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<document id="690A75C0F41DDE792090AEFD0500F831" ID-DOI="10.11646/zootaxa.4039.1.1" ID-GBIF-Dataset="4ecc5394-2569-4cbd-8790-a3a4301d304a" ID-ISSN="1175-5326" ID-Zenodo-Dep="289932" ID-ZooBank="CB081EA2-3F93-4FD3-8AC8-B24F772833FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461319549927" checkinUser="plazi" docAuthor="Plant, Adrian R." docDate="2015" docId="D96487F9992BFF9007E94FE8ACA08709" docLanguage="en" docName="zt04039p056.pdf" docOrigin="Zootaxa 4039 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemerodromia systoechon Plant, 2015, sp. nov." docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="255DFF819901FFBB077E4E17A842800A" masterDocTitle="Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?" masterLastPageNumber="56" masterPageNumber="1" pageNumber="43" updateTime="1698618296215" updateUser="plazi">
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<mods:title id="4974FB7519F205B92A64FC5B04173778">Diversity of Hemerodromia Meigen, 1822 (Diptera: Empididae) in Thailand, the tip of a tropical iceberg?</mods:title>
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<mods:namePart id="DE8B86BC45A84F2588C01178F7FCB497">Plant, Adrian R.</mods:namePart>
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<mods:date id="967DFE731AFB857CF38C84FA4C4926B9">2015</mods:date>
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<mods:identifier id="0199BC7C6CF79F17922DDFA45D42A1D4" type="DOI">10.11646/zootaxa.4039.1.1</mods:identifier>
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<treatment id="D96487F9992BFF9007E94FE8ACA08709" ID-DOI="http://doi.org/10.5281/zenodo.6121245" ID-GBIF-Taxon="127693977" ID-Zenodo-Dep="6121245" LSID="urn:lsid:plazi:treatment:D96487F9992BFF9007E94FE8ACA08709" httpUri="http://treatment.plazi.org/id/D96487F9992BFF9007E94FE8ACA08709" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection id="19D76564992BFF9107E94FE8A92E8231" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="517236EF992BFF9107E94FE8AA7D8210" blockId="42.[151,575,511,571]" box="[151,575,511,538]" pageId="42" pageNumber="43">
<heading id="0A3A8183992BFF9107E94FE8AA7D8210" bold="true" box="[151,575,511,538]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="63B9EAFD992BFF9107E94FE8AA7D8210" bold="true" box="[151,575,511,538]" pageId="42" pageNumber="43">
<taxonomicName id="96CD4D6C992BFF9107E94FE8A9988213" ID-CoL="9HVC2" box="[151,474,511,537]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="systoechon" status="sp. nov.">
<emphasis id="63B9EAFD992BFF9107E94FE8A9988213" bold="true" box="[151,474,511,537]" italics="true" pageId="42" pageNumber="43">Hemerodromia systoechon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788A5786992BFF91069F4C17AA7D8210" box="[481,575,512,538]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="517236EF992BFF9107E94C35A92E8231" blockId="42.[151,575,511,571]" box="[151,364,546,571]" pageId="42" pageNumber="43">
(
<figureCitation id="C9F62A6A992BFF9107E14C35A8A58230" box="[159,231,546,571]" captionStart="FIGURES 6 8" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[166,1424,199,1893]" captionTargetId="figure@8.[151,1435,193,1905]" captionTargetPageId="8" captionText="FIGURES 6 8. Habitus of Hemerodromia spp. 6, H. attenuata sp. nov. ♂; 7, H. systoechon sp. nov. ♂; 8, H. songsee sp. nov. ♀." httpUri="https://zenodo.org/record/289933/files/figure.png" pageId="42" pageNumber="43">Figs 7</figureCitation>
,
<figureCitation id="C9F62A6A992BFF91078D4C35A9798231" box="[243,315,546,571]" captionStart="FIGURES 66 67" captionStartId="43.[151,264,948,971]" captionTargetBox="[159,1418,214,914]" captionTargetId="figure@43.[152,1435,193,927]" captionTargetPageId="43" captionText="FIGURES 66 67. Male terminalia of Hemerodromia systoechon sp. nov. 66, lateral view of terminalia; 67, dorsal view of cerci. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium." httpUri="https://zenodo.org/record/289956/files/figure.png" pageId="42" pageNumber="43">66, 67</figureCitation>
,
<figureCitation id="C9F62A6A992BFF9106394C35A9268231" box="[327,356,546,571]" captionStart="FIGURES 91 98" captionStartId="49.[151,264,1887,1910]" captionTargetBox="[281,1307,208,1839]" captionTargetId="figure@49.[264,1331,193,1856]" captionTargetPageId="49" captionText="FIGURES 91 98. Distribution of Hemerodromia spp. in Thailand at a resolution of 0.25 ° of latitude and longitude with symbols scaled to 0.5 °. 91, H. ocellata sp. nov.; 92, H. oriens sp. nov.; 93, H. phahompokensis sp. nov.; 94, H. songsee sp. nov.; 95, H. systoechon sp. nov.; 96, H. yunnanensis Yang &amp; Yang; 97, H. zetalutea sp. nov.; 98, Species richness of Hemerodromia spp. in Thailand at a resolution of 1 ° with symbols plotted at same resolution." httpUri="https://zenodo.org/record/289961/files/figure.png" pageId="42" pageNumber="43">95</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="19D76564992BFF9107E94C7DAC6882F8" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="517236EF992BFF9107E94C7DAC6882F8" blockId="42.[151,1437,618,2028]" pageId="42" pageNumber="43">
<emphasis id="63B9EAFD992BFF9107E94C7DA9488289" bold="true" box="[151,266,618,643]" pageId="42" pageNumber="43">Diagnosis</emphasis>
. Scutum black with lateral margins narrowly and postpronotal area yellowish; Prothoracic collar dorsally, scutellum and mediotergite black; pleura yellowish but sutures about laterotergite and base of halter blackish. Wing with cell bm+dm long, ending distinctly beyond level of R1; distance between ends of R2+3 and R4 long, ~1.8X long as R4. Male cercus with distinctly spinose process on inner face.
</paragraph>
</subSubSection>
<subSubSection id="19D76564992BFF9107B94D17A8BE87E6" pageId="42" pageNumber="43" type="description">
<paragraph id="517236EF992BFF9107B94D17A8BE87E6" blockId="42.[151,1437,618,2028]" pageId="42" pageNumber="43">
<emphasis id="63B9EAFD992BFF9107B94D17A9138313" bold="true" box="[199,337,768,793]" pageId="42" pageNumber="43">Description</emphasis>
.
<emphasis id="63B9EAFD992BFF9106204D17A9D98313" bold="true" box="[350,411,768,793]" pageId="42" pageNumber="43">Male</emphasis>
: Body length
<quantity id="96359B0A992BFF9105474D17AA818312" box="[569,707,768,793]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="42" pageNumber="43" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5 mm</quantity>
; wing length
<quantity id="96359B0A992BFF9104264D17ABA08312" box="[856,994,768,792]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="42" pageNumber="43" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
.
<emphasis id="63B9EAFD992BFF9104924D17AC698313" bold="true" box="[1004,1067,768,793]" pageId="42" pageNumber="43">Head</emphasis>
. Black, dusted greyish, strongest on upper occiput; antenna and mouthparts yellow; all setae whitish; one pair reclinate ocl; postocular setae uniseriate, merging with two pairs small vtl; a few very fine setulae behind vtl; frontal setulae apparently absent. Antenna with postpedicel 1.5X long as wide, stylus of similar length; scape with distinct fine dorsal seta.
<emphasis id="63B9EAFD992BFF9102424D7BADD7838F" bold="true" box="[1340,1429,876,901]" pageId="42" pageNumber="43">Thorax</emphasis>
. Dusted greyish. Scutum with ground colour brown or brownish black, usually with markedly darker area posterolaterally immediately above root of wing (viewed laterodorsally, appearing somewhat triangular with posterior apex darker); lateral margin distinctly yellowish, becoming wider anteriorly to include postpronotal area, demarcation between brown and yellow parts well defined; posterior margins laterally, narrowly diffuse yellowish. Prothoracic collar dorsally, scutellum and mediotergite black. Pleura yellow (
<figureCitation id="C9F62A6A992BFF9103424A37ACC48432" box="[1084,1158,1056,1081]" captionStart="FIGURES 6 8" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[166,1424,199,1893]" captionTargetId="figure@8.[151,1435,193,1905]" captionTargetPageId="8" captionText="FIGURES 6 8. Habitus of Hemerodromia spp. 6, H. attenuata sp. nov. ♂; 7, H. systoechon sp. nov. ♂; 8, H. songsee sp. nov. ♀." httpUri="https://zenodo.org/record/289933/files/figure.png" pageId="42" pageNumber="43">Fig. 7</figureCitation>
), fused anepisternum + katepisternum and meron sometimes vaguely darkened ventrally; sutures about laterotergite and base of halter blackish. All setae yellowish; minute apart from a pair of distinct sct and one smaller npl.
<emphasis id="63B9EAFD992BFF9103E94A7FAC8D848B" bold="true" box="[1175,1231,1128,1153]" pageId="42" pageNumber="43">Legs</emphasis>
. Yellowish white, apical tarsal segment hardly darker. C1 ~1.2X longer than distance between C1 and C2, all setulae minute excepting a few slightly longer apicals. F1 ~ 1.2X long as C1, ~ 4
<date id="2573102F992BFF9104E34AA7ABEB84C3" box="[925,937,1200,1225]" pageId="42" pageNumber="43">5</date>
X long as wide, slightly constricted 0.20.4 from base; femoral formula ~ 57/1820+12/1620/67, denticles black, rows converging apically; spines yellow, longest basally with several additional distinct spines at base. T1 ~ 0.7X long as F1, evenly curved, ventral face shallowly concave; with one row of ~ 1620 sharply pointed spinose setae ventrally; a very small bluntly pointed ventroapical spur with strong ventroapical erect black spinose seta arising from near its base. Mid and hind legs slender with only small setulae but mid metatarsus with somewhat longer setulae ventrally, T3 with a short row of small setae posterodorsally and a comb of short setae posteroapically.
<emphasis id="63B9EAFD992BFF91032F4B9FACD085AB" bold="true" box="[1105,1170,1416,1441]" pageId="42" pageNumber="43">Wing</emphasis>
. Membrane with dark microtrichia but hyaline basally. Veins dark but whitish near base and C yellowish brown with posterior margin very narrowly black. Marginal setulae pale yellow. R2+3 slightly curved, joining C ~ 0.60.7X distance between end of R1 and R4; length of C between ends of R2+3 and R4 quite long, ~1.8X long as R4. R4 sublinear, only slightly Sshaped, angle with R5 at extreme base ~ 70°; R5 ~2.02.2X long as R4; R4+5 fork distal to M1+2 fork by ~
<date id="2573102F992BFF9102524809AD0C863C" box="[1324,1358,1566,1590]" pageId="42" pageNumber="43">1X</date>
length of R4; R5 and M1 slightly curved, convergent distally; cell bm+dm long, ending distinctly beyond level of R1. Halter pale yellowish white.
<emphasis id="63B9EAFD992BFF9106E3487CAA53868E" bold="true" box="[413,529,1643,1668]" pageId="42" pageNumber="43">Abdomen</emphasis>
. Tergites black with tergite 1 yellow; vent blackish becoming progressively yellowish basally on sternites 13; tergites with only indistinct setulae on posterior margins, but long brown setae on tergite 8 posteriorly. Sternites with setae on posterior margins brownish, becoming longer on posterior segments where also present on disc.
<emphasis id="63B9EAFD992BFF91069548C0AA2C86FA" bold="true" box="[491,622,1751,1776]" pageId="42" pageNumber="43">Terminalia</emphasis>
. Black with dark setae. Cercus elongate, extending as far as tip of epandrium in lateral view (
<figureCitation id="C9F62A6A992BFF9106AB48ECAA6F871E" box="[469,557,1787,1812]" captionStart="FIGURES 66 67" captionStartId="43.[151,264,948,971]" captionTargetBox="[159,1418,214,914]" captionTargetId="figure@43.[152,1435,193,927]" captionTargetPageId="43" captionText="FIGURES 66 67. Male terminalia of Hemerodromia systoechon sp. nov. 66, lateral view of terminalia; 67, dorsal view of cerci. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium." httpUri="https://zenodo.org/record/289956/files/figure.png" pageId="42" pageNumber="43">Fig. 66</figureCitation>
); broad proximally in dorsal view (
<figureCitation id="C9F62A6A992BFF9104B148ECAC65871E" box="[975,1063,1787,1812]" captionStart="FIGURES 66 67" captionStartId="43.[151,264,948,971]" captionTargetBox="[159,1418,214,914]" captionTargetId="figure@43.[152,1435,193,927]" captionTargetPageId="43" captionText="FIGURES 66 67. Male terminalia of Hemerodromia systoechon sp. nov. 66, lateral view of terminalia; 67, dorsal view of cerci. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium." httpUri="https://zenodo.org/record/289956/files/figure.png" pageId="42" pageNumber="43">Fig. 67</figureCitation>
); distinctly setose on outer face distally. Viewed dorsally an elongate inner process apparently originating from inner face of cercus bearing 56 short strong spines. Epandrium elongate, bluntly pointed apically, with 12 strong setae subapically and numerous smaller setae on distal 0.5. Hypandrium elongate hemispherical, with distinct setae posteroventrally.
<emphasis id="63B9EAFD992BFF9102434970ADD6878A" bold="true" box="[1341,1428,1895,1920]" pageId="42" pageNumber="43">Female</emphasis>
: Similar to male but F1 without additional yellow spines at base or spines indistinct; abdomen more extensively yellowish proximally on vent, often only darkened on sternites 6 laterally and 7 laterally and medially. Terminalia blackish.
</paragraph>
</subSubSection>
<caption id="05B26667992AFF9007E94DA3AB2083E3" httpUri="https://zenodo.org/record/289956/files/figure.png" pageId="43" pageNumber="44" targetBox="[159,1418,214,914]" targetPageId="43">
<paragraph id="517236EF992AFF9007E94DA3AB2083E3" blockId="43.[151,1435,948,1001]" pageId="43" pageNumber="44">
<emphasis id="63B9EAFD992AFF9007E94DA3A90D83C3" bold="true" box="[151,335,948,971]" pageId="43" pageNumber="44">FIGURES 6667</emphasis>
. Male terminalia of
<taxonomicName id="96CD4D6C992AFF9005574DA3AB7B83C0" box="[553,825,948,970]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="systoechon" status="sp. nov.">
<emphasis id="63B9EAFD992AFF9005574DA3AB7B83C0" box="[553,825,948,970]" italics="true" pageId="43" pageNumber="44">Hemerodromia systoechon</emphasis>
</taxonomicName>
<emphasis id="63B9EAFD992AFF90043C4DA2ABF583C3" bold="true" box="[834,951,948,970]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="788A5786992AFF90043C4DA2ABD783C0" box="[834,917,949,970]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
66
</emphasis>
, lateral view of terminalia;
<emphasis id="63B9EAFD992AFF9003A34DA3ACB483C3" bold="true" box="[1245,1270,948,969]" pageId="43" pageNumber="44">67</emphasis>
, dorsal view of cerci. Abbreviations: cer = cerci, epa = epandrium, hyp = hypandrium.
</paragraph>
</caption>
<subSubSection id="19D76564992AFF9007B94A02A9408578" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="517236EF992AFF9007B94A02A9408578" blockId="43.[151,1437,1045,1796]" pageId="43" pageNumber="44">
<emphasis id="63B9EAFD992AFF9007B94A02A92F8424" bold="true" box="[199,365,1045,1070]" pageId="43" pageNumber="44">
<typeStatus id="8E76884D992AFF9007B94A02A9408424" box="[199,258,1045,1070]" pageId="43" pageNumber="44">Type</typeStatus>
material
</emphasis>
.
<emphasis id="63B9EAFD992AFF9006054A02AA588424" bold="true" box="[379,538,1045,1070]" pageId="43" pageNumber="44">
<typeStatus id="8E76884D992AFF9006054A02AA588424" box="[379,538,1045,1070]" pageId="43" pageNumber="44" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
♂,
<collectingCountry id="29DA767F992AFF90053D4A01AA9A8424" box="[579,728,1046,1070]" name="Thailand" pageId="43" pageNumber="44">THAILAND</collectingCountry>
: Chiang Mai Province, Doi Inthanon NP, Summit Marsh,
<quantity id="96359B0A992AFF9002234A02A8ED845B" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="m" value="2500.0">2,500 m</quantity>
,
<geoCoordinate id="34F95028992AFF9007C44A2EA9088458" box="[186,330,1081,1106]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.58935">18°35.361'N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF90062B4A2EA9A08458" box="[341,482,1081,1106]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="98.48595">98°29.157'E</geoCoordinate>
,
<date id="2573102F992AFF9006934A2EAAFC8458" box="[493,702,1081,1106]" pageId="43" pageNumber="44" value="2006-07-22" valueMax="2006-08-02" valueMin="2006-07-22">22.vii.2.viii.2006</date>
, MT, Y. Areeluck [T118](
<collectionCode id="37DCAE2A992AFF9004A34A2DAC6D8458" box="[989,1071,1082,1106]" pageId="43" pageNumber="44">QSBG</collectionCode>
).
<emphasis id="63B9EAFD992AFF90033F4A2EACA78458" bold="true" box="[1089,1253,1081,1106]" pageId="43" pageNumber="44">
<typeStatus id="8E76884D992AFF90033F4A2EACA78458" box="[1089,1253,1081,1106]" pageId="43" pageNumber="44" type="paratype">PARATYPES</typeStatus>
</emphasis>
:
<specimenCount id="47CBFD66992AFF90038A4A2DAD5B8458" box="[1268,1305,1082,1106]" pageId="43" pageNumber="44" type="generic">2♂</specimenCount>
, 15♀, same data as
<typeStatus id="8E76884D992AFF9007934A4AA910847C" box="[237,338,1117,1142]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
,
<date id="2573102F992AFF9006204A4AAA62847C" box="[350,544,1117,1142]" pageId="43" pageNumber="44" value="2006-06-29" valueMax="2006-07-02" valueMin="2006-06-29">29.vi.2.vii.2006</date>
,
<date id="2573102F992AFF9005534A4AAA9B847C" box="[557,729,1117,1142]" pageId="43" pageNumber="44" value="2006-07-15" valueMax="2006-07-22" valueMin="2006-07-15">1522.vii.2006</date>
,
<date id="2573102F992AFF90059B4A4AAB3F847C" box="[741,893,1117,1142]" pageId="43" pageNumber="44" value="2006-08-02" valueMax="2006-08-09" valueMin="2006-08-02">29.viii.2006</date>
,
<date id="2573102F992AFF9004F44A4AAC6F847C" box="[906,1069,1117,1142]" pageId="43" pageNumber="44" value="2007-04-15" valueMax="2007-04-22" valueMin="2007-04-15">1522.iv.2007</date>
,
<date id="2573102F992AFF9003474A4AACAA847C" box="[1081,1256,1117,1142]" pageId="43" pageNumber="44" value="2007-04-29" valueMax="2007-05-06" valueMin="2007-04-29">29.iv.6.v.2007</date>
,
<date id="2573102F992AFF90038A4A4AADC0847C" box="[1268,1410,1117,1142]" pageId="43" pageNumber="44" value="2007-05-08" valueMax="2007-05-15" valueMin="2007-05-08">815.v.2007</date>
;
<specimenCount id="47CBFD66992AFF9002F04A49A8EC8493" pageId="43" pageNumber="44" type="generic">2 ♂</specimenCount>
, Summit Forest,
<quantity id="96359B0A992AFF90060B4A96A9978493" box="[373,469,1153,1178]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="43" pageNumber="44" unit="m" value="2500.0">2,500 m</quantity>
,
<geoCoordinate id="34F95028992AFF90069D4A96AA318490" box="[483,627,1153,1178]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.58935">18°35.361'N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF9005FF4A96AB4C8490" box="[641,782,1153,1178]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="98.48595">98°29.157'E</geoCoordinate>
,
<date id="2573102F992AFF9004624A96ABFC8490" box="[796,958,1153,1178]" pageId="43" pageNumber="44" value="2007-04-15" valueMax="2007-04-22" valueMin="2007-04-15">1522.iv.2007</date>
, MT, Y. Areeluck;
<specimenCount id="47CBFD66992AFF9003E34A95AC808490" box="[1181,1218,1154,1178]" pageId="43" pageNumber="44" type="generic">2♂</specimenCount>
, 5♀, Kew Maepan Trail,
<quantity id="96359B0A992AFF90079D4AB1A90884B7" box="[227,330,1190,1214]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="43" pageNumber="44" unit="m" value="2200.0">2,200 m</quantity>
,
<geoCoordinate id="34F95028992AFF9006204AB2A9AD84B4" box="[350,495,1189,1214]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.5527">18°33.162'N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF90057D4AB1AAC084B4" box="[515,642,1190,1214]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="9" value="98.48016">98°28.81'E</geoCoordinate>
,
<date id="2573102F992AFF9005E84AB2AB7984B4" box="[662,827,1189,1214]" pageId="43" pageNumber="44" value="2006-09-21" valueMax="2006-09-27" valueMin="2006-09-21">2127.ix.2006</date>
,
<date id="2573102F992AFF9004314AB2ABBF84B4" box="[847,1021,1189,1214]" pageId="43" pageNumber="44" value="2007-04-29" valueMax="2007-05-06" valueMin="2007-04-29">29.iv.6.v.2007</date>
,
<date id="2573102F992AFF90036C4AB1ACD284B4" box="[1042,1168,1190,1214]" pageId="43" pageNumber="44" value="2007-05-01" valueMax="2007-05-08" valueMin="2007-05-01">18.v.2007</date>
,
<date id="2573102F992AFF9003DB4AB2AD7084B4" box="[1189,1330,1189,1214]" pageId="43" pageNumber="44" value="2007-05-08" valueMax="2007-05-15" valueMin="2007-05-08">815.v.2007</date>
, MT, Y. Areeluck;
<specimenCount id="47CBFD66992AFF90066C4ADDA97584E8" box="[274,311,1226,1250]" pageId="43" pageNumber="44" type="generic">1♂</specimenCount>
, Checkpoint 2,
<quantity id="96359B0A992AFF9006844ADDAA1C84EB" box="[506,606,1226,1250]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="43" pageNumber="44" unit="m" value="1700.0">1,700 m</quantity>
,
<geoCoordinate id="34F95028992AFF90050E4ADEAB4284E8" box="[624,768,1225,1250]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.525984">18°31.559'N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF90046F4ADDABDC84E8" box="[785,926,1226,1250]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="98.499016">98°29.941'E</geoCoordinate>
,
<date id="2573102F992AFF9004D14ADEAC0F84E8" box="[943,1101,1225,1250]" pageId="43" pageNumber="44" value="2006-07-08" valueMax="2006-07-15" valueMin="2006-07-08">815.vii.2006</date>
, MT, Y. Areeluck;
<specimenCount id="47CBFD66992AFF9002454ADEAD2284E8" box="[1339,1376,1225,1250]" pageId="43" pageNumber="44" type="generic">6♂</specimenCount>
, 7♀, Summit trap3 (Angka),
<quantity id="96359B0A992AFF9006DB4AFAAA46850F" box="[421,516,1261,1286]" metricMagnitude="3" metricUnit="m" metricValue="2.545" pageId="43" pageNumber="44" unit="m" value="2545.0">2,545 m</quantity>
,
<geoCoordinate id="34F95028992AFF90056E4AFAAAEE850C" box="[528,684,1261,1286]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.588917">18°35'20.1&quot;N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF9005C64AFAAB12850C" box="[696,848,1261,1286]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="98.4848">98°29'05.3&quot;E</geoCoordinate>
,
<date id="2573102F992AFF9004224AFAABBF850C" box="[860,1021,1261,1286]" pageId="43" pageNumber="44" value="2014-04-01" valueMax="2014-05-02" valueMin="2014-04-01">1.iv.2.v.2014</date>
,
<date id="2573102F992AFF9003774AF9ACD4850C" box="[1033,1174,1262,1286]" pageId="43" pageNumber="44" value="2014-05-02" valueMax="2014-05-29" valueMin="2014-05-02">229.v.2014</date>
, MT, W. Srisuka
<emphasis id="63B9EAFD992AFF9002254AF9ADCF850C" box="[1371,1421,1261,1286]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
.; 1♀, Summit trap1 (back of office),
<quantity id="96359B0A992AFF9005594B06AAC48523" box="[551,646,1297,1322]" metricMagnitude="3" metricUnit="m" metricValue="2.534" pageId="43" pageNumber="44" unit="m" value="2534.0">2,534 m</quantity>
,
<geoCoordinate id="34F95028992AFF9005ED4B06AB6C8520" box="[659,814,1297,1322]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.58689">18°35'12.8&quot;N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF9004444B05AB908520" box="[826,978,1298,1322]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="98.487274">98°29'14.2&quot;E</geoCoordinate>
,
<date id="2573102F992AFF9004A14B06ACD48520" box="[991,1174,1297,1322]" pageId="43" pageNumber="44" value="2014-05-29" valueMax="2014-07-01" valueMin="2014-05-29">29.v.1.vii.2014</date>
, MT, W. Srisuka
<emphasis id="63B9EAFD992AFF9002224B04ADCF8520" box="[1372,1421,1297,1322]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
.; 1 ♀, Checkpoint 2 trap1,
<quantity id="96359B0A992AFF9006BA4B22AA648547" box="[452,550,1333,1358]" metricMagnitude="3" metricUnit="m" metricValue="1.639" pageId="43" pageNumber="44" unit="m" value="1639.0">1,639 m</quantity>
,
<geoCoordinate id="34F95028992AFF90054B4B22AA928544" box="[565,720,1333,1358]" direction="north" orientation="latitude" pageId="43" pageNumber="44" precision="1" value="18.52764">18°31'39.5&quot;N</geoCoordinate>
,
<geoCoordinate id="34F95028992AFF9005A14B22AB358544" box="[735,887,1333,1358]" direction="east" orientation="longitude" pageId="43" pageNumber="44" precision="1" value="98.499916">98°29'59.7&quot;E</geoCoordinate>
,
<date id="2573102F992AFF9004F84B21AC518544" box="[902,1043,1334,1358]" pageId="43" pageNumber="44" value="2014-05-02" valueMax="2014-05-29" valueMin="2014-05-02">229.v.2014</date>
, MT, W. Srisuka
<emphasis id="63B9EAFD992AFF90039C4B21AD578544" box="[1250,1301,1333,1358]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
. (
<collectionCode id="37DCAE2A992AFF9002514B21AD3E8544" box="[1327,1404,1334,1358]" pageId="43" pageNumber="44">QSBG</collectionCode>
&amp;
<collectionCode id="37DCAE2A992AFF9007E94B4DA8B58578" box="[151,247,1370,1394]" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="43" pageNumber="44">NMWC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="19D76564992AFF9007B94B6AABC285B0" pageId="43" pageNumber="44" type="etymology">
<paragraph id="517236EF992AFF9007B94B6AABC285B0" blockId="43.[151,1437,1045,1796]" pageId="43" pageNumber="44">
<emphasis id="63B9EAFD992AFF9007B94B6AA904859C" bold="true" box="[199,326,1405,1430]" pageId="43" pageNumber="44">Etymology</emphasis>
. The specific epithet is from the Greek
<taxonomicName id="96CD4D6C992AFF90046A4B6AABCD859C" box="[788,911,1405,1430]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="systoechon">
<emphasis id="63B9EAFD992AFF90046A4B6AABCD859C" box="[788,911,1405,1430]" italics="true" pageId="43" pageNumber="44">systoechon</emphasis>
</taxonomicName>
(single row) in reference to the single row of ventral setulae on the front tibia. It is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="19D76564992AFF9007B94BD2ACA08709" pageId="43" pageNumber="44" type="discussion">
<paragraph id="517236EF992AFF9007B94BD2ACA08709" blockId="43.[151,1437,1045,1796]" pageId="43" pageNumber="44">
<emphasis id="63B9EAFD992AFF9007B94BD2A97685D4" bold="true" box="[199,308,1477,1502]" pageId="43" pageNumber="44">Remarks</emphasis>
. The thoracic colour pattern of this species and
<taxonomicName id="96CD4D6C992AFF9004364BD0ACB285D4" authority="Grootaert, Yang &amp; Saigusa" authorityName="Grootaert, Yang &amp; Saigusa" box="[840,1264,1477,1502]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutata">
<emphasis id="63B9EAFD992AFF9004364BD0ABFC85D4" box="[840,958,1479,1502]" italics="true" pageId="43" pageNumber="44">H. acutata</emphasis>
Grootaert, Yang &amp; Saigusa
</taxonomicName>
is quite similar in some individuals, especially in females of the latter species in which the scutellum is dark. In such cases
<taxonomicName id="96CD4D6C992AFF9002FF4BFCA950862C" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="systoechon" status="sp. nov.">
<emphasis id="63B9EAFD992AFF9002FF4BFCA950862C" italics="true" pageId="43" pageNumber="44">H. systoechon</emphasis>
</taxonomicName>
<emphasis id="63B9EAFD992AFF900667481AA931862C" bold="true" box="[281,371,1549,1574]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="788A5786992AFF900667481AA931862C" box="[281,371,1549,1574]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="96CD4D6C992AFF9005D84818AB5F862C" box="[678,797,1551,1574]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutata">
<emphasis id="63B9EAFD992AFF9005D84818AB5F862C" box="[678,797,1551,1574]" italics="true" pageId="43" pageNumber="44">H. acutata</emphasis>
</taxonomicName>
by having cell bm+dm long, obviously reaching beyond the apex of R1 (level with R
<quantity id="96359B0A992AFF9006B54857A9AD8640" box="[459,495,1585,1614]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="43" pageNumber="44" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="96CD4D6C992AFF90068B4824AA298640" box="[501,619,1587,1610]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutata">
<emphasis id="63B9EAFD992AFF90068B4824AA298640" box="[501,619,1587,1610]" italics="true" pageId="43" pageNumber="44">H. acutata</emphasis>
</taxonomicName>
) and length of C between ends of R2+3 and R4 quite long, about 1.8X long as R4. Both
<taxonomicName id="96CD4D6C992AFF90065F484EA9DB867A" box="[289,409,1625,1648]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutata">
<emphasis id="63B9EAFD992AFF90065F484EA9DB867A" box="[289,409,1625,1648]" italics="true" pageId="43" pageNumber="44">H. acutata</emphasis>
</taxonomicName>
and
<taxonomicName id="96CD4D6C992AFF9006AA484EAA31867B" box="[468,627,1624,1649]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="systoechon" status="sp. nov.">
<emphasis id="63B9EAFD992AFF9006AA484EAA31867B" box="[468,627,1624,1649]" italics="true" pageId="43" pageNumber="44">H. systoechon</emphasis>
</taxonomicName>
<emphasis id="63B9EAFD992AFF900502484FAA9A867B" bold="true" box="[636,728,1624,1649]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="788A5786992AFF900502484FAA9A867B" box="[636,728,1624,1649]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have a single row of denticles ventrally on the front tibia.
<taxonomicName id="96CD4D6C992AFF9002FE484EA9508692" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="systoechon" status="sp. nov.">
<emphasis id="63B9EAFD992AFF9002FE484EA9508692" italics="true" pageId="43" pageNumber="44">H. systoechon</emphasis>
</taxonomicName>
<emphasis id="63B9EAFD992AFF9006674868A9308692" bold="true" box="[281,370,1663,1688]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="788A5786992AFF9006674868A9308692" box="[281,370,1663,1688]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the mountain Doi Inthanon, Chiang Mai Province, northern
<collectingCountry id="29DA767F992AFF9003824868AD228692" box="[1276,1376,1663,1688]" name="Thailand" pageId="43" pageNumber="44">Thailand</collectingCountry>
(
<figureCitation id="C9F62A6A992AFF9002114868A8F586B6" captionStart="FIGURES 91 98" captionStartId="49.[151,264,1887,1910]" captionTargetBox="[281,1307,208,1839]" captionTargetId="figure@49.[264,1331,193,1856]" captionTargetPageId="49" captionText="FIGURES 91 98. Distribution of Hemerodromia spp. in Thailand at a resolution of 0.25 ° of latitude and longitude with symbols scaled to 0.5 °. 91, H. ocellata sp. nov.; 92, H. oriens sp. nov.; 93, H. phahompokensis sp. nov.; 94, H. songsee sp. nov.; 95, H. systoechon sp. nov.; 96, H. yunnanensis Yang &amp; Yang; 97, H. zetalutea sp. nov.; 98, Species richness of Hemerodromia spp. in Thailand at a resolution of 1 ° with symbols plotted at same resolution." httpUri="https://zenodo.org/record/289961/files/figure.png" pageId="43" pageNumber="44">Fig. 95</figureCitation>
). 95% of specimens were captured at
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in moist hill evergreen forest or
<taxonomicName id="96CD4D6C992AFF90031248B4AD5386B6" box="[1132,1297,1699,1724]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="43" pageNumber="44" phylum="Tracheophyta" rank="genus">
<emphasis id="63B9EAFD992AFF90031248B4AD5386B6" box="[1132,1297,1699,1724]" italics="true" pageId="43" pageNumber="44">Rhododendron</emphasis>
</taxonomicName>
/
<taxonomicName id="96CD4D6C992AFF90025848B4ADDE86B6" box="[1318,1436,1699,1724]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="43" pageNumber="44" phylum="Bryophyta" rank="genus">
<emphasis id="63B9EAFD992AFF90025848B4ADDE86B6" box="[1318,1436,1699,1724]" italics="true" pageId="43" pageNumber="44">Sphagnum</emphasis>
</taxonomicName>
biotopes close to the summit. All specimens have been taken between April and September (mostly AprilJuly) thus adult phenology correlates with the late hot dry period and Southwest Monsoon rainy season.
</paragraph>
</subSubSection>
</treatment>
</document>