290 lines
39 KiB
XML
290 lines
39 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3792852" ID-GBIF-Dataset="142a25ed-47c3-4361-bc24-6f1b3e8dba5c" ID-GBIF-Taxon="163755037" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792852" ID-ZooBank="664C49F1-5384-43C4-8BF1-CE76AC11D32E" approvalRequired="10" approvalRequired_for_treatments="10" checkinTime="1587998787238" checkinUser="plazi" docAuthor="Klimaszewski, Jan, Savard, Karine, Pelletier, Georges & Webster, Reginald" docDate="2008" docId="D43E034BFFB0415CFF17FB9A6661FE37" docLanguage="en" docName="ZK_article_1943.pdf" docOrigin="ZooKeys 2 (2)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Gnypeta carbonaria" docType="treatment" docVersion="13" lastPageId="48" lastPageNumber="58" masterDocId="28077B33FF9D416CFF9BFFC76718FFEC" masterDocTitle="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution" masterLastPageNumber="84" masterPageNumber="11" pageId="45" pageNumber="56" updateTime="1643574247635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klimaszewski, Jan</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada & C 136 Eddc & A- 86 C</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Savard, Karine</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada & C 136 Eddc & A- 86 C</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pelletier, Georges</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada & C 136 Eddc & A- 86 C</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Webster, Reginald</mods:namePart>
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<mods:affiliation>24 Millstream Drive, Fredericton, NB, Canada & F-</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2008</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2008-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>11</mods:start>
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<mods:end>84</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.2.4</mods:identifier>
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<mods:identifier type="GBIF-Dataset">142a25ed-47c3-4361-bc24-6f1b3e8dba5c</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576410</mods:identifier>
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<mods:identifier type="ZooBank">664C49F1-5384-43C4-8BF1-CE76AC11D32E</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792852" ID-GBIF-Taxon="163755037" ID-Zenodo-Dep="3792852" LSID="urn:lsid:plazi:treatment:D43E034BFFB0415CFF17FB9A6661FE37" httpUri="http://treatment.plazi.org/id/D43E034BFFB0415CFF17FB9A6661FE37" lastPageId="48" lastPageNumber="58" pageId="45" pageNumber="56">
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<subSubSection box="[140,608,1116,1143]" pageId="45" pageNumber="56" type="nomenclature">
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||
<paragraph blockId="45.[140,608,1116,1179]" box="[140,608,1116,1143]" pageId="45" pageNumber="56">
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||
<heading box="[140,608,1116,1143]" fontSize="11" level="2" pageId="45" pageNumber="56" reason="3">
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<emphasis bold="true" box="[140,608,1116,1143]" pageId="45" pageNumber="56">
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13.
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<taxonomicName authority="(Mannerheim)" baseAuthorityName="Mannerheim" baseAuthorityYear="1830" box="[185,608,1116,1143]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="56" phylum="Arthropoda" rank="species" species="carbonaria">
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<emphasis bold="true" box="[185,426,1116,1143]" italics="true" pageId="45" pageNumber="56">Gnypeta carbonaria</emphasis>
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(Mannerheim)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,503,1152,1179]" pageId="45" pageNumber="56" type="description">
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<paragraph blockId="45.[140,608,1116,1179]" box="[140,503,1152,1179]" pageId="45" pageNumber="56">
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(
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<figureCitation box="[147,232,1152,1179]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="45" pageNumber="56">Figs 15</figureCitation>
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,
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<figureCitation box="[245,326,1152,1179]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="45" pageNumber="56">33 a, b</figureCitation>
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,
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<figureCitation box="[337,438,1152,1178]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="45" pageNumber="56">139-147</figureCitation>
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,
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<figureCitation box="[450,495,1152,1178]" captionStart="Fig" captionStartId="61.[140,174,979,1001]" captionTargetBox="[140,1106,515,855]" captionTargetPageId="61" captionText="Fig. 199. Collection localities in Canada and Alaska of G. carbonaria and G. saccharina." figureDoi="http://doi.org/10.5281/zenodo.3774152" httpUri="https://zenodo.org/record/3774152/files/figure.png" pageId="45" pageNumber="56">199</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="56" type="reference_group">
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<paragraph blockId="45.[140,1108,1222,1390]" pageId="45" pageNumber="56">
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<treatmentCitationGroup pageId="45" pageNumber="56">
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<treatmentCitation author="Mannerheim CG" box="[140,662,1222,1249]" page="75" pageId="45" pageNumber="56" year="1830">
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<taxonomicName authority="Mannerheim 1830: 75" authorityName="Mannerheim" authorityPageNumber="75" authorityYear="1830" box="[140,662,1222,1249]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="56" phylum="Arthropoda" rank="species" species="carbonaria">
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<emphasis box="[140,389,1222,1248]" italics="true" pageId="45" pageNumber="56">Bolitochara carbonaria</emphasis>
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<bibRefCitation author="Mannerheim" box="[398,662,1222,1249]" firstAuthor="Mannerheim" page="75" pageId="45" pageNumber="56" pagination="415 - 501" refId="ref23744" refString="Mannerheim CG (1830) Precis d'un nouvel arrangement de la famille des brachelytres de l'ordre des insectes coleopteres. Memoires de l'Academie Imperiale des Sciences de St. - Petersburg 1: 415 - 501. Sep. St. Petersburg, 87 pp." type="book chapter" year="1830">Mannerheim 1830: 75</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation author="Palm T" box="[675,862,1222,1249]" page="141" pageId="45" pageNumber="56" year="1966">
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<bibRefCitation author="Palm" box="[675,862,1222,1249]" firstAuthor="Palm" page="141" pageId="45" pageNumber="56" pagination="136 - 141" refId="ref23890" refString="Palm T (1966) De svenska Gnypeta - arterna (Col. Staphylinidae, Aleocharinae). Entomologisk Tidskrift 87: 136 - 141." type="journal article" year="1966">Palm 1966: 141</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Smetana A" box="[876,1104,1223,1249]" page="489" pageId="45" pageNumber="56" year="2004">
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<bibRefCitation author="Smetana" box="[876,1104,1223,1249]" firstAuthor="Smetana" page="489" pageId="45" pageNumber="56" pagination="353 - 494" refId="ref24122" refString="Smetana A (2004) Subfamily Aleocharinae Fleming, 1821 In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 2. Apollo Books, Stenstrup. 353 - 494." type="book chapter" year="2004">Smetana 2004: 489</bibRefCitation>
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</treatmentCitation>
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.
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<emphasis bold="true" box="[185,412,1257,1284]" pageId="45" pageNumber="56">TYPE LOCALITY:</emphasis>
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<collectingCountry box="[418,551,1258,1284]" name="Finland" pageId="45" pageNumber="56">FINLAND</collectingCountry>
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: in original description: “Ad Willnäs in Finlandia australi semel capta”.
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<typeStatus box="[428,535,1293,1319]" pageId="45" pageNumber="56">Holotype</typeStatus>
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not examined.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="45" pageNumber="56" type="materials_examined">
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<paragraph blockId="45.[140,1108,1222,1390]" box="[140,363,1328,1354]" pageId="45" pageNumber="56">
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<emphasis bold="true" box="[140,363,1328,1354]" pageId="45" pageNumber="56">
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<emphasis box="[140,241,1328,1354]" inLineHeading="true" pageId="45" pageNumber="56" reason="1">Material</emphasis>
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examined
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</emphasis>
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</paragraph>
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<paragraph blockId="45.[140,1108,1222,1390]" box="[185,592,1363,1390]" pageId="45" pageNumber="56">Specimens are listed in Appendix A.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="47" lastPageNumber="58" pageId="45" pageNumber="56" type="diagnosis">
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<paragraph blockId="45.[140,1108,1433,1671]" box="[140,259,1433,1459]" pageId="45" pageNumber="56">
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<emphasis bold="true" box="[140,259,1433,1459]" inLineHeading="true" pageId="45" pageNumber="56" reason="1">Diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="45.[140,1108,1433,1671]" pageId="45" pageNumber="56">
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This species can be recognized by the following combination of characters: body length 2.7-3.0 mm (
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<figureCitation box="[296,380,1504,1531]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="45" pageNumber="56">Fig. 15</figureCitation>
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); elytra at suture slightly longer than pronotum and about 1/4 wider than maximum width of pronotum (
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<figureCitation box="[637,722,1539,1566]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="45" pageNumber="56">Fig. 15</figureCitation>
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); abdomen at base distinctly narrower than elytra and subparallel (
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<figureCitation box="[537,622,1574,1601]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="45" pageNumber="56">Fig. 15</figureCitation>
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); antennal article 4 strongly elongate, 5-7 moderately elongate, 8-10 subquadrate to slightly transverse (
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<figureCitation box="[854,998,1610,1637]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="45" pageNumber="56">Figs 33 a, b</figureCitation>
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); median lobe of aedeagus with apical part triangularly produced with sinuate edges in lateral
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3774120" ID-Zenodo-Dep="3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="46" pageNumber="57" startId="46.[140,184,1590,1612]" targetBox="[152,1103,246,1521]" targetPageId="46">
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<paragraph blockId="46.[140,1108,1590,1676]" pageId="46" pageNumber="57">
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<emphasis bold="true" box="[140,286,1590,1612]" pageId="46" pageNumber="57">Figs 139-147.</emphasis>
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Genital structures of
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<taxonomicName authorityName="Palm" authorityYear="1966" box="[492,637,1590,1612]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="57" phylum="Arthropoda" rank="species" species="brincki">
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<emphasis box="[492,637,1590,1612]" italics="true" pageId="46" pageNumber="57">Gnypeta brincki</emphasis>
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</taxonomicName>
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: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8.
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</paragraph>
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</caption>
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<paragraph blockId="47.[140,1108,166,404]" pageId="47" pageNumber="58">
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view (
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<figureCitation box="[206,304,166,193]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 139</figureCitation>
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); spermatheca club-shaped, apical part of capsule approximately spherical connected to narrow and short tubular part (
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<figureCitation box="[683,781,201,228]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 145</figureCitation>
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); stem narrow, tubular, short and slightly swollen posteriorly (
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<figureCitation box="[515,615,237,264]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 145</figureCitation>
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); male tergite 8 truncate apically and with slight apical emargination (
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<figureCitation box="[447,545,272,298]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 143</figureCitation>
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); female sternite 8 with small apical emargination (
|
||
<figureCitation box="[148,249,307,333]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 147</figureCitation>
|
||
). The antennal proportions; distinct shape of spermatheca; medially emarginated female sternite 8, and the shape of the apical portion of the median lobe of the aedeagus are the best characters for recognition of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="47" pageNumber="58" type="description">
|
||
<paragraph blockId="47.[140,1108,448,1108]" box="[140,282,448,474]" pageId="47" pageNumber="58">
|
||
<emphasis bold="true" box="[140,282,448,474]" inLineHeading="true" pageId="47" pageNumber="58" reason="1">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="47.[140,1108,448,1108]" pageId="47" pageNumber="58">
|
||
Body length 2.7-3.0 mm; dark brown to black, elytra often rust brown medially, and legs and/or tarsi rust brown (
|
||
<figureCitation box="[470,553,518,545]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="47" pageNumber="58">Fig. 15</figureCitation>
|
||
); integument moderately glossy; pubescence yellowish grey and moderately long and dense; antennal article 4 strongly elongate, 5-7 moderately elongate, 8-10 subquadrate to slightly transverse (
|
||
<figureCitation box="[841,980,589,616]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="47" pageNumber="58">Figs 33 a, b</figureCitation>
|
||
); head and pronotum of about the same width (
|
||
<figureCitation box="[553,636,624,651]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="47" pageNumber="58">Fig. 15</figureCitation>
|
||
); elytra and abdomen (less so) wider than either head or pronotum; head rounded posteriorly; pronotum broadest in apical third, pubescence directed anterad along midline and laterad elsewhere; elytra at suture slightly longer than pronotum and about 1/4 wider than maximum width of pronotum, pubescence directed obliquely postero-laterad, in wavy pattern medially on each side of disc (
|
||
<figureCitation box="[227,309,800,827]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="47" pageNumber="58">Fig. 15</figureCitation>
|
||
); abdomen subparallel, distinctly narrower than elytra at base (
|
||
<figureCitation box="[1011,1092,800,827]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="47" pageNumber="58">Fig. 15</figureCitation>
|
||
); metatarsus with basal article slightly longer than the following article.
|
||
<emphasis bold="true" box="[933,993,835,861]" pageId="47" pageNumber="58">Male</emphasis>
|
||
. Tergite 8 transverse and truncate apically with small median emargination (
|
||
<figureCitation box="[897,997,870,897]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 143</figureCitation>
|
||
). Sternite 8 elongate and broadly rounded apically (
|
||
<figureCitation box="[630,732,906,932]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 144</figureCitation>
|
||
). Median lobe of aedeagus with triangularly produced apical part of tubus in lateral view (
|
||
<figureCitation box="[782,879,941,967]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 139</figureCitation>
|
||
); bulbus moderately large in dorsal view (
|
||
<figureCitation box="[372,471,976,1002]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 140</figureCitation>
|
||
); internal sac with complex structures as illustrated (
|
||
<figureCitation captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Figs 139-141</figureCitation>
|
||
).
|
||
<emphasis bold="true" box="[263,347,1011,1037]" pageId="47" pageNumber="58">Female</emphasis>
|
||
. Tergite 8 truncate apically (
|
||
<figureCitation box="[676,777,1011,1038]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 146</figureCitation>
|
||
). Sternite 8 broadly rounded posteriorly with small median emargination (
|
||
<figureCitation box="[648,747,1046,1073]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 147</figureCitation>
|
||
). Spermatheca with tubular and short, club-shaped capsule (
|
||
<figureCitation box="[452,550,1081,1108]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 145</figureCitation>
|
||
); stem short and broadly tubular (
|
||
<figureCitation box="[936,1034,1081,1108]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="47" pageNumber="58">Fig. 145</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="47" pageNumber="58" type="distribution">
|
||
<paragraph blockId="47.[140,1108,1152,1390]" box="[140,412,1152,1178]" pageId="47" pageNumber="58">
|
||
<emphasis bold="true" box="[140,291,1152,1178]" inLineHeading="true" pageId="47" pageNumber="58" reason="1">Distribution</emphasis>
|
||
(Fig. 199)
|
||
</paragraph>
|
||
<paragraph blockId="47.[140,1108,1152,1390]" pageId="47" pageNumber="58">
|
||
<taxonomicName baseAuthorityName="Mannerheim" baseAuthorityYear="1830" box="[140,354,1187,1214]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="58" phylum="Arthropoda" rank="species" species="carbonaria">
|
||
<emphasis box="[140,354,1187,1214]" italics="true" pageId="47" pageNumber="58">Gnypeta carbonaria</emphasis>
|
||
</taxonomicName>
|
||
is a Holarctic species and is here reported from
|
||
<collectingCountry box="[892,978,1187,1213]" name="Canada" pageId="47" pageNumber="58">Canada</collectingCountry>
|
||
and Alaska for the first time. It was previously known from the Palaearctic region and was recorded from Europe, Northern Africa, and Asia including eastern and western Siberia (
|
||
<bibRefCitation author="Smetana" box="[147,317,1293,1319]" firstAuthor="Smetana" pageId="47" pageNumber="58" pagination="353 - 494" refId="ref24122" refString="Smetana A (2004) Subfamily Aleocharinae Fleming, 1821 In: Lobl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera, Vol. 2. Apollo Books, Stenstrup. 353 - 494." type="book chapter" year="2004">Smetana 2004</bibRefCitation>
|
||
). We discovered this species to be transcontinental in North America with records from Alaska,
|
||
<collectingRegion box="[451,699,1328,1354]" country="Canada" name="Northwest Territories" pageId="47" pageNumber="58">Northwest Territories</collectingRegion>
|
||
,
|
||
<collectingRegion box="[714,800,1328,1354]" country="Canada" name="Alberta" pageId="47" pageNumber="58">Alberta</collectingRegion>
|
||
,
|
||
<collectingRegion box="[815,974,1328,1354]" country="Canada" name="Saskatchewan" pageId="47" pageNumber="58">Saskatchewan</collectingRegion>
|
||
,
|
||
<collectingRegion box="[989,1103,1328,1354]" country="Canada" name="Manitoba" pageId="47" pageNumber="58">Manitoba</collectingRegion>
|
||
,
|
||
<collectingRegion box="[140,234,1363,1389]" country="Canada" name="Ontario" pageId="47" pageNumber="58">Ontario</collectingRegion>
|
||
,
|
||
<collectingRegion box="[245,336,1363,1389]" country="Canada" name="Quebec" pageId="47" pageNumber="58">Quebec</collectingRegion>
|
||
, and
|
||
<collectingRegion box="[397,582,1363,1390]" country="Canada" name="New Brunswick" pageId="47" pageNumber="58">New Brunswick</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="47.[140,1108,1433,1531]" box="[140,467,1433,1460]" pageId="47" pageNumber="58">
|
||
<emphasis bold="true" box="[140,467,1433,1460]" pageId="47" pageNumber="58">
|
||
<emphasis box="[140,265,1433,1459]" inLineHeading="true" pageId="47" pageNumber="58" reason="1">Collection</emphasis>
|
||
and habitat data
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="47.[140,1108,1433,1531]" pageId="47" pageNumber="58">
|
||
In
|
||
<collectingCountry box="[172,258,1469,1495]" name="Canada" pageId="47" pageNumber="58">Canada</collectingCountry>
|
||
and Alaska, adults were collected in July and August from wet ground near a bog, in rotten fungus, and in a UV light trap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="48" lastPageNumber="59" pageId="47" pageNumber="58" type="discussion">
|
||
<paragraph blockId="47.[140,1108,1574,1671]" box="[140,271,1574,1600]" pageId="47" pageNumber="58">
|
||
<emphasis bold="true" box="[140,271,1574,1600]" inLineHeading="true" pageId="47" pageNumber="58" reason="1">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="47.[140,1108,1574,1671]" lastBlockId="48.[140,1108,166,475]" lastPageId="48" lastPageNumber="59" pageId="47" pageNumber="58">
|
||
We compared North American specimens of
|
||
<taxonomicName baseAuthorityName="Mannerheim" baseAuthorityYear="1830" box="[681,840,1610,1636]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="58" phylum="Arthropoda" rank="species" species="carbonaria">
|
||
<emphasis box="[681,840,1610,1636]" italics="true" pageId="47" pageNumber="58">G. carbonaria</emphasis>
|
||
</taxonomicName>
|
||
with those from
|
||
<collectingCountry name="Sweden" pageId="47" pageNumber="58">Sweden</collectingCountry>
|
||
and found no distinct external or genital morphological differences between the two populations. We also compared the newly designated female
|
||
<typeStatus box="[969,1074,166,192]" pageId="48" pageNumber="59">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Casey" authorityName="Casey" authorityYear="1911" box="[140,449,202,228]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="59" phylum="Arthropoda" rank="species" species="brunnescens">
|
||
<emphasis box="[140,372,202,228]" italics="true" pageId="48" pageNumber="59">Gnypeta brunnescens</emphasis>
|
||
Casey
|
||
</taxonomicName>
|
||
with Canadian and Swedish specimens of
|
||
<taxonomicName baseAuthorityName="Mannerheim" baseAuthorityYear="1830" box="[950,1108,202,228]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="59" phylum="Arthropoda" rank="species" species="carbonaria">
|
||
<emphasis box="[950,1108,202,228]" italics="true" pageId="48" pageNumber="59">G. carbonaria</emphasis>
|
||
</taxonomicName>
|
||
and found no differences between female genital features. However, Casey’s type of
|
||
<taxonomicName authorityName="Casey" authorityYear="1911" box="[175,346,272,298]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="59" phylum="Arthropoda" rank="species" species="brunnescens">
|
||
<emphasis box="[175,346,272,298]" italics="true" pageId="48" pageNumber="59">G. brunnescens</emphasis>
|
||
</taxonomicName>
|
||
differs from both populations by the light brown body colour instead of dark brown or black and by a slightly different pubescence pattern on the pronotum, which may be caused by the preparation of the specimen. Due to these differences, we hesitate to formally synonymize
|
||
<taxonomicName authorityName="Casey" authorityYear="1911" box="[702,871,378,404]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="59" phylum="Arthropoda" rank="species" species="brunnescens">
|
||
<emphasis box="[702,871,378,404]" italics="true" pageId="48" pageNumber="59">G. brunnescens</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName baseAuthorityName="Mannerheim" baseAuthorityYear="1830" box="[942,1100,378,404]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="59" phylum="Arthropoda" rank="species" species="carbonaria">
|
||
<emphasis box="[942,1100,378,404]" italics="true" pageId="48" pageNumber="59">G. carbonaria</emphasis>
|
||
</taxonomicName>
|
||
. Additional specimens of
|
||
<taxonomicName authorityName="Casey" authorityYear="1911" box="[426,592,413,439]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="59" phylum="Arthropoda" rank="species" species="brunnescens">
|
||
<emphasis box="[426,592,413,439]" italics="true" pageId="48" pageNumber="59">G. brunnescens</emphasis>
|
||
</taxonomicName>
|
||
are needed from the type locality to confirm its taxonomic status.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |