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<document id="34542A55C2F20094E8E40AF3B4DAD9CD" ID-CLB-Dataset="276470" ID-DOI="10.1111/j.1096-3642.2012.00860.x" ID-GBIF-Dataset="cc1867a9-801f-40ae-9613-e41e91dfa8fd" ID-ISSN="0024-4082" ID-Zenodo-Dep="5409033" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630349847782" checkinUser="marcus" docAuthor="Edgecombe, Gregory D., Vahtera, Varpu, Stock, Stuart R., Kallonen, Aki, Xiao, Xianghui, Rack, Alexander &amp; Giribet, Gonzalo" docDate="2012" docId="A37287B67F513840FCCDFE66FB46FA25" docLanguage="en" docName="zoj860.pdf" docOrigin="Zoological Journal of the Linnean Society 166 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2012.00860.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Scolopocryptops simojovelensis Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet 2012, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="776" masterDocId="5F4BFFCE7F543848FFB3FFB7FFA8FFE7" masterDocTitle="A scolopocryptopid centipede (Chilopoda: Scolopendromorpha) from Mexican amber: synchrotron microtomography and phylogenetic placement using a combined morphological and molecular data set" masterLastPageNumber="786" masterPageNumber="768" pageNumber="773" updateTime="1699245749930" updateUser="ExternalLinkService">
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<mods:title id="4ED508416F415EF5DFF461012198D36F">A scolopocryptopid centipede (Chilopoda: Scolopendromorpha) from Mexican amber: synchrotron microtomography and phylogenetic placement using a combined morphological and molecular data set</mods:title>
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<taxonomicName id="ECDB4D237F51384DFCCDFE66FA9EFE01" authority="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet, 2012" authorityName="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet" authorityYear="2012" box="[894,1334,465,486]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="5" pageNumber="778" phylum="Arthropoda" rank="species" species="simojovelensis" status="sp. nov.">SCOLOPOCRYPTOPS SIMOJOVELENSIS</taxonomicName>
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(
<figureCitation id="B3E02A257F51384DFBEDFE45FB6CFDEF" box="[1118,1220,498,520]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="6.[164,243,1324,1343]" captionStartId-1="7.[144,223,1510,1529]" captionStartId-2="8.[164,243,925,944]" captionTargetBox-0="[323,1283,196,1293]" captionTargetBox-1="[303,1263,194,1480]" captionTargetBox-2="[422,1186,199,893]" captionTargetId-0="figure-225@6.[323,1283,196,1293]" captionTargetId-1="figure-120@7.[303,1263,194,1480]" captionTargetId-2="figure-450@8.[419,1188,196,895]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." captionText-1="Figure 2. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm." captionText-2="Figure 3. Scolopocryptops simojovelensis sp. nov. Visualizations of synchrotron tomography data of holotype. A, B, dorsolateral and oblique anterodorsal views of head. C, ventral view of forcipules. D, lateral view of coxopleuron of left leg 23, anterior to left. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5409035" figureDoi-1="http://doi.org/10.5281/zenodo.5409037" figureDoi-2="http://doi.org/10.5281/zenodo.5409039" httpUri-0="https://zenodo.org/record/5409035/files/figure.png" httpUri-1="https://zenodo.org/record/5409037/files/figure.png" httpUri-2="https://zenodo.org/record/5409039/files/figure.png" pageId="5" pageNumber="773">FIGS 1–3</figureCitation>
)
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Diagnosis: 
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lacking margination on cephalic plate; paramedian sutures terminating on tergite 21; anterior margin of forcipular coxosternite lacking lateral tooth or bulge, median part of margin projecting anterior to lateral part; coxopleural process long, terminating in a robust pointed spine; two tibial spurs on legs 1–19, ventral spur only on leg 20; accessory spurs on legs 1–22 
<emphasis id="19AFEAB27F51384DFB24FD58FB0AFCE3" box="[1175,1186,751,772]" italics="true" pageId="5" pageNumber="773">c</emphasis>
. 20% length of the pretarsus.
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<materialsCitation id="9BB33CFD7F51384DFC95FCE3FB47FC40" ID-GBIF-Occurrence="4435344301" collectionCode="AMNH" collectorName="de Allende" country="Mexico" location="American Museum of Natural History" municipality="Simojovel de Allende" pageId="5" pageNumber="773" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
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:
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Ch-SH7, 
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amber arthropod collection, from 
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Simojovel 
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, 
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, 
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.
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<emphasis id="19AFEAB27F51384DFC95FC6FFC0CFC0A" box="[806,932,984,1005]" italics="true" pageId="5" pageNumber="773">Etymology:</emphasis>
For Simojovel de Allende, the most prolific source of 
<collectingRegion id="E91FF8427F51384DFC24FC41FC5CFBEB" box="[919,1012,1014,1036]" country="Mexico" name="Chiapas" pageId="5" pageNumber="773">Chiapas</collectingRegion>
amber.
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<emphasis id="19AFEAB27F51384DFC95FB89FC06FBB4" box="[806,942,1086,1107]" italics="true" pageId="5" pageNumber="773">Description:</emphasis>
Length of body (anterior margin of cephalic plate to posterior margin of tergite 23) 
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. Cephalic plate lacking margination either laterally or posteriorly, its posterior margin overlying tergite 1 (
<figureCitation id="B3E02A257F51384DFC10FB0FFC54FB2A" box="[931,1020,1208,1230]" captionStart="Figure 1" captionStartId="6.[164,243,1324,1343]" captionTargetBox="[323,1283,196,1293]" captionTargetId="figure-225@6.[323,1283,196,1293]" captionTargetPageId="6" captionText="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5409035" httpUri="https://zenodo.org/record/5409035/files/figure.png" pageId="5" pageNumber="773">Figs 1C</figureCitation>
, 
<figureCitation id="B3E02A257F51384DFBBCFB0FFB99FB29" box="[1039,1073,1208,1230]" captionStart="Figure 3" captionStartId="8.[164,243,925,944]" captionTargetBox="[422,1186,199,893]" captionTargetId="figure-450@8.[419,1188,196,895]" captionTargetPageId="8" captionText="Figure 3. Scolopocryptops simojovelensis sp. nov. Visualizations of synchrotron tomography data of holotype. A, B, dorsolateral and oblique anterodorsal views of head. C, ventral view of forcipules. D, lateral view of coxopleuron of left leg 23, anterior to left. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409039" httpUri="https://zenodo.org/record/5409039/files/figure.png" pageId="5" pageNumber="773">3A</figureCitation>
). Seventeen articles in right antenna (left antenna preserves only a few basal articles, the remainder eroded from the inclusion); moderately long setae numerous over length of articles to at least article 8; articles 2 and 3 bearing a similar, moderate number of setae dorsally (neither is sparsely setose); short, dense setae on article 4 and more distal parts of antenna.
</paragraph>
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Anterior margin of forcipular coxosternite partly covered (
<figureCitation id="B3E02A257F51384DFC3DFA7CFC4AFA06" box="[910,994,1483,1505]" captionStart="Figure 3" captionStartId="8.[164,243,925,944]" captionTargetBox="[422,1186,199,893]" captionTargetId="figure-450@8.[419,1188,196,895]" captionTargetPageId="8" captionText="Figure 3. Scolopocryptops simojovelensis sp. nov. Visualizations of synchrotron tomography data of holotype. A, B, dorsolateral and oblique anterodorsal views of head. C, ventral view of forcipules. D, lateral view of coxopleuron of left leg 23, anterior to left. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409039" httpUri="https://zenodo.org/record/5409039/files/figure.png" pageId="5" pageNumber="773">Fig. 3C</figureCitation>
), only visible on lateral half of right side; exposed part of margin approximately straight, its more medial part anterior to its lateral part, lacking lateral tooth or lateral bulge.
</paragraph>
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Complete pair of paramedian sutures on tergites (TT)16-21 (
<figureCitation id="B3E02A257F51384DFC02F9D3FB83F99E" box="[945,1067,1636,1658]" captionStart="Figure 2" captionStartId="7.[144,223,1510,1529]" captionTargetBox="[303,1263,194,1480]" captionTargetId="figure-120@7.[303,1263,194,1480]" captionTargetPageId="7" captionText="Figure 2. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409037" httpUri="https://zenodo.org/record/5409037/files/figure.png" pageId="5" pageNumber="773">Fig. 2A, B</figureCitation>
), anterior to this to at least T8 (and apparently as far as T5) a pair of lines of surficial crust run parallel to the course of the paramedian sutures on TT16-21 and evidently represent deposits along the sutures; paramedian sutures absent on TT1-3 and 22. Posterior part of tergites bearing several subtransverse anastomosing ridges (
<figureCitation id="B3E02A257F51384DFC9DF88DFC2BF8B7" box="[814,899,1850,1872]" captionStart="Figure 2" captionStartId="7.[144,223,1510,1529]" captionTargetBox="[303,1263,194,1480]" captionTargetId="figure-120@7.[303,1263,194,1480]" captionTargetPageId="7" captionText="Figure 2. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409037" httpUri="https://zenodo.org/record/5409037/files/figure.png" pageId="5" pageNumber="773">Fig. 2B</figureCitation>
), more prominent on posterior trunk segments to 
<emphasis id="19AFEAB27F51384DFC25F8EEFC00F889" box="[918,936,1881,1902]" italics="true" pageId="5" pageNumber="773">c.</emphasis>
segment 21.
</paragraph>
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<emphasis id="19AFEAB27F52384EFF17FA9BFD5FFAD8" bold="true" box="[164,759,1323,1343]" pageId="6" pageNumber="774">
Figure 1. 
<taxonomicName id="ECDB4D237F52384EFEAAFA9CFD21FAD9" authority="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet, 2012" authorityName="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet" authorityYear="2012" box="[281,649,1323,1342]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="774" phylum="Arthropoda" rank="species" species="simojovelensis" status="sp. nov.">
<emphasis id="19AFEAB27F52384EFEAAFA9CFD21FAD9" bold="true" box="[281,649,1323,1342]" italics="true" pageId="6" pageNumber="774">Scolopocryptops simojovelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="029C57C97F52384EFD2BFA9BFD5FFAD8" box="[664,759,1324,1343]" pageId="6" pageNumber="774" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm.
</paragraph>
</caption>
<paragraph id="2B6436A07F52384EFF0FFA7CFC3BF9DA" blockId="6.[164,779,1483,1903]" lastBlockId="6.[826,1441,1483,1597]" pageId="6" pageNumber="774">
Paired tibial spurs on legs 1–19, smaller anterodorsal spur and larger ventral spur; ventral spur only on leg 20 (
<figureCitation id="B3E02A257F52384EFF4FF9BFFEE7F9F9" box="[252,335,1544,1566]" captionStart="Figure 1" captionStartId="6.[164,243,1324,1343]" captionTargetBox="[323,1283,196,1293]" captionTargetId="figure-225@6.[323,1283,196,1293]" captionTargetPageId="6" captionText="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5409035" httpUri="https://zenodo.org/record/5409035/files/figure.png" pageId="6" pageNumber="774">Fig. 1B</figureCitation>
), lacking on legs 21–23. Single tarsal spur on legs 1–21 (
<figureCitation id="B3E02A257F52384EFEC8F990FE58F9DA" box="[379,496,1575,1597]" captionStart="Figure 1" captionStartId="6.[164,243,1324,1343]" captionTargetBox="[323,1283,196,1293]" captionTargetId="figure-225@6.[323,1283,196,1293]" captionTargetPageId="6" captionText="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5409035" httpUri="https://zenodo.org/record/5409035/files/figure.png" pageId="6" pageNumber="774">Fig. 1B, C</figureCitation>
); tarsal spurs lacking on legs 22 and 23. Pretarsus of legs 1–22 with moderately developed pair of accessory spurs (
<figureCitation id="B3E02A257F52384EFDC3F9D3FD6DF99D" box="[624,709,1636,1658]" captionStart="Figure 1" captionStartId="6.[164,243,1324,1343]" captionTargetBox="[323,1283,196,1293]" captionTargetId="figure-225@6.[323,1283,196,1293]" captionTargetPageId="6" captionText="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5409035" httpUri="https://zenodo.org/record/5409035/files/figure.png" pageId="6" pageNumber="774">Fig. 1D</figureCitation>
); ultimate leg with small accessory spurs. Sternite of segment 23 with evenly concave posterior margin. Coxopleural pore field extending close to base of coxopleural process ventrad, posterior margin of pore field sinuous but without a re-entrant field devoid of pores dorsad (
<figureCitation id="B3E02A257F52384EFEFEF8ABFE0BF8D5" box="[333,419,1820,1842]" captionStart="Figure 3" captionStartId="8.[164,243,925,944]" captionTargetBox="[422,1186,199,893]" captionTargetId="figure-450@8.[419,1188,196,895]" captionTargetPageId="8" captionText="Figure 3. Scolopocryptops simojovelensis sp. nov. Visualizations of synchrotron tomography data of holotype. A, B, dorsolateral and oblique anterodorsal views of head. C, ventral view of forcipules. D, lateral view of coxopleuron of left leg 23, anterior to left. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409039" httpUri="https://zenodo.org/record/5409039/files/figure.png" pageId="6" pageNumber="774">Fig. 3D</figureCitation>
). Dorsomedial spinose process of ultimate leg prefemur more than half the length of strong ventral spinose process (
<figureCitation id="B3E02A257F52384EFD81F8EEFD1BF889" box="[562,691,1881,1903]" captionStart="Figure 2" captionStartId="7.[144,223,1510,1529]" captionTargetBox="[303,1263,194,1480]" captionTargetId="figure-120@7.[303,1263,194,1480]" captionTargetPageId="7" captionText="Figure 2. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409037" httpUri="https://zenodo.org/record/5409037/files/figure.png" pageId="6" pageNumber="774">Fig. 2C, D</figureCitation>
). Setal density on distal articles of ultimate leg (
<figureCitation id="B3E02A257F52384EFAF6FA7CFA31FA06" box="[1349,1433,1483,1505]" captionStart="Figure 2" captionStartId="7.[144,223,1510,1529]" captionTargetBox="[303,1263,194,1480]" captionTargetId="figure-120@7.[303,1263,194,1480]" captionTargetPageId="7" captionText="Figure 2. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409037" httpUri="https://zenodo.org/record/5409037/files/figure.png" pageId="6" pageNumber="774">Fig. 2D</figureCitation>
) apparently similarly sparse to that on proximal part (no evidence for clustered ‘bottle brush’ setae on any article).
</paragraph>
<paragraph id="2B6436A07F52384FFC89F9D2FC0AF888" blockId="6.[826,1442,1636,1903]" lastBlockId="7.[806,1421,1697,1903]" lastPageId="7" lastPageNumber="775" pageId="6" pageNumber="774">
<emphasis id="19AFEAB27F52384EFC89F9D2FC12F99D" box="[826,954,1637,1658]" italics="true" pageId="6" pageNumber="774">Discussion:</emphasis>
Although the new species is known from a single specimen, it exhibits distinct states for characters that, in combination, are routinely used in species diagnosis in 
<taxonomicName id="ECDB4D237F52384EFB97F977FB71F932" box="[1060,1241,1728,1749]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="6" pageNumber="774" phylum="Arthropoda" rank="genus">
<emphasis id="19AFEAB27F52384EFB97F977FB71F932" box="[1060,1241,1728,1749]" italics="true" pageId="6" pageNumber="774">Scolopocryptops</emphasis>
</taxonomicName>
. It is readily distinguished from all North American and Asian species of the genus by the absence of margination on the cephalic plate, a character that is invariant in species. Some of the states cited in the species diagnosis exhibit variability in geographically widespread extant species, but the condition in the fossil allows distinction from other species. For example, the paramedian sutures terminate on any of tergites 
<quantity id="EC239B457F53384FFD20F968FD5FF913" box="[659,759,1758,1780]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="7" pageNumber="775" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
<taxonomicName id="ECDB4D237F53384FFF23F94AFE8DF8F4" authorityName="Humbert &amp; Saussure" authorityYear="1869" box="[144,293,1789,1811]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="19AFEAB27F53384FFF23F94AFE8DF8F4" box="[144,293,1789,1811]" italics="true" pageId="7" pageNumber="775">S. mexicanus</emphasis>
</taxonomicName>
, but the state in the fossil (terminating on tergite 21) differs from that seen in 
<taxonomicName id="ECDB4D237F53384FFDFAF8ABFD71F8D6" baseAuthorityName="Kraepelin" baseAuthorityYear="1903" box="[585,729,1820,1841]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="macrodon">
<emphasis id="19AFEAB27F53384FFDFAF8ABFD71F8D6" box="[585,729,1820,1841]" italics="true" pageId="7" pageNumber="775">S. macrodon</emphasis>
</taxonomicName>
or 
<taxonomicName id="ECDB4D237F53384FFF23F88DFE08F8B7" authorityName="Newport" authorityYear="1845" box="[144,416,1850,1872]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="miersii">
<emphasis id="19AFEAB27F53384FFF23F88DFE08F8B7" box="[144,416,1850,1872]" italics="true" pageId="7" pageNumber="775">Scolopocryptops miersii</emphasis>
</taxonomicName>
(terminating on tergite 22). Likewise, the posterior-most leg with two tibial spurs may be either 18 or 
<quantity id="EC239B457F53384FFBB9F916FBECF950" box="[1034,1092,1697,1719]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="7" pageNumber="775" unit="in" value="19.0">19 in</quantity>
<taxonomicName id="ECDB4D237F53384FFBF8F915FB48F950" authorityName="Humbert &amp; Saussure" authorityYear="1869" box="[1099,1248,1698,1719]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="19AFEAB27F53384FFBF8F915FB48F950" box="[1099,1248,1698,1719]" italics="true" pageId="7" pageNumber="775">S. mexicanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="ECDB4D237F53384FFAAAF915FA2EF950" authorityName="Newport" authorityYear="1845" box="[1305,1414,1698,1719]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="miersii">
<emphasis id="19AFEAB27F53384FFAAAF915FA2EF950" box="[1305,1414,1698,1719]" italics="true" pageId="7" pageNumber="775">S. miersii</emphasis>
</taxonomicName>
, but the fossil (with two spurs on leg 19) is distinguished from 
<taxonomicName id="ECDB4D237F53384FFC75F968FBFDF914" baseAuthorityName="Kraepelin" baseAuthorityYear="1903" box="[966,1109,1758,1780]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="macrodon">
<emphasis id="19AFEAB27F53384FFC75F968FBFDF914" box="[966,1109,1758,1780]" italics="true" pageId="7" pageNumber="775">S. macrodon</emphasis>
</taxonomicName>
(two spurs on leg 20). The shape of the coxopleural process and the development of the pretarsal accessory spurs are stable within species, and, in combination, the states are diagnostic.
</paragraph>
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<paragraph id="2B6436A07F53384FFF23FA51FCE0F988" blockId="7.[144,1422,1510,1647]" pageId="7" pageNumber="775">
<emphasis id="19AFEAB27F53384FFF23FA51FD66FA1E" bold="true" box="[144,718,1510,1529]" pageId="7" pageNumber="775">
Figure 2. 
<taxonomicName id="ECDB4D237F53384FFEB7FA51FDC6FA1E" authority="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet, 2012" authorityName="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet" authorityYear="2012" box="[260,622,1510,1529]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="7" pageNumber="775" phylum="Arthropoda" rank="species" species="simojovelensis" status="sp. nov.">
<emphasis id="19AFEAB27F53384FFEB7FA51FDC6FA1E" bold="true" box="[260,622,1510,1529]" italics="true" pageId="7" pageNumber="775">Scolopocryptops simojovelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="029C57C97F53384FFDC5FA51FD66FA1E" box="[630,718,1510,1529]" pageId="7" pageNumber="775" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm.
</paragraph>
</caption>
<caption id="7FA466287F5C3840FF17FC2AFDD5FC0C" ID-DOI="http://doi.org/10.5281/zenodo.5409039" ID-Zenodo-Dep="5409039" httpUri="https://zenodo.org/record/5409039/files/figure.png" pageId="8" pageNumber="776" startId="8.[164,243,925,944]" targetBox="[422,1186,199,893]" targetPageId="8">
<paragraph id="2B6436A07F5C3840FF17FC2AFDD5FC0C" blockId="8.[164,1441,924,1003]" pageId="8" pageNumber="776">
<emphasis id="19AFEAB27F5C3840FF17FC2AFD4DFC57" bold="true" box="[164,741,924,944]" pageId="8" pageNumber="776">
Figure 3. 
<taxonomicName id="ECDB4D237F5C3840FEAAFC2BFD2BFC48" authority="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet, 2012" authorityName="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet" authorityYear="2012" box="[281,643,924,943]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="simojovelensis" status="sp. nov.">
<emphasis id="19AFEAB27F5C3840FEAAFC2BFD2BFC48" bold="true" box="[281,643,924,943]" italics="true" pageId="8" pageNumber="776">Scolopocryptops simojovelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="029C57C97F5C3840FD3FFC2AFD4DFC57" box="[652,741,925,944]" pageId="8" pageNumber="776" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Visualizations of synchrotron tomography data of holotype. A, B, dorsolateral and oblique anterodorsal views of head. C, ventral view of forcipules. D, lateral view of coxopleuron of left leg 23, anterior to left. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="2B6436A07F5C3840FF0FFBA8FB46FA25" blockId="8.[164,779,1055,1903]" lastBlockId="8.[826,1441,1055,1474]" pageId="8" pageNumber="776">
The fossil can be reliably distinguished from each of the five extant Neotropical species of 
<taxonomicName id="ECDB4D237F5C3840FDFCFB8AFCACFBB5" box="[591,772,1085,1106]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="genus">
<emphasis id="19AFEAB27F5C3840FDFCFB8AFCACFBB5" box="[591,772,1085,1106]" italics="true" pageId="8" pageNumber="776">Scolopocryptops</emphasis>
</taxonomicName>
. Perhaps the most pertinent comparisons are with the two most geographically widespread species, 
<taxonomicName id="ECDB4D237F5C3840FF17FB2EFDD5FB48" authority="Humbert &amp; Saussure, 1869" authorityName="Humbert &amp; Saussure" authorityYear="1869" box="[164,637,1177,1199]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="19AFEAB27F5C3840FF17FB2EFE91FB48" box="[164,313,1177,1199]" italics="true" pageId="8" pageNumber="776">S. mexicanus</emphasis>
Humbert &amp; Saussure, 1869
</taxonomicName>
, and 
<taxonomicName id="ECDB4D237F5C3840FD08FB2EFE76FB2A" authority="Newport, 1845" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="melanostoma">
<emphasis id="19AFEAB27F5C3840FD08FB2EFEB9FB2A" italics="true" pageId="8" pageNumber="776">S. melanostoma</emphasis>
Newport, 1845
</taxonomicName>
, because their modern distributions include the area inhabited by 
<taxonomicName id="ECDB4D237F5C3840FD28FB60FF54FAED" authorityName="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet" authorityYear="2012" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="simojovelensis">
<emphasis id="19AFEAB27F5C3840FD28FB60FF54FAED" italics="true" pageId="8" pageNumber="776">S. simojovelensis</emphasis>
</taxonomicName>
in the Miocene. Both of these species occur in southern 
<collectingCountry id="53CC76307F5C3840FEACFAA4FEC7FACE" box="[287,367,1299,1321]" name="Mexico" pageId="8" pageNumber="776">Mexico</collectingCountry>
and range throughout Central America and the Caribbean, throughout which they are the only extant members of 
<taxonomicName id="ECDB4D237F5C3840FD9CFAE6FE00FA62" authority="(Chagas Junior, 2008)" baseAuthorityName="Chagas Junior" baseAuthorityYear="2008" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="subFamily" subFamily="Scolopocryptopinae">
Scolopocryptopinae (
<bibRefCitation id="4F4A4B517F5C3840FF1FFAD8FE09FA62" author="Chagas Junior A" box="[172,417,1391,1413]" pageId="8" pageNumber="776" refId="ref9047" refString="Chagas Junior A. 2008. Revisao sistematica e analise filogenetica dos Scolopocryptopinae (Chilopoda, Scolopendromorpha). PhD thesis, Universidade Federal do Rio de Janeiro." type="book" year="2008">Chagas Juniór, 2008</bibRefCitation>
)
</taxonomicName>
. Compared to 
<taxonomicName id="ECDB4D237F5C3840FDE5FAC7FCA2FA63" authorityName="Nrewport" authorityYear="1845" box="[598,778,1391,1413]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="melanostoma">
<emphasis id="19AFEAB27F5C3840FDE5FAC7FCA2FA63" box="[598,778,1391,1413]" italics="true" pageId="8" pageNumber="776">S. melanostoma</emphasis>
</taxonomicName>
the fossil has more strongly developed pretarsal accessory spurs (
<figureCitation id="B3E02A257F5C3840FE95FA1BFED3FA24" box="[294,379,1452,1475]" captionStart="Figure 1" captionStartId="6.[164,243,1324,1343]" captionTargetBox="[323,1283,196,1293]" captionTargetId="figure-225@6.[323,1283,196,1293]" captionTargetPageId="6" captionText="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5409035" httpUri="https://zenodo.org/record/5409035/files/figure.png" pageId="8" pageNumber="776">Fig. 1D</figureCitation>
), has the medial part of the forcipular coxosternal margin anterior to the lateral part (versus the opposite in 
<taxonomicName id="ECDB4D237F5C3840FE7FFA5DFD28FA18" authorityName="Nrewport" authorityYear="1845" box="[460,640,1514,1535]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="melanostoma">
<emphasis id="19AFEAB27F5C3840FE7FFA5DFD28FA18" box="[460,640,1514,1535]" italics="true" pageId="8" pageNumber="776">S. melanostoma</emphasis>
</taxonomicName>
), possesses both dorsal and ventral tibial spurs on leg 19 (versus a ventral spur only on leg 
<quantity id="EC239B457F5C3840FE78F990FDA0F9DA" box="[459,520,1575,1597]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="8" pageNumber="776" unit="in" value="19.0">19 in</quantity>
<taxonomicName id="ECDB4D237F5C3840FDA1F990FD6EF9DB" authorityName="Nrewport" authorityYear="1845" box="[530,710,1575,1596]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="melanostoma">
<emphasis id="19AFEAB27F5C3840FDA1F990FD6EF9DB" box="[530,710,1575,1596]" italics="true" pageId="8" pageNumber="776">S. melanostoma</emphasis>
</taxonomicName>
), and has a ventral tibial spur on leg 20 (
<figureCitation id="B3E02A257F5C3840FD81F9F2FD2EF9BB" box="[562,646,1605,1628]" captionStart="Figure 1" captionStartId="6.[164,243,1324,1343]" captionTargetBox="[323,1283,196,1293]" captionTargetId="figure-225@6.[323,1283,196,1293]" captionTargetPageId="6" captionText="Figure 1. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, dorsolateral view of complete specimen. B, distal part of right leg 20, showing tibial spur (ti) and tarsal spur (ta). C, dorsolateral view of cephalic plate and right antenna. D, distal part of tarsus and pretarsus of right leg 20, showing accessory spurs (ac). Scale bars: A = 5 mm; B = 0.5 mm; C = 1 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5409035" httpUri="https://zenodo.org/record/5409035/files/figure.png" pageId="8" pageNumber="776">Fig. 1B</figureCitation>
). Following 
<bibRefCitation id="4F4A4B517F5C3840FF17F9D3FE08F99D" author="Chagas Junior A" box="[164,416,1636,1658]" pageId="8" pageNumber="776" refId="ref9047" refString="Chagas Junior A. 2008. Revisao sistematica e analise filogenetica dos Scolopocryptopinae (Chilopoda, Scolopendromorpha). PhD thesis, Universidade Federal do Rio de Janeiro." type="book" year="2008">Chagas Juniór (2008)</bibRefCitation>
, we apply the name 
<taxonomicName id="ECDB4D237F5C3840FD25F9D3FF65F97F" authorityName="Humbert &amp; Saussure" authorityYear="1869" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="19AFEAB27F5C3840FD25F9D3FF65F97F" italics="true" pageId="8" pageNumber="776">S. mexicanus</emphasis>
</taxonomicName>
to Neotropical material that has generally been identified as 
<taxonomicName id="ECDB4D237F5C3840FEE5F916FF4EF931" authority="(Linne, 1767)" baseAuthorityName="Linne" baseAuthorityYear="1767" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="ferrugineus">
<emphasis id="19AFEAB27F5C3840FEE5F916FD0AF951" box="[342,674,1697,1718]" italics="true" pageId="8" pageNumber="776">Scolopocryptops ferrugineus</emphasis>
(Linné, 1767)
</taxonomicName>
. Compared to 
<taxonomicName id="ECDB4D237F5C3840FE23F977FD8DF932" authorityName="Humbert &amp; Saussure" authorityYear="1869" box="[400,549,1728,1749]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="19AFEAB27F5C3840FE23F977FD8DF932" box="[400,549,1728,1749]" italics="true" pageId="8" pageNumber="776">S. mexicanus</emphasis>
</taxonomicName>
, the 
<collectingRegion id="E91FF8427F5C3840FDD0F977FD68F931" box="[611,704,1728,1750]" country="Mexico" name="Chiapas" pageId="8" pageNumber="776">Chiapas</collectingRegion>
specimen lacks a lateral tooth on the forcipular coxosternal margin, and has a substantially longer coxopleural process (both a longer spinose distal part as well as a more sloping posterior margin when seen in lateral view; 
<figureCitation id="B3E02A257F5C3840FF5CF8EEFEEBF889" box="[239,323,1881,1903]" captionStart="Figure 2" captionStartId="7.[144,223,1510,1529]" captionTargetBox="[303,1263,194,1480]" captionTargetId="figure-120@7.[303,1263,194,1480]" captionTargetPageId="7" captionText="Figure 2. Scolopocryptops simojovelensis sp. nov. Holotype AMNH Ch-SH7. A, nearly dorsal view of tergites 17–22; arrows on TT17 and 18 indicate complete paramedian sutures. B, dorsolateral view of tergites 17–20; inset shows anastomizing ridges parallel to posterior margin on tergite 19. C, dorsal view of segment 23, showing tergite (T23), coxopleural process (cp), dorsomedial spinose process (ds) and ventral spinose process (vs) of prefemur. D, dorsolateral view of leg pairs 21–23. Scale bars: A, B, D = 1 mm; C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409037" httpUri="https://zenodo.org/record/5409037/files/figure.png" pageId="8" pageNumber="776">Fig. 2C</figureCitation>
; 
<bibRefCitation id="4F4A4B517F5C3840FEEBF8EEFE52F889" author="Attems CG" box="[344,506,1881,1902]" pageId="8" pageNumber="776" refId="ref8860" refString="Attems CG. 1930. Myriapoda 2. Scolopendromorpha. Berlin: Walter de Gruyter." type="book" year="1930">Attems, 1930</bibRefCitation>
: fig. 347 for 
<taxonomicName id="ECDB4D237F5C3840FD2CF8EEFCDEFBD3" baseAuthorityName="Linne" baseAuthorityYear="1767" box="[671,886,1055,1902]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="ferrugineus">
<emphasis id="19AFEAB27F5C3840FD2CF8EEFCDEFBD3" box="[671,886,1055,1902]" italics="true" pageId="8" pageNumber="776">S. ferrugineus</emphasis>
</taxonomicName>
). It appears to differ from both of these species in that antennal articles 2 and 3 bear numerous setae dorsally. Compared to the Brazilian 
<taxonomicName id="ECDB4D237F5C3840FB5FFBEBFBCEFB77" authority="Chagas, 2004" authorityName="Chagas" authorityYear="2004" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="piauhyensis">
<emphasis id="19AFEAB27F5C3840FB5FFBEBFC69FB68" italics="true" pageId="8" pageNumber="776">Scolopocryptops piauhyensis</emphasis>
<bibRefCitation id="4F4A4B517F5C3840FC7AFBCDFBCEFB77" author="Chagas Junior A" box="[969,1126,1146,1168]" pageId="8" pageNumber="776" pagination="81 - 83" refId="ref9016" refString="Chagas Junior A. 2004. A new Scolopocryptopid centipede from northeastern Brazil (Chilopoda: Scolopendromorpha). Studies on Neotropical Fauna and Environment 39: 81 - 83." type="journal article" year="2004">Chagas, 2004</bibRefCitation>
</taxonomicName>
, the new species has a less concave posterior margin of the ultimate sternite, and that sternite does not conceal as much of the coxopleural pore field. 
<taxonomicName id="ECDB4D237F5C3840FC6AFB61FCDBFAEC" authority="Manfredi, 1957" authorityName="Manfredi" authorityYear="1957" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="guacharensis">
<emphasis id="19AFEAB27F5C3840FC6AFB61FA82FB0C" box="[985,1322,1238,1259]" italics="true" pageId="8" pageNumber="776">Scolopocryptops guacharensis</emphasis>
Manfredi, 1957
</taxonomicName>
, from 
<collectingCountry id="53CC76307F5C3840FC7CFB42FBEBFAEC" box="[975,1091,1269,1291]" name="Venezuela" pageId="8" pageNumber="776">Venezuela</collectingCountry>
(revised by 
<bibRefCitation id="4F4A4B517F5C3840FB58FB42FC28FACE" author="Chagas Junior A" pageId="8" pageNumber="776" pagination="65 - 67" refId="ref8978" refString="Chagas Junior A. 2003 b. A review of the status of Scolopocryptops ferrugineus guacharensis (Chilopoda: Scolopendromorpha: Scolopocryptopidae) from Venezuela. Boletin de la Sociedad Entomologica Aragonesa 33: 65 - 67." type="journal article" year="2003">Chagas Juniór, 2003b</bibRefCitation>
) has a much smaller dorsomedial spinose process on the ultimate leg prefemur, possesses an external tooth on the forcipular coxosternal margin, has paramedian sutures terminating on tergite 20 (versus 
<quantity id="EC239B457F5C3840FC2AFA39FC7FFA43" box="[921,983,1422,1444]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="8" pageNumber="776" unit="in" value="21.0">21 in</quantity>
<taxonomicName id="ECDB4D237F5C3840FC57FA39FB0BFA44" authorityName="Edgecombe &amp; Vahtera &amp; Stock &amp; Kallonen &amp; Xiao &amp; Rack &amp; Giribet" authorityYear="2012" box="[996,1187,1422,1443]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="8" pageNumber="776" phylum="Arthropoda" rank="species" species="simojovelensis">
<emphasis id="19AFEAB27F5C3840FC57FA39FB0BFA44" box="[996,1187,1422,1443]" italics="true" pageId="8" pageNumber="776">S. simojovelensis</emphasis>
</taxonomicName>
), and has a strongly setose tibia and tarsus on leg pair 23.
</paragraph>
</subSubSection>
</treatment>
</document>