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<mods:title id="66BCA2E581ED6EADFB00C8ABF189D671">DNA barcode and phylogeography of six new high altitude wingless Niphadomimus (Coleoptera: Curculionidae: Molytinae) from Southwest China</mods:title>
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<treatment id="D44787FEFFECFFDB53D127DB82CEF98C" ID-DOI="http://doi.org/10.5281/zenodo.5129171" ID-GBIF-Taxon="183731286" ID-Zenodo-Dep="5129171" LSID="urn:lsid:plazi:treatment:D44787FEFFECFFDB53D127DB82CEF98C" httpUri="http://treatment.plazi.org/id/D44787FEFFECFFDB53D127DB82CEF98C" lastPageId="5" lastPageNumber="155" pageId="4" pageNumber="155">
<subSubSection id="14F46563FFECFFDA53D127DB8301FA0C" box="[151,544,1474,1500]" pageId="4" pageNumber="155" type="nomenclature">
<paragraph id="5C5136E8FFECFFDA53D127DB8301FA0C" blockId="4.[151,544,1474,1500]" box="[151,544,1474,1500]" pageId="4" pageNumber="155">
<heading id="07198184FFECFFDA53D127DB8301FA0C" bold="true" box="[151,544,1474,1500]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
<taxonomicName id="9BEE4D6BFFECFFDA53D127DB8301FA0C" ID-CoL="8HN4C" authority="Zherikhin, 1987" authorityName="Zherikhin" authorityYear="1987" box="[151,544,1474,1500]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="6E9AEAFAFFECFFDA53D127DB8301FA0C" bold="true" box="[151,544,1474,1500]" pageId="4" pageNumber="155">
<emphasis id="6E9AEAFAFFECFFDA53D127DB8070FA0C" bold="true" box="[151,337,1474,1500]" italics="true" pageId="4" pageNumber="155">Niphadomimus</emphasis>
<bibRefCitation id="387F4B19FFECFFDA521E27DB8301FA0C" author="Zherikhin, V. V." box="[344,544,1474,1500]" pageId="4" pageNumber="155" pagination="1 - 43" refId="ref13814" refString="Zherikhin, V. V. (1987) Curculionidae from the Nepal Himalayas. Part 1. Molytinae (Insecta: Coleoptera). Stuttgarter Beitraege zur Naturkunde, Serie A (Biologie), 411, 1 - 43." type="journal article" year="1987">Zherikhin, 1987</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="14F46563FFECFFDA53D1241080B9F9EE" pageId="4" pageNumber="155" type="reference_group">
<paragraph id="5C5136E8FFECFFDA53D1241080B9F9EE" blockId="4.[151,1436,1545,1598]" pageId="4" pageNumber="155">
<bibRefCitation id="387F4B19FFECFFDA53D124108048F9CF" author="Zherikhin, V. V." box="[151,361,1545,1567]" pageId="4" pageNumber="155" pagination="1 - 43" refId="ref13814" refString="Zherikhin, V. V. (1987) Curculionidae from the Nepal Himalayas. Part 1. Molytinae (Insecta: Coleoptera). Stuttgarter Beitraege zur Naturkunde, Serie A (Biologie), 411, 1 - 43." type="journal article" year="1987">Zherikhin, 1987: 15</bibRefCitation>
(species included:
<taxonomicName id="9BEE4D6BFFECFFDA517C24138391F9CF" ID-CoL="47FT5" authorityName="Zherikhin" authorityYear="1987" box="[570,688,1546,1567]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="6E9AEAFAFFECFFDA517C24138391F9CF" box="[570,688,1546,1567]" italics="true" pageId="4" pageNumber="155">nigriventris</emphasis>
</taxonomicName>
,
<taxonomicName id="9BEE4D6BFFECFFDA5184241383D6F9CF" ID-CoL="47FT4" authorityName="Zherikhin" authorityYear="1987" box="[706,759,1546,1567]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="6E9AEAFAFFECFFDA5184241383D6F9CF" box="[706,759,1546,1567]" italics="true" pageId="4" pageNumber="155">niger</emphasis>
</taxonomicName>
).
<typeStatus id="8355884AFFECFFDA505724138264F9CF" box="[785,837,1546,1567]" pageId="4" pageNumber="155">Type</typeStatus>
species:
<taxonomicName id="9BEE4D6BFFECFFDA50ED24108459F9CF" ID-CoL="47FT5" authority="Zherikhin, 1987" authorityName="Zherikhin" authorityYear="1987" box="[939,1400,1545,1567]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="6E9AEAFAFFECFFDA50ED241085E4F9CF" box="[939,1221,1545,1567]" italics="true" pageId="4" pageNumber="155">Niphadomimus nigriventris</emphasis>
<bibRefCitation id="387F4B19FFECFFDA579624108459F9CF" author="Zherikhin, V. V." box="[1232,1400,1545,1567]" pageId="4" pageNumber="155" pagination="1 - 43" refId="ref13814" refString="Zherikhin, V. V. (1987) Curculionidae from the Nepal Himalayas. Part 1. Molytinae (Insecta: Coleoptera). Stuttgarter Beitraege zur Naturkunde, Serie A (Biologie), 411, 1 - 43." type="journal article" year="1987">Zherikhin, 1987</bibRefCitation>
</taxonomicName>
by original designation.
</paragraph>
</subSubSection>
<subSubSection id="14F46563FFECFFDA53D1247985D2F861" pageId="4" pageNumber="155" type="diagnosis">
<paragraph id="5C5136E8FFECFFDA53D124798416F899" blockId="4.[151,1437,1632,2038]" pageId="4" pageNumber="155">
<emphasis id="6E9AEAFAFFECFFDA53D124798030F9A8" box="[151,273,1632,1657]" inLineHeading="true" pageId="4" pageNumber="155" reason="1">
<emphasis id="6E9AEAFAFFECFFDA53D12479802BF9A9" bold="true" box="[151,266,1632,1657]" pageId="4" pageNumber="155">Diagnosis</emphasis>
.
</emphasis>
Adults of the genus
<taxonomicName id="9BEE4D6BFFECFFDA52BA24798384F9A9" authorityName="Zherikhin" authorityYear="1987" box="[508,677,1632,1657]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="6E9AEAFAFFECFFDA52BA24798384F9A9" box="[508,677,1632,1657]" italics="true" pageId="4" pageNumber="155">Niphadomimus</emphasis>
</taxonomicName>
can be immediately recognized from other wingless Asian weevils by the following unique combination of characters: (1.) rostrum longer than wide; (2.) prosternum anterior of procoxae with a shallow and broad depression (
<figureCitation id="C4D52A6DFFECFFDA51FF24BC822DF96E" box="[697,780,1701,1726]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="4" pageNumber="155">Fig. 11</figureCitation>
); (3.) mesoventrum at middle anterad of mesocoxae with distinct and sizable projection pointed ventrad, vertically sloped anteriorly and oblique posteriorly (
<figureCitation id="C4D52A6DFFECFFDA57BD24D1846CF931" box="[1275,1357,1736,1761]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="4" pageNumber="155">Fig. 11</figureCitation>
). In its natural habitat
<taxonomicName id="9BEE4D6BFFECFFDA521324F280DEF8D4" authorityName="Zherikhin" authorityYear="1987" box="[341,511,1771,1796]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="6E9AEAFAFFECFFDA521324F280DEF8D4" box="[341,511,1771,1796]" italics="true" pageId="4" pageNumber="155">Niphadomimus</emphasis>
</taxonomicName>
co-occurs with similarly wingless
<taxonomicName id="9BEE4D6BFFECFFDA50F824F285E9F8D4" authority="Schoenherr" authorityName="Schoenherr" box="[958,1224,1771,1796]" class="Insecta" family="Curculionidae" genus="Colobodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="6E9AEAFAFFECFFDA50F824F28517F8D4" box="[958,1078,1771,1796]" italics="true" pageId="4" pageNumber="155">Colobodes</emphasis>
Schoenherr
</taxonomicName>
and
<taxonomicName id="9BEE4D6BFFECFFDA565224F2803DF8F6" authority="Schenkling." authorityName="Schenkling." class="Insecta" family="Curculionidae" genus="Niphadonyx" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="6E9AEAFAFFECFFDA565224F284BAF8D4" box="[1300,1435,1771,1796]" italics="true" pageId="4" pageNumber="155">Niphadonyx</emphasis>
Schenkling.
</taxonomicName>
Adults of these two genera are most easily separated from those of
<taxonomicName id="9BEE4D6BFFECFFDA5769251485F8F8F6" authorityName="Zherikhin" authorityYear="1987" box="[1071,1241,1805,1830]" class="Insecta" family="Curculionidae" genus="Niphadomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="6E9AEAFAFFECFFDA5769251485F8F8F6" box="[1071,1241,1805,1830]" italics="true" pageId="4" pageNumber="155">Niphadomimus</emphasis>
</taxonomicName>
by either widely separate procoxae and or by the claws having a sizable inner lobe (Meregalli 2013, fig. 2M), respectively.
</paragraph>
<paragraph id="5C5136E8FFECFFDA5381254A85D2F861" blockId="4.[151,1437,1632,2038]" pageId="4" pageNumber="155">The original generic description by Zherikhin appears adequate, even though it did not refer to the internal structures, such as the proventiculus, or male and female genitalia. These and a few other characters are mentioned below in an attempt to elaborate the original Zherikhin description, which should also be consulted.</paragraph>
</subSubSection>
<subSubSection id="14F46563FFECFFDB538125A282CEF98C" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" type="description">
<paragraph id="5C5136E8FFECFFDB538125A282CEF98C" blockId="4.[151,1437,1632,2038]" lastBlockId="5.[151,1437,1048,1628]" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
<emphasis id="6E9AEAFAFFECFFDA538125A28078F804" bold="true" box="[199,345,1979,2004]" inLineHeading="true" pageId="4" pageNumber="155" reason="1">Description.</emphasis>
Fully apterous genus of
<taxonomicName id="9BEE4D6BFFECFFDA513825A283D0F804" box="[638,753,1979,2004]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="subFamily" subFamily="Molytinae">Molytinae</taxonomicName>
with elytra fused to each other and to metathorax. Length
<quantity id="9B169B0DFFECFFDA53D125C48065F825" box="[151,324,2013,2038]" metricMagnitude="-3" metricUnit="m" metricValue="4.359999999999999" metricValueMax="6.27" metricValueMin="2.45" pageId="4" pageNumber="155" unit="mm" value="4.359999999999999" valueMax="6.27" valueMin="2.45">2.456.27 mm</quantity>
. Colour dark, from reddish-brown to almost black; legs and elytra with bluish or reddish hue, sometimes partly bicoloured (
<figureCitation id="C4D52A6DFFEDFFDB52B426018351FBE1" box="[498,624,1048,1073]" captionStart="FIGURE 9" captionStartId="12.[151,250,1356,1378]" captionTargetBox="[171,1436,542,1313]" captionTargetId="figure-395@12.[157,1442,537,1318]" captionTargetPageId="12" captionText="FIGURE 9. Niphadomimus nigriventris, female, #4697. AB: habitus; C: sternite 9; D: apical hemisternite 9 (= “stylus”); E: sternite 8." figureDoi="http://doi.org/10.5281/zenodo.4921817" httpUri="https://zenodo.org/record/4921817/files/figure.png" pageId="5" pageNumber="156">Fig. 9AB</figureCitation>
), never metallic or shiny. Antennal funicle with seven antennomeres. Procoxal cavities adjacent to each other (
<figureCitation id="C4D52A6DFFEDFFDB513B262283F2FB84" box="[637,723,1083,1108]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11</figureCitation>
); prosternum anterad of procoxal cavities with depression at middle (
<figureCitation id="C4D52A6DFFEDFFDB53B226448067FBA6" box="[244,326,1117,1142]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11</figureCitation>
), this depression normally in form of wide longitudinal groove marked by ridge on each side (
<figureCitation id="C4D52A6DFFEDFFDB5629264481EBFB49" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11H</figureCitation>
). Ventral surface of thorax and abdomen with numerous punctures (
<figureCitation id="C4D52A6DFFEDFFDB508D2699853FFB49" box="[971,1054,1152,1177]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11</figureCitation>
). Mesoventrum at middle anterad of mesocoxae with distinct and sizable projection pointed ventrad, vertically sloped anteriorly and oblique posteriorly (
<figureCitation id="C4D52A6DFFEDFFDB526026DC805EFB0E" box="[294,383,1221,1246]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11</figureCitation>
). Entire length of meso-metaventral suture between metacoxae formed by deep narrow transverse groove (
<figureCitation id="C4D52A6DFFEDFFDB523626F180E3FAD1" box="[368,450,1256,1281]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11</figureCitation>
) often separated at middle by septum into two equal parts (
<figureCitation id="C4D52A6DFFEDFFDB573726F185EEFAD1" box="[1137,1231,1256,1281]" captionStart="FIGURE 11" captionStartId="14.[151,250,1467,1489]" captionTargetBox="[151,1436,589,1445]" captionTargetId="figure-470@14.[151,1436,589,1445]" captionTargetPageId="14" captionText="FIGURE 11. Niphadomimus spp., pro, meso- and metathorax, ventral (AG, I) and left ventro-lateral (H). A: Niphadomimus alcyone sp. n., holotype, female, #2493; B: N. alcyone sp. n., paratype, female, #4429; C: N. celaeno sp. n., holotype, male, #2476; D: N electra sp. n., paratype, female, #0879; E: N. maia sp. n., holotype, male, #2730; F: N. merope sp. n., holotype, female #2201; G: N. nigriventris, male, #0796; H: N. nigriventris, female, #4697; I: N. sterope sp. n., holotype, male, #4475." figureDoi="http://doi.org/10.5281/zenodo.4921821" httpUri="https://zenodo.org/record/4921821/files/figure.png" pageId="5" pageNumber="156">Fig. 11I</figureCitation>
). All femora with single variously developed femoral tooth at distal third. Scutellum with minute externally visible part. Elytra with nine punctate striae, odd interstriae with our without tubercles, some of which might be markedly enlarged (
<figureCitation id="C4D52A6DFFEDFFDB5629273481FFFAB9" captionStart="FIGURE 4" captionStartId="7.[151,250,1494,1516]" captionTargetBox="[168,1417,183,1457]" captionTargetId="figure-138@7.[160,1420,179,1472]" captionTargetPageId="7" captionText="FIGURE 4. Niphadomimus electra sp. n., holotype, male, #2727. AD: habitus; E: sternites 8 and 9; FH: aedeagus and tegmen, lateral (F), dorsal (G) and ventral (H)." figureDoi="http://doi.org/10.5281/zenodo.4921807" httpUri="https://zenodo.org/record/4921807/files/figure.png" pageId="5" pageNumber="156">Fig. 4AC</figureCitation>
). Proventriculus sclerotized and easily distinguishable (
<figureCitation id="C4D52A6DFFEDFFDB501627498289FAB9" box="[848,936,1360,1385]" captionStart="FIGURE 8" captionStartId="11.[151,250,1928,1950]" captionTargetBox="[159,1436,633,1872]" captionTargetId="figure-173@11.[151,1436,617,1882]" captionTargetPageId="11" captionText="FIGURE 8. Niphadomimus nigriventris, male, #0796. AD: habitus; E: proventriculus; F: sternites 8 and 9; GI: aedeagus and tegmen, lateral (G), dorsal (H) and ventral (I)." figureDoi="http://doi.org/10.5281/zenodo.4921815" httpUri="https://zenodo.org/record/4921815/files/figure.png" pageId="5" pageNumber="156">Fig. 8E</figureCitation>
).
<emphasis id="6E9AEAFAFFEDFFDB50FA2749854EFAB9" bold="true" box="[956,1135,1360,1385]" pageId="5" pageNumber="156">Male genitalia.</emphasis>
Male abdominal sclerite 9 variously asymmetrically V-shaped (
<figureCitation id="C4D52A6DFFEDFFDB5173276A83AAFA5C" box="[565,651,1395,1420]" captionStart="FIGURE 3" captionStartId="6.[151,250,1431,1453]" captionTargetBox="[157,1421,164,1383]" captionTargetId="figure-188@6.[151,1436,151,1403]" captionTargetPageId="6" captionText="FIGURE 3. Niphadomimus celaeno sp. n., holotype, male, #2476. AD: habitus; E: sternites 8 and 9; FH: aedeagus and tegmen, lateral (F), dorsal (G) and ventral (H)." figureDoi="http://doi.org/10.5281/zenodo.4921805" httpUri="https://zenodo.org/record/4921805/files/figure.png" pageId="5" pageNumber="156">Fig. 3E</figureCitation>
), with long apodeme (= “spiculum gastrale”); sternite 8 consisting of two hemisternites (
<figureCitation id="C4D52A6DFFEDFFDB5236278C80E8FA7E" box="[368,457,1429,1454]" captionStart="FIGURE 3" captionStartId="6.[151,250,1431,1453]" captionTargetBox="[157,1421,164,1383]" captionTargetId="figure-188@6.[151,1436,151,1403]" captionTargetPageId="6" captionText="FIGURE 3. Niphadomimus celaeno sp. n., holotype, male, #2476. AD: habitus; E: sternites 8 and 9; FH: aedeagus and tegmen, lateral (F), dorsal (G) and ventral (H)." figureDoi="http://doi.org/10.5281/zenodo.4921805" httpUri="https://zenodo.org/record/4921805/files/figure.png" pageId="5" pageNumber="156">Fig. 3E</figureCitation>
). Aedeagus symmetrical, longer than wide, almost parallel sided and evenly rounded, without internal sclerites and with partly sclerotized walls on endophallus proximad to it (
<figureCitation id="C4D52A6DFFEDFFDB57EF27A18405FA01" box="[1193,1316,1464,1489]" captionStart="FIGURE 3" captionStartId="6.[151,250,1431,1453]" captionTargetBox="[157,1421,164,1383]" captionTargetId="figure-188@6.[151,1436,151,1403]" captionTargetPageId="6" captionText="FIGURE 3. Niphadomimus celaeno sp. n., holotype, male, #2476. AD: habitus; E: sternites 8 and 9; FH: aedeagus and tegmen, lateral (F), dorsal (G) and ventral (H)." figureDoi="http://doi.org/10.5281/zenodo.4921805" httpUri="https://zenodo.org/record/4921805/files/figure.png" pageId="5" pageNumber="156">Fig. 3FH</figureCitation>
); aedeagal apodemes about 1.5x as long as aedeagus; tegmen fully closed dorsally, with two asetose and weakly sclerotized parameral lobes each longer than wide.
<emphasis id="6E9AEAFAFFEDFFDB511827E4820DF9C6" bold="true" box="[606,812,1533,1558]" pageId="5" pageNumber="156">Female genitalia.</emphasis>
Sternite 8 Y-shaped with setose apical lobes (
<figureCitation id="C4D52A6DFFEDFFDB567C27E484B3F9C6" box="[1338,1426,1533,1558]" captionStart="FIGURE 1" captionStartId="4.[151,250,1265,1287]" captionTargetBox="[151,1428,193,1220]" captionTargetId="figure-281@4.[143,1428,187,1221]" captionTargetPageId="4" captionText="FIGURE 1. Niphadomimus alcyone sp. n., holotype, female, #2493. A-D: habitus; E: apical part of basal hemisternite 9 (“coxite”) and much smaller apical cylindrical hemisternite 9 (=“stylus”); F: sternites 8 and IX; G: sternite 8 (sternite 9 in the background)." figureDoi="http://doi.org/10.5281/zenodo.4921801" httpUri="https://zenodo.org/record/4921801/files/figure.png" pageId="5" pageNumber="156">Fig. 1G</figureCitation>
) and long apodeme; sternite 9 consisting of two hemisternites each formed by larger basal piece (= “coxite”) and smaller apical piece (= “stylus”) (
<figureCitation id="C4D52A6DFFEDFFDB5149245A83A5F98C" box="[527,644,1603,1628]" captionStart="FIGURE 1" captionStartId="4.[151,250,1265,1287]" captionTargetBox="[151,1428,193,1220]" captionTargetId="figure-281@4.[143,1428,187,1221]" captionTargetPageId="4" captionText="FIGURE 1. Niphadomimus alcyone sp. n., holotype, female, #2493. A-D: habitus; E: apical part of basal hemisternite 9 (“coxite”) and much smaller apical cylindrical hemisternite 9 (=“stylus”); F: sternites 8 and IX; G: sternite 8 (sternite 9 in the background)." figureDoi="http://doi.org/10.5281/zenodo.4921801" httpUri="https://zenodo.org/record/4921801/files/figure.png" pageId="5" pageNumber="156">Fig. 1E, F</figureCitation>
); spermatheca present (
<figureCitation id="C4D52A6DFFEDFFDB50CD245A82C2F98C" box="[907,995,1603,1628]" captionStart="FIGURE 2" captionStartId="5.[151,250,978,1000]" captionTargetBox="[152,1422,155,935]" captionTargetId="figure-400@5.[149,1434,155,941]" captionTargetPageId="5" captionText="FIGURE 2. Niphadomimus alcyone sp. n., paratype, female, #4429. AD: habitus; E: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4921803" httpUri="https://zenodo.org/record/4921803/files/figure.png" pageId="5" pageNumber="156">Fig. 2E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>