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<mods:title id="EC12554CE5DA41607C519D8087AEDAF5">Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae)</mods:title>
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<mods:namePart id="0CD5AF76ECAF110D6E61195A134E2736">Almeida-Silva, Lina M.</mods:namePart>
<mods:affiliation id="F18FC6849C23EFE3730ED11D6800C6F1">Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.</mods:affiliation>
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<mods:namePart id="F2C26B4E6AE4CC00C346AC4F578ECDDD">Brescovit, Antonio D.</mods:namePart>
<mods:affiliation id="20BC62F142BA274093B2116846D9D576">Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. &amp; Laboratório de Coleções Zoológicas-LCZ, Instituto Butantan, Avenida Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil &amp; Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.</mods:affiliation>
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<mods:title id="DC6D9E7A283B5172D5AA33E57E9513A5">Zootaxa</mods:title>
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<mods:date id="2423CDD3B3F5C99F126A2B94D9690DBA">2024</mods:date>
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<treatment id="B86D8799FF9EFFF88CDDFD26FD85720D" ID-DOI="http://doi.org/10.5281/zenodo.10850417" ID-Zenodo-Dep="10850417" LSID="urn:lsid:plazi:treatment:B86D8799FF9EFFF88CDDFD26FD85720D" httpUri="http://treatment.plazi.org/id/B86D8799FF9EFFF88CDDFD26FD85720D" lastPageId="30" lastPageNumber="181" pageId="29" pageNumber="180">
<subSubSection id="78DE6504FF9EFFFB8CDDFD26FD8B7324" box="[151,584,762,789]" pageId="29" pageNumber="180" type="nomenclature">
<paragraph id="307B368FFF9EFFFB8CDDFD26FD8B7324" blockId="29.[151,584,762,825]" box="[151,584,762,789]" pageId="29" pageNumber="180">
<heading id="6B3381E3FF9EFFFB8CDDFD26FD8B7324" bold="true" box="[151,584,762,789]" fontSize="11" level="1" pageId="29" pageNumber="180" reason="1">
<taxonomicName id="F7C44D0CFF9EFFFB8CDDFD26FD8B7324" ID-CoL="3GQMJ" authority="(Keyserling, 1878)" baseAuthorityName="Keyserling" baseAuthorityYear="1878" box="[151,584,762,789]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="02B0EA9DFF9EFFFB8CDDFD26FD8B7324" bold="true" box="[151,584,762,789]" pageId="29" pageNumber="180">
<emphasis id="02B0EA9DFF9EFFFB8CDDFD26FE9E7325" bold="true" box="[151,349,762,789]" italics="true" pageId="29" pageNumber="180">Goeldia obscura</emphasis>
(
<bibRefCitation id="54554B7EFF9EFFFB8D26FD26FD837324" author="Keyserling, E." box="[364,576,762,788]" pageId="29" pageNumber="180" pagination="571 - 624" refId="ref35210" refString="Keyserling, E. (1878) Spinnen aus Uruguay und einigen anderen Gegenden Amerikas. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 27, 571 - 624." type="journal article" year="1878">Keyserling, 1878</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="78DE6504FF9EFFFB8CDDFCC3FEE47309" box="[151,295,798,825]" pageId="29" pageNumber="180" type="description">
<paragraph id="307B368FFF9EFFFB8CDDFCC3FEE47309" blockId="29.[151,584,762,825]" box="[151,295,798,825]" pageId="29" pageNumber="180">
(
<figureCitation id="A8FF2A0AFF9EFFFB8CD5FCC3FF357309" box="[159,246,799,825]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Figs 18</figureCitation>
,
<figureCitation id="A8FF2A0AFF9EFFFB8D48FCC2FEDC7309" box="[258,287,798,825]" captionStart="FIGURE 25" captionStartId="39.[151,250,1041,1065]" captionTargetBox="[184,1404,188,1011]" captionTargetId="figure-21@39.[177,1411,181,1018]" captionTargetPageId="39" captionText="FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities." figureDoi="http://doi.org/10.5281/zenodo.10847445" httpUri="https://zenodo.org/record/10847445/files/figure.png" pageId="29" pageNumber="180">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF9EFFFB8CDDFCBFFD9B73E7" pageId="29" pageNumber="180" type="reference_group">
<paragraph id="307B368FFF9EFFFB8CDDFCBFFDB073AA" blockId="29.[151,1436,867,983]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="10D411A1FF9EFFFB8CDDFCBFFDB073AA" pageId="29" pageNumber="180">
<treatmentCitation id="B165109EFF9EFFFB8CDDFCBFFD8F734B" author="Keyserling, E." box="[151,588,867,891]" page="591" pageId="29" pageNumber="180" year="1878">
<taxonomicName id="F7C44D0CFF9EFFFB8CDDFCBFFD8F734B" authority="Keyserling, 1878: 591" authorityName="Keyserling" authorityPageNumber="591" authorityYear="1878" box="[151,588,867,891]" class="Arachnida" family="Titanoecidae" genus="Titanoeca" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="02B0EA9DFF9EFFFB8CDDFCBFFE98734B" bold="true" box="[151,347,867,891]" italics="true" pageId="29" pageNumber="180">Titanoeca obscura</emphasis>
<materialsCitation id="80AC3CD2FF9EFFFB8D2EFCBFFD8F734B" box="[356,588,867,891]" country="Colombia" latitude="4.598889" location="Santa Fe" longLatPrecision="21" longitude="-74.08083" pageId="29" pageNumber="180" specimenCount="1" specimenCount-female="1" stateProvince="Bogota" typeStatus="holotype">
<bibRefCitation id="54554B7EFF9EFFFB8D2EFCBFFD8F734B" author="Keyserling, E." box="[356,588,867,891]" pageId="29" pageNumber="180" pagination="571 - 624" refId="ref35210" refString="Keyserling, E. (1878) Spinnen aus Uruguay und einigen anderen Gegenden Amerikas. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 27, 571 - 624." type="journal article" year="1878">Keyserling, 1878: 591</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
pl. 14, fig. 15 (
<typeStatus id="EF7F882DFF9EFFFB8EBDFCBFFC92734B" box="[759,849,867,891]" pageId="29" pageNumber="180" type="holotype">holotype</typeStatus>
<specimenCount id="26C2FD06FF9EFFFB8F11FCB8FCAD734B" box="[859,878,868,891]" count="1" pageId="29" pageNumber="180" type="female">♀</specimenCount>
from
<location id="351B6054FF9EFFFB8FF8FCBFFBCE734B" LSID="urn:lsid:plazi:treatment:B86D8799FF9EFFF88CDDFD26FD85720D:351B6054FF9EFFFB8FF8FCBFFBCE734B" box="[946,1037,867,891]" country="Colombia" latitude="4.598889" longLatPrecision="21" longitude="-74.08083" name="Santa Fe" pageId="29" pageNumber="180" stateProvince="Bogota">Santa Fé</location>
de
<collectingRegion id="F200F86DFF9EFFFB8872FCBFFB42734B" box="[1080,1153,867,891]" country="Colombia" name="Bogota" pageId="29" pageNumber="180">Bogotá</collectingRegion>
(
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<geoCoordinate id="55F05048FF9EFFFB8946FCBFFA52734B" box="[1292,1425,867,891]" degrees="74" direction="west" minutes="04" orientation="longitude" pageId="29" pageNumber="180" precision="15" seconds="51" value="-74.08083">74°04’51”W</geoCoordinate>
),
<collectingCountry id="48D3761FFF9EFFFB8C8DFC5EFEED73AA" box="[199,302,898,922]" name="Colombia" pageId="29" pageNumber="180">Colombia</collectingCountry>
, deposited in NHM, examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="307B368FFF9EFFFB8CDDFC7CFD497388" blockId="29.[151,1436,867,983]" box="[151,650,928,953]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="10D411A1FF9EFFFB8CDDFC7CFD497388" box="[151,650,928,953]" pageId="29" pageNumber="180">
<taxonomicName id="F7C44D0CFF9EFFFB8CDDFC7CFEBE7388" authority="Petrunkevitch 1911" box="[151,381,928,952]" class="Arachnida" family="Amaurobiidae" genus="Amaurobius" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="02B0EA9DFF9EFFFB8CDDFC7CFEBE7388" bold="true" box="[151,381,928,952]" italics="true" pageId="29" pageNumber="180">Amaurobius obscurus</emphasis>
</taxonomicName>
:
<treatmentCitation id="B165109EFF9EFFFB8DC1FC7CFD457388" author="Petrunkevitch, A." box="[395,646,928,953]" page="105" pageId="29" pageNumber="180" year="1911">
<bibRefCitation id="54554B7EFF9EFFFB8DC1FC7CFD457388" author="Petrunkevitch, A." box="[395,646,928,953]" pageId="29" pageNumber="180" pagination="1 - 791" refId="ref35763" refString="Petrunkevitch, A. (1911) A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History, 29, 1 - 791. https: // doi. org / 10.5962 / bhl. title. 23819" type="journal article" year="1911">Petrunkevitch 1911: 105</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="307B368FFF9EFFFB8CDDFC63FD9B73E7" blockId="29.[151,1436,867,983]" box="[151,600,959,983]" pageId="29" pageNumber="180">
<treatmentCitationGroup id="10D411A1FF9EFFFB8CDDFC63FD9B73E7" box="[151,600,959,983]" pageId="29" pageNumber="180">
<taxonomicName id="F7C44D0CFF9EFFFB8CDDFC63FEFF73E7" ID-CoL="3GQMC" baseAuthorityName="Keyserling" baseAuthorityYear="1878" box="[151,316,959,983]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="funesta">
<emphasis id="02B0EA9DFF9EFFFB8CDDFC63FEFF73E7" bold="true" box="[151,316,959,983]" italics="true" pageId="29" pageNumber="180">Goeldia funesta</emphasis>
</taxonomicName>
:
<treatmentCitation id="B165109EFF9EFFFB8D00FC63FDD373E7" author="Lehtinen, P. T." box="[330,528,959,983]" page="236" pageId="29" pageNumber="180" year="1967">
<bibRefCitation id="54554B7EFF9EFFFB8D00FC63FDD373E7" author="Lehtinen, P. T." box="[330,528,959,983]" pageId="29" pageNumber="180" pagination="199 - 468" refId="ref35363" refString="Lehtinen, P. T. (1967) Classification of the Cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4 (3), 199 - 468." type="journal article" year="1967">Lehtinen 1967: 236</bibRefCitation>
</treatmentCitation>
(Part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF9EFFFB8CDDFBD9FCC675B7" pageId="29" pageNumber="180" type="discussion">
<paragraph id="307B368FFF9EFFFB8CDDFBD9FBD37457" blockId="29.[151,1437,1029,1992]" pageId="29" pageNumber="180">
<emphasis id="02B0EA9DFF9EFFFB8CDDFBD9FEC87410" bold="true" box="[151,267,1029,1056]" pageId="29" pageNumber="180">Remarks.</emphasis>
The description of a male presented by
<bibRefCitation id="54554B7EFF9EFFFB8EF7FBD9FC92742F" author="Leech, R. E." box="[701,849,1029,1055]" pageId="29" pageNumber="180" pagination="1 - 182" refId="ref35316" refString="Leech, R. E. (1972) A revision of the Nearctic Amaurobiidae (Arachnida: Araneida). Memoirs of the Entomological Society of Canada, 84, 1 - 182. https: // doi. org / 10.4039 / entm 10484 fv" type="journal article" year="1972">Leech (1972)</bibRefCitation>
for
<taxonomicName id="F7C44D0CFF9EFFFB8F36FBD9FBF5742F" baseAuthorityName="Keyserling" baseAuthorityYear="1878" box="[892,1078,1029,1055]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="02B0EA9DFF9EFFFB8F36FBD9FBF5742F" box="[892,1078,1029,1055]" italics="true" pageId="29" pageNumber="180">Goeldia obscura</emphasis>
</taxonomicName>
refers to the specimen described by
<bibRefCitation id="54554B7EFF9EFFFB8CF6FBF5FE487473" author="Keyserling, E." box="[188,395,1065,1091]" pageId="29" pageNumber="180" refId="ref35239" refString="Keyserling, E. (1891) Die Spinnen Amerikas. Brasilianische. Spinnen. Vol. 3. Verlag von Bauer &amp; Raspe (E. Kuster), Nurnberg, 278 pp." type="book" year="1891">Keyserling (1891)</bibRefCitation>
and it was erroneously interpreted as the male of
<taxonomicName id="F7C44D0CFF9EFFFB8F85FBF6FAEC7473" authority="(Keyserling, 1878)" baseAuthorityName="Keyserling" baseAuthorityYear="1878" box="[975,1327,1065,1092]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="02B0EA9DFF9EFFFB8F85FBF6FB8E7473" box="[975,1101,1065,1092]" italics="true" pageId="29" pageNumber="180">G. obscura</emphasis>
(
<bibRefCitation id="54554B7EFF9EFFFB8814FBF5FAE47473" author="Keyserling, E." box="[1118,1319,1065,1091]" pageId="29" pageNumber="180" pagination="571 - 624" refId="ref35210" refString="Keyserling, E. (1878) Spinnen aus Uruguay und einigen anderen Gegenden Amerikas. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 27, 571 - 624." type="journal article" year="1878">Keyserling, 1878</bibRefCitation>
)
</taxonomicName>
. Here we present the first description of the male of
<taxonomicName id="F7C44D0CFF9EFFFB8E39FB91FBCF7457" authority="(Keyserling, 1878)" baseAuthorityName="Keyserling" baseAuthorityYear="1878" box="[627,1036,1101,1127]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="02B0EA9DFF9EFFFB8E39FB91FCED7457" box="[627,814,1101,1127]" italics="true" pageId="29" pageNumber="180">Goeldia obscura</emphasis>
(
<bibRefCitation id="54554B7EFF9EFFFB8F77FB91FBC77457" author="Keyserling, E." box="[829,1028,1101,1127]" pageId="29" pageNumber="180" pagination="571 - 624" refId="ref35210" refString="Keyserling, E. (1878) Spinnen aus Uruguay und einigen anderen Gegenden Amerikas. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 27, 571 - 624." type="journal article" year="1878">Keyserling, 1878</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="307B368FFF9EFFFB8C8DFBADFCC675B7" blockId="29.[151,1437,1029,1992]" pageId="29" pageNumber="180">
<emphasis id="02B0EA9DFF9EFFFB8C8DFBADFDF174BC" bold="true" box="[199,562,1137,1164]" pageId="29" pageNumber="180">Additional material examined.</emphasis>
<materialsCitation id="80AC3CD2FF9EFFFB8E72FBADFDFF7480" collectionCode="CAS" collectorName="V. &amp; B. Roth &amp;" country="Colombia" latitude="1.9166667" location="San Agustin" longLatPrecision="1307" longitude="-76.23333" pageId="29" pageNumber="180" specimenCode="CAS 9025938, CAS 9025935" specimenCount="2" specimenCount-male="2" stateProvince="Huila">
<collectingCountry id="48D3761FFF9EFFFB8E72FBADFD0E74BB" box="[568,717,1137,1163]" name="Colombia" pageId="29" pageNumber="180">COLOMBIA</collectingCountry>
.
<emphasis id="02B0EA9DFF9EFFFB8E90FBADFCE074BB" box="[730,803,1137,1163]" italics="true" pageId="29" pageNumber="180">
<collectingRegion id="F200F86DFF9EFFFB8E90FBADFCDE74BB" box="[730,797,1137,1163]" country="Colombia" name="Huila" pageId="29" pageNumber="180">Huila</collectingRegion>
:
</emphasis>
<location id="351B6054FF9EFFFB8F60FBADFC7274BB" LSID="urn:lsid:plazi:treatment:B86D8799FF9EFFF88CDDFD26FD85720D:351B6054FF9EFFFB8F60FBADFC7274BB" box="[810,945,1137,1163]" country="Colombia" latitude="1.9166667" longLatPrecision="1307" longitude="-76.23333" name="San Agustin" pageId="29" pageNumber="180" stateProvince="Huila">San Agustín</location>
(
<geoCoordinate id="55F05048FF9EFFFB8F8AFBADFBFF74BC" box="[960,1084,1137,1164]" degrees="1" direction="north" minutes="52" orientation="latitude" pageId="29" pageNumber="180" precision="15" seconds="45" value="1.8791667">1°52’45”N</geoCoordinate>
<geoCoordinate id="55F05048FF9EFFFB880BFBADFB1774BB" box="[1089,1236,1137,1163]" degrees="76" direction="west" minutes="16" orientation="longitude" pageId="29" pageNumber="180" precision="15" seconds="06" value="-76.26833">76°16’06”W</geoCoordinate>
),
<specimenCount id="26C2FD06FF9EFFFB88ADFBADFACF74BB" box="[1255,1292,1137,1163]" count="1" pageId="29" pageNumber="180" type="male">1♂</specimenCount>
, 24.III.1994,
<collectorName id="9D315359FF9EFFFB8CDDFB4AFEF5749F" box="[151,310,1173,1200]" pageId="29" pageNumber="180">V. &amp; B. Roth</collectorName>
coll. (
<specimenCode id="60629EF4FF9EFFFB8DCCFB49FDF27480" box="[390,561,1173,1200]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="180" type="Museum">CAS 9025938</specimenCode>
);
</materialsCitation>
<materialsCitation id="80AC3CD2FF9EFFFB8E02FB49FDB774E4" collectionCode="CAS" collectorName="V. &amp; B. Roth" country="Colombia" latitude="1.9166667" location="San Agustin" longLatPrecision="1307" longitude="-76.23333" pageId="29" pageNumber="180" specimenCode="CAS 9025938, CAS 9025935" specimenCount="2" specimenCount-male="2" stateProvince="Huila">
San Agustín, Archaeological site of San Agustín (
<geoCoordinate id="55F05048FF9EFFFB88DBFB49FB277480" box="[1169,1252,1173,1200]" degrees="1" direction="north" minutes="55" orientation="latitude" pageId="29" pageNumber="180" precision="925" value="1.9166667">1°55’N</geoCoordinate>
<geoCoordinate id="55F05048FF9EFFFB88BAFB49FA99749F" box="[1264,1370,1173,1199]" degrees="76" direction="west" minutes="14" orientation="longitude" pageId="29" pageNumber="180" precision="925" value="-76.23333">76°14’W</geoCoordinate>
),
<specimenCount id="26C2FD06FF9EFFFB8938FB49FA54749F" box="[1394,1431,1173,1199]" count="1" pageId="29" pageNumber="180" type="male">1♂</specimenCount>
, 9.IX.1980,
<collectorName id="9D315359FF9EFFFB8D50FB65FEB574E3" box="[282,374,1209,1235]" pageId="29" pageNumber="180">C. Gold</collectorName>
coll. (
<specimenCode id="60629EF4FF9EFFFB8D89FB65FDAF74E4" box="[451,620,1209,1236]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="180" type="Museum">CAS 9025935</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="80AC3CD2FF9EFFFB8EC8FB65FD5C74C7" country="Colombia" county="Mosquera" elevation="1600" latitude="4.7308335" location="Cachipay" longLatPrecision="21" longitude="-74.4375" municipality="Laguna de la Herrera" pageId="29" pageNumber="180" specimenCount="2" specimenCount-female="2" stateProvince="Cundinamarca">
<emphasis id="02B0EA9DFF9EFFFB8EC8FB65FCF074E3" box="[642,819,1209,1235]" italics="true" pageId="29" pageNumber="180">
<collectingRegion id="F200F86DFF9EFFFB8EC8FB65FCEF74E3" box="[642,812,1209,1235]" country="Colombia" name="Cundinamarca" pageId="29" pageNumber="180">Cundinamarca</collectingRegion>
:
</emphasis>
<collectingCounty id="D91A4E03FF9EFFFB8F77FB66FC7374E4" box="[829,944,1210,1236]" pageId="29" pageNumber="180">Mosquera</collectingCounty>
,
<collectorName id="9D315359FF9EFFFB8FF4FB66FB7774E4" box="[958,1204,1209,1236]" pageId="29" pageNumber="180">
<collectingMunicipality id="D01FACF5FF9EFFFB8FF4FB66FB7774E4" box="[958,1204,1209,1236]" pageId="29" pageNumber="180">Laguna de la Herrera</collectingMunicipality>
</collectorName>
(
<geoCoordinate id="55F05048FF9EFFFB888DFB65FAD874E3" box="[1223,1307,1209,1235]" degrees="4" direction="north" minutes="42" orientation="latitude" pageId="29" pageNumber="180" precision="925" value="4.7">4°42’N</geoCoordinate>
<geoCoordinate id="55F05048FF9EFFFB896FFB65FA4C74E3" box="[1317,1423,1209,1235]" degrees="74" direction="west" minutes="18" orientation="longitude" pageId="29" pageNumber="180" precision="925" value="-74.3">74°18’W</geoCoordinate>
),
<specimenCount id="26C2FD06FF9EFFFB8CDDFB01FF7974C7" box="[151,186,1245,1271]" count="1" pageId="29" pageNumber="180" type="female">1♀</specimenCount>
, VII.1991,
<collectorName id="9D315359FF9EFFFB8D0AFB02FE6A74C7" box="[320,425,1245,1272]" pageId="29" pageNumber="180">E. Flórez</collectorName>
coll. (ICN-Ar 6067);
</materialsCitation>
<materialsCitation id="80AC3CD2FF9EFFFB8EE2FB01FDB0752B" country="Colombia" county="Mosquera" elevation="1600" latitude="4.7308335" location="Cachipay" longLatPrecision="21" longitude="-74.4375" municipality="Laguna de la Herrera" pageId="29" pageNumber="180" specimenCount="2" specimenCount-female="2" stateProvince="Cundinamarca">
<location id="351B6054FF9EFFFB8EE2FB01FCD174C7" LSID="urn:lsid:plazi:treatment:B86D8799FF9EFFF88CDDFD26FD85720D:351B6054FF9EFFFB8EE2FB01FCD174C7" box="[680,786,1245,1271]" country="Colombia" county="Mosquera" latitude="4.7308335" longLatPrecision="21" longitude="-74.4375" municipality="Laguna de la Herrera" name="Cachipay" pageId="29" pageNumber="180" stateProvince="Cundinamarca">Cachipay</location>
(
<geoCoordinate id="55F05048FF9EFFFB8F6EFB01FC5C74C8" box="[804,927,1245,1272]" degrees="4" direction="north" minutes="43" orientation="latitude" pageId="29" pageNumber="180" precision="15" seconds="51" value="4.7308335">4°43’51”N</geoCoordinate>
<geoCoordinate id="55F05048FF9EFFFB8FE3FB01FBFF74C8" box="[937,1084,1245,1272]" degrees="74" direction="west" minutes="26" orientation="longitude" pageId="29" pageNumber="180" precision="15" seconds="15" value="-74.4375">74°26’15”W</geoCoordinate>
),
<collectorName id="9D315359FF9EFFFB881AFB01FB0274C7" box="[1104,1217,1245,1271]" pageId="29" pageNumber="180">Alto Tolú</collectorName>
, elev. 1600 m,
<specimenCount id="26C2FD06FF9EFFFB8938FB01FA5674C7" box="[1394,1429,1245,1271]" count="1" pageId="29" pageNumber="180" type="female">1♀</specimenCount>
, 5.XII.1996,
<collectorName id="9D315359FF9EFFFB8D6BFADEFE4B752B" box="[289,392,1281,1308]" pageId="29" pageNumber="180">E. Floréz</collectorName>
coll. (ICN-Ar 1611)
</materialsCitation>
;
<materialsCitation id="80AC3CD2FF9EFFFB8ECAFADDFDE57570" country="Colombia" elevation="2600" latitude="4.7077775" location="Bogota" longLatPrecision="21" longitude="-74.23278" pageId="29" pageNumber="180" specimenCount="29" specimenCount-female="9" stateProvince="Bogota">
<collectingRegion id="F200F86DFF9EFFFB8ECAFADDFD12752B" box="[640,721,1281,1307]" country="Colombia" name="Bogota" pageId="29" pageNumber="180">Bogotá</collectingRegion>
(4°35’56”N 74°04’51”W), elev. 2600 m,
<specimenCount id="26C2FD06FF9EFFFB88E1FADDFB0C752B" box="[1195,1231,1281,1307]" count="3" pageId="29" pageNumber="180" type="female">3♀</specimenCount>
,
<specimenCount id="26C2FD06FF9EFFFB8897FADDFADF752B" box="[1245,1308,1281,1307]" count="2" pageId="29" pageNumber="180" type="generic">2 juv.</specimenCount>
20.V.1979,
<collectorName id="9D315359FF9EFFFB8CDDFAF9FEF1750F" box="[151,306,1317,1343]" pageId="29" pageNumber="180">C. Hernandez</collectorName>
coll. (ICN-Ar 5349);
</materialsCitation>
<materialsCitation id="80AC3CD2FF9EFFFB8E65FAFAFCC275B7" country="Colombia" elevation="2600" latitude="4.7077775" location="Bogota" longLatPrecision="21" longitude="-74.23278" pageId="29" pageNumber="180" specimenCount="29" specimenCount-female="9" stateProvince="Bogota">
Mosquera (
<geoCoordinate id="55F05048FF9EFFFB8EFAFAF9FCEE750F" box="[688,813,1317,1343]" degrees="4" direction="north" minutes="42" orientation="latitude" pageId="29" pageNumber="180" precision="15" seconds="28" value="4.7077775">4°42’28”N</geoCoordinate>
<geoCoordinate id="55F05048FF9EFFFB8F7FFAF9FC047570" box="[821,967,1317,1344]" degrees="74" direction="west" minutes="13" orientation="longitude" pageId="29" pageNumber="180" precision="15" seconds="58" value="-74.23278">74°13’58”W</geoCoordinate>
),
<collectorName id="9D315359FF9EFFFB8F97FAF9FBA4750F" box="[989,1127,1317,1343]" pageId="29" pageNumber="180">Mondoñedo</collectorName>
, elev. 2600 m,
<specimenCount id="26C2FD06FF9EFFFB895CFAF9FAF8750F" box="[1302,1339,1317,1343]" count="4" pageId="29" pageNumber="180" type="female">4♀</specimenCount>
,
<specimenCount id="26C2FD06FF9EFFFB8903FAF9FA5A750F" box="[1353,1433,1317,1343]" count="11" pageId="29" pageNumber="180" type="generic">11 juv.</specimenCount>
, 6.IV.2002,
<collectorName id="9D315359FF9EFFFB8D59FA95FEAC7553" box="[275,367,1353,1379]" pageId="29" pageNumber="180">C. Niño</collectorName>
,
<collectorName id="9D315359FF9EFFFB8D30FA95FE147554" box="[378,471,1353,1380]" pageId="29" pageNumber="180">G. Hora</collectorName>
,
<collectorName id="9D315359FF9EFFFB8DA8FA96FDA37553" box="[482,608,1353,1380]" pageId="29" pageNumber="180">J. Martinez</collectorName>
&amp;
<collectorName id="9D315359FF9EFFFB8EC8FA96FCDB7553" box="[642,792,1353,1380]" pageId="29" pageNumber="180">A. Castañeda</collectorName>
coll. (ICN-Ar 2082);
<specimenCount id="26C2FD06FF9EFFFB8844FA95FBF07553" box="[1038,1075,1353,1379]" count="2" pageId="29" pageNumber="180" type="female">2♀</specimenCount>
,
<specimenCount id="26C2FD06FF9EFFFB8875FA96FB427553" box="[1087,1153,1353,1380]" count="7" pageId="29" pageNumber="180" type="generic">7 juv.</specimenCount>
, 23.III.2002,
<collectorName id="9D315359FF9EFFFB895DFA96FA5B7553" box="[1303,1432,1353,1380]" pageId="29" pageNumber="180">J. Martinez</collectorName>
,
<collectorName id="9D315359FF9EFFFB8CDDFAB1FF3075B7" box="[151,243,1389,1415]" pageId="29" pageNumber="180">C. Niño</collectorName>
,
<collectorName id="9D315359FF9EFFFB8CB4FAB1FE9E75B8" box="[254,349,1389,1416]" pageId="29" pageNumber="180">G. Mura</collectorName>
&amp;
<collectorName id="9D315359FF9EFFFB8D35FAB2FDD575B7" box="[383,534,1389,1416]" pageId="29" pageNumber="180">A. Costañeda</collectorName>
coll. (ICN-Ar 2083)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF9EFFFB8C8DFA4DFD25760B" pageId="29" pageNumber="180" type="diagnosis">
<paragraph id="307B368FFF9EFFFB8C8DFA4DFD25760B" blockId="29.[151,1437,1029,1992]" pageId="29" pageNumber="180">
<emphasis id="02B0EA9DFF9EFFFB8C8DFA4DFE82759C" bold="true" box="[199,321,1425,1452]" pageId="29" pageNumber="180">Diagnosis.</emphasis>
Males of
<taxonomicName id="F7C44D0CFF9EFFFB8DF6FA4DFDB9759B" baseAuthorityName="Keyserling" baseAuthorityYear="1878" box="[444,634,1425,1451]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="180" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="02B0EA9DFF9EFFFB8DF6FA4DFDB9759B" box="[444,634,1425,1451]" italics="true" pageId="29" pageNumber="180">Goeldia obscura</emphasis>
</taxonomicName>
differ from other species by the short, thick, and distally constricted patellar apophysis (
<figureCitation id="A8FF2A0AFF9EFFFB8D25FA69FE1775FF" box="[367,468,1461,1487]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18C</figureCitation>
); and by the tegular process distally narrowed and ventrally curved (
<figureCitation id="A8FF2A0AFF9EFFFB888CFA69FAEE75FF" box="[1222,1325,1461,1487]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18A</figureCitation>
). Females are recognized by the wide median field of the epigynum not enclosing the copulatory openings (
<figureCitation id="A8FF2A0AFF9EFFFB88B6FA05FAA075C3" box="[1276,1379,1497,1523]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18E</figureCitation>
); the copulatory ducts distally bent (
<figureCitation id="A8FF2A0AFF9EFFFB8E48FA21FDA97627" box="[514,618,1533,1559]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18F</figureCitation>
); the vertical spermathecae shorter than the copulatory ducts, and the horizontal spermathecae ventrally curved (
<figureCitation id="A8FF2A0AFF9EFFFB8E3FF9FDFD1B760B" box="[629,728,1569,1595]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF9EFFF88C8DF999FB897229" lastPageId="30" lastPageNumber="181" pageId="29" pageNumber="180" type="description">
<paragraph id="307B368FFF9EFFFB8C8DF999FDE877F7" blockId="29.[151,1437,1029,1992]" pageId="29" pageNumber="180">
<emphasis id="02B0EA9DFF9EFFFB8C8DF999FE9A7650" bold="true" box="[199,345,1605,1632]" pageId="29" pageNumber="180">Description.</emphasis>
Male (CAS 9025935): Coloration: carapace orange, palp brown and femora black, other leg segments yellowish brown. Total length 4.9. Carapace: length 2.25, width 1.65, height 0.68. Opisthosoma: length 2.25, width 2.2. Clypeus: height 0.16. Eye diameters and interdistances: AME 0.05; ALE 0.10; PME 0.08; PLE 0.06; AME–AME 0.11; AME–ALE 0.11; ALE–PLE 0.05; PME–PLE 0.2; PME–PME 0.13; AME–PME 0.13. Chelicerae paturon: length 0.10. Leg measurements: I: Femur 1.73/ patella 0.65/ tibia 1.68/ metatarsus 1.63/ tarsus 0.88/ total 6.55; II: 1.75/ 0.65/ 1.63/ 1.45/ 0.68/ 6.15; III: 1.50/ 0.63/ 1.38/ 1.23/ 0.68/ 5.40; IV: 1.70/ 0.68/ 1.80/ 1.45/ 0.63/ 6.25; Leg formula: 1423; Leg spination: femur I–II: p1ap; tibia II v0-0-1, p1ap; III v1-0-1 or v1-0-2, p1ap, r1ap; IV v1-0-1 or v0-1-2, p1ap, r1ap; metatarsus I v2-2-
<quantity id="F73C9B6AFF9EFFFB8EF0F89DFD1D776B" box="[698,734,1857,1883]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="180" unit="m" value="1.0">1m</quantity>
, p1ap, r1ap; II v2-2-1map, r1ap, p2ap; III v2-2-
<quantity id="F73C9B6AFF9EFFFB894AF89DFAE0776B" box="[1280,1315,1857,1883]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="180" unit="m" value="1.0">1m</quantity>
, p1-1-1ap, r0-0-1ap, d1-0-1rap or d0-0-1p; IV v2-2-2-
<quantity id="F73C9B6AFF9EFFFB8EC2F8B9FD68774F" box="[648,683,1893,1919]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="180" unit="m" value="1.0">1m</quantity>
, p0-2-2ap, r1ap, d2ap. Patellar apophysis dorsally projected (
<figureCitation id="A8FF2A0AFF9EFFFB8925F8B9FF2F7793" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18B–D</figureCitation>
). MLT with apex membranous (
<figureCitation id="A8FF2A0AFF9EFFFB8E1DF855FD7D7793" box="[599,702,1929,1955]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18C</figureCitation>
). Spermatic duct bending at 12 o’clock.
<emphasis id="02B0EA9DFF9EFFFB88C2F856FADD7793" box="[1160,1310,1929,1955]" italics="true" pageId="29" pageNumber="180">Pars pendula</emphasis>
thickening at the tip of the embolus (
<figureCitation id="A8FF2A0AFF9EFFFB8DFCF871FDDD77F7" box="[438,542,1965,1991]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="29" pageNumber="180">Fig. 18D</figureCitation>
).
</paragraph>
<paragraph id="307B368FFF9DFFF88C8DFF4BFE5D71C5" blockId="30.[151,1437,150,573]" pageId="30" pageNumber="181">
Female (CAS 9025998). Coloration: carapace brown with the edge darker; chelicerae, labium sternum and endites brown; coxae and femora brown; other leg segments yellowish brown; opisthosoma pale brown. Total length: 5.40. Carapace: length 2.1, width 1.6, height 1.03. Opisthosoma: length 3.5, width 2.2. Clypeus: height 0.18. Eye diameters and interdistances: AME 0.07; ALE 0.11; PME 0.10; PLE 0.10; AME–AME 0.10; AME–ALE 0.13; ALE–PLE 0.06; PME–PLE 0.23; PME–PME 0.15; AME–PME 0.11. Chelicerae paturon: length 1.06. Leg measurements: I: femur 1.56/ patella 0.59/ tibia 1.53/ metatarsus 1.22/ tarsus 0.72/ total 5.63; II: 1.47/ 0.69/ 1.34/ 1.09/ 0.69/ 5.28; III: 1.34/ 0.66/ 0.97/ 0.94/ 0.50/ 4.41; IV: 1.69/ 0.75/ 1.47/ 1.09/ 0.53/ 5.53. Leg formula: 1423. Leg spination: femur I p1ap; metatarsus I–III v1map, p1ap, r1ap; IV v0-1-1map, r1ap, p1ap. Median field of epigynum wider than long but not covering the copulatory openings (
<figureCitation id="A8FF2A0AFF9DFFF88F69FE6BFC4471E1" box="[803,903,439,465]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="30" pageNumber="181">Fig. 18E</figureCitation>
). Fertilization ducts apart from each other twice its length (
<figureCitation id="A8FF2A0AFF9DFFF88D44FE07FE5271C5" box="[270,401,475,501]" captionStart="FIGURE 18" captionStartId="30.[151,250,1852,1876]" captionTargetBox="[158,1430,606,1822]" captionTargetId="figure-192@30.[151,1436,597,1829]" captionTargetPageId="30" captionText="FIGURE 18.A–F. Goeldia obscura (Keyserling, 1878).A–D. Male left palp; E–F Female Epigynum:A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847423" httpUri="https://zenodo.org/record/10847423/files/figure.png" pageId="30" pageNumber="181">Fig. 18E–F</figureCitation>
).
</paragraph>
<paragraph id="307B368FFF9DFFF88C8DFE22FB897229" blockId="30.[151,1437,150,573]" box="[199,1098,510,537]" pageId="30" pageNumber="181">
<emphasis id="02B0EA9DFF9DFFF88C8DFE22FEFC7229" bold="true" box="[199,319,510,537]" pageId="30" pageNumber="181">Variation.</emphasis>
Length (
<specimenCount id="26C2FD06FF9DFFF88DEFFE23FE1E7229" box="[421,477,511,537]" count="10" pageId="30" pageNumber="181" type="female">10 ♀</specimenCount>
): total 4.82–6.15; carapace 1.8–2.6; femur I 1.48–1.70.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FF9DFFF88C8DFDFEFD85720D" box="[199,582,546,573]" pageId="30" pageNumber="181" type="distribution">
<paragraph id="307B368FFF9DFFF88C8DFDFEFD85720D" blockId="30.[151,1437,150,573]" box="[199,582,546,573]" pageId="30" pageNumber="181">
<emphasis id="02B0EA9DFF9DFFF88C8DFDFEFEA3720D" bold="true" box="[199,352,546,573]" pageId="30" pageNumber="181">Distribution.</emphasis>
<collectingCountry id="48D3761FFF9DFFF88D2DFDFFFE16720D" box="[359,469,547,573]" name="Colombia" pageId="30" pageNumber="181">Colombia</collectingCountry>
(
<figureCitation id="A8FF2A0AFF9DFFF88DAFFDFFFDF9720D" box="[485,570,546,573]" captionStart="FIGURE 25" captionStartId="39.[151,250,1041,1065]" captionTargetBox="[184,1404,188,1011]" captionTargetId="figure-21@39.[177,1411,181,1018]" captionTargetPageId="39" captionText="FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities." figureDoi="http://doi.org/10.5281/zenodo.10847445" httpUri="https://zenodo.org/record/10847445/files/figure.png" pageId="30" pageNumber="181">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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