="[342,595,1195,1221]" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="plurima">
|
||
<emphasis id="63A7EAD0FFA86C64F8073E831F79FB2E" box="[342,595,1195,1221]" italics="true" pageId="36" pageNumber="37">Harebayaspis plurima</emphasis>
|
||
</taxonomicName>
|
||
is most similar among known species to
|
||
<taxonomicName id="96D34D41FFA86C64FD663E8319A9FB2E" authorityName="Edgecombe, Chatterton, Vaccari, and Waisfeld" authorityYear="1999" box="[1079,1155,1195,1221]" class="Trilobita" family="Pilekiidae" genus="Macrogrammus" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rafi">
|
||
<emphasis id="63A7EAD0FFA86C64FD663E8319A9FB2E" box="[1079,1155,1195,1221]" italics="true" pageId="36" pageNumber="37">H. rafi</emphasis>
|
||
</taxonomicName>
|
||
, particularly in cephalic features. The eyes are similarly close to the glabella, with relatively short eye ridges. The portion of the fixigena abaxial to the palpebral lobe is quite narrow. In both
|
||
<taxonomicName id="96D34D41FFA86C64FBB03EDB1EAFFAE6" authorityName="Fortey" authorityYear="1980" box="[737,901,1267,1293]" class="Trilobita" family="Pilekiidae" genus="Parapilekia" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="jacquelinae">
|
||
<emphasis id="63A7EAD0FFA86C64FBB03EDB1EAFFAE6" box="[737,901,1267,1293]" italics="true" pageId="36" pageNumber="37">H. jacquelinae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="96D34D41FFA86C64FAEB3EDB1930FAE7" authorityName="Edgecombe, Chatterton, Vaccari, and Waisfeld" authorityYear="1999" box="[954,1050,1267,1292]" class="Trilobita" family="Pilekiidae" genus="Macrogrammus" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pengi">
|
||
<emphasis id="63A7EAD0FFA86C64FAEB3EDB1930FAE7" box="[954,1050,1267,1292]" italics="true" pageId="36" pageNumber="37">H. pengi</emphasis>
|
||
</taxonomicName>
|
||
the eye is set further abaxially, the eye ridge is longer, and the abaxial portion of the fixigena is much wider in dorsal view (in large part because it is directed more laterally instead of ventrally). Librigenae of
|
||
<taxonomicName id="96D34D41FFA86C64FA763F131E88FABE" authorityName="Adrain & Pérez-Peris" authorityYear="2021" box="[807,930,1339,1365]" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="plurima">
|
||
<emphasis id="63A7EAD0FFA86C64FA763F131E88FABE" box="[807,930,1339,1365]" italics="true" pageId="36" pageNumber="37">H. plurima</emphasis>
|
||
</taxonomicName>
|
||
have almost the same dimensions as those of
|
||
<taxonomicName id="96D34D41FFA86C64F9C63F771DC8FA92" authorityName="Edgecombe, Chatterton, Vaccari, and Waisfeld" authorityYear="1999" box="[151,226,1375,1401]" class="Trilobita" family="Pilekiidae" genus="Macrogrammus" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rafi">
|
||
<emphasis id="63A7EAD0FFA86C64F9C63F771DC8FA92" box="[151,226,1375,1401]" italics="true" pageId="36" pageNumber="37">H. rafi</emphasis>
|
||
</taxonomicName>
|
||
, and both differ from the wider field seen in
|
||
<taxonomicName id="96D34D41FFA86C64FBB73F771E6DFA93" authorityName="Edgecombe, Chatterton, Vaccari, and Waisfeld" authorityYear="1999" box="[742,839,1375,1400]" class="Trilobita" family="Pilekiidae" genus="Macrogrammus" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pengi">
|
||
<emphasis id="63A7EAD0FFA86C64FBB73F771E6DFA93" box="[742,839,1375,1400]" italics="true" pageId="36" pageNumber="37">H. pengi</emphasis>
|
||
</taxonomicName>
|
||
. The librigenae differ in the larger caecal pits developed on the field in
|
||
<taxonomicName id="96D34D41FFA86C64F8263FAB1CD8FA76" authorityName="Adrain & Pérez-Peris" authorityYear="2021" box="[375,498,1411,1437]" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="plurima">
|
||
<emphasis id="63A7EAD0FFA86C64F8263FAB1CD8FA76" box="[375,498,1411,1437]" italics="true" pageId="36" pageNumber="37">H. plurima</emphasis>
|
||
</taxonomicName>
|
||
. The species differ most obviously in the presence of seven pygidial segments in
|
||
<taxonomicName id="96D34D41FFA86C64FCD03FAB1DDAFA2A" authorityName="Adrain & Pérez-Peris" authorityYear="2021" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="plurima">
|
||
<emphasis id="63A7EAD0FFA86C64FCD03FAB1DDAFA2A" italics="true" pageId="36" pageNumber="37">H. plurima</emphasis>
|
||
</taxonomicName>
|
||
versus four in
|
||
<taxonomicName id="96D34D41FFA86C64F8C63F8F1CCBFA2A" authorityName="Edgecombe, Chatterton, Vaccari, and Waisfeld" authorityYear="1999" box="[407,481,1447,1473]" class="Trilobita" family="Pilekiidae" genus="Macrogrammus" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rafi">
|
||
<emphasis id="63A7EAD0FFA86C64F8C63F8F1CCBFA2A" box="[407,481,1447,1473]" italics="true" pageId="36" pageNumber="37">H. rafi</emphasis>
|
||
</taxonomicName>
|
||
. Even here, though, comparison of just the rear four pygidial segments of
|
||
<taxonomicName id="96D34D41FFA86C64FC703F8F18B7FA2A" authorityName="Adrain & Pérez-Peris" authorityYear="2021" box="[1313,1437,1447,1473]" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="plurima">
|
||
<emphasis id="63A7EAD0FFA86C64FC703F8F18B7FA2A" box="[1313,1437,1447,1473]" italics="true" pageId="36" pageNumber="37">H. plurima</emphasis>
|
||
</taxonomicName>
|
||
with the entire pygidium of
|
||
<taxonomicName id="96D34D41FFA86C64F8833FE31F36FA0E" authorityName="Edgecombe, Chatterton, Vaccari, and Waisfeld" authorityYear="1999" box="[466,540,1483,1509]" class="Trilobita" family="Pilekiidae" genus="Macrogrammus" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="rafi">
|
||
<emphasis id="63A7EAD0FFA86C64F8833FE31F36FA0E" box="[466,540,1483,1509]" italics="true" pageId="36" pageNumber="37">H. rafi</emphasis>
|
||
</taxonomicName>
|
||
reveals substantial similarity, with faint paired axial tubercles and similar spine positions. Though they are broken in many specimens, the spines are apparently relatively longer in
|
||
<taxonomicName id="96D34D41FFA86C64FDAB3FC7185FF9E2" authorityName="Adrain & Pérez-Peris" authorityYear="2021" box="[1274,1397,1519,1545]" class="Trilobita" family="Pliomeridae" genus="Harebayaspis" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="plurima">
|
||
<emphasis id="63A7EAD0FFA86C64FDAB3FC7185FF9E2" box="[1274,1397,1519,1545]" italics="true" pageId="36" pageNumber="37">H. plurima</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |