226 lines
18 KiB
XML
226 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.363.6482" ID-GBIF-Dataset="36e5551d-3881-45a2-810a-386e517c86fc" ID-PMC="PMC3867181" ID-Pensoft-Pub="1313-2970-363-1" ID-PubMed="24363601" ID-ZBK="780695A9CB2E4FECA0560BA6F456185E" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-363-1" ModsDocOrigin="ZooKeys 363" ModsDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" checkinTime="1451246560986" checkinUser="pensoft" docAuthor="Mathis, Wayne N. & Zatwarnicki, Tadeusz" docDate="2013" docId="7874544450583BE86D5D2E014C9EFBA2" docLanguage="en" docName="ZooKeys 363: 1-161" docOrigin="ZooKeys 363" docSource="http://dx.doi.org/10.3897/zookeys.363.6482" docTitle="Hydrochasma annae Mathis & Zatwarnicki, 2013, sp. n." docType="treatment" docUuid="ED9D3074-6AEF-428D-AEEB-A9A6DC9211D1" docUuidSource="ZooBank" docVersion="4" lastPageNumber="76" masterDocId="FFF2FFCA0333765855689824FFCDB839" masterDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="73" updateTime="1668157177207" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>363</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>161</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.363.6482</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.363.6482</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-363-1</mods:identifier>
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<mods:identifier type="ZBK">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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<mods:identifier type="ZooBank">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050156" LSID="urn:lsid:zoobank.org:act:ED9D3074-6AEF-428D-AEEB-A9A6DC9211D1" httpUri="http://treatment.plazi.org/id/7874544450583BE86D5D2E014C9EFBA2" lastPageId="75" lastPageNumber="76" pageId="72" pageNumber="73">
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<subSubSection pageId="72" pageNumber="73" type="nomenclature">
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<paragraph pageId="72" pageNumber="73">
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25.
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<taxonomicName LSID="http://zoobank.org/ED9D3074-6AEF-428D-AEEB-A9A6DC9211D1" class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
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<taxonomicNameLabel pageId="72" pageNumber="73">sp. n.</taxonomicNameLabel>
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Figs 137-141
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="diagnosis">
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<paragraph pageId="72" pageNumber="73">Diagnosis.</paragraph>
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<paragraph lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
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This species is distinguished from congeners by the following combination of characters: Small to moderately small shore flies, body length 1.75-2.20 mm.
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<pageBreakToken pageId="73" pageNumber="74" start="start">Head</pageBreakToken>
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: Antenna mostly dark gray; parafacial silvery white, concolorous with facial coloration. Gena moderately high, height usually slightly higher than length of basal flagellomere; gena-to-eye ratio 0.24-0.26. Thorax: Wing hyaline with costal vein ratio 0.75-0.76; M vein ratio 0.51-0.53. Forecoxa gray to silvery gray; hindtibia with only pale, yellowish setae ventroapically. Abdomen: Tergites 2-4 with brownish gray, relatively narrow, medial stripe, lateral margin of darkened areas lacking wedge-like markings; tergite 5 of male gray. Male terminalia (Figs 137-140): Combined structures generally elongate, in posterior view (Fig. 137) height about 2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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width, narrowly diamond shaped; epandrium with dorsal arch above cerci attenuate, not connected, becoming slightly wider to dorsal 1/3, thereafter ventrally tapered to shallowly bilobed ventral apex, generally bearing setulae, in lateral view (Fig. 138) mostly parallel sided with anterior shallowly sinuous, apical 1/3 tapered to moderately rounded apex; cerci moderately long, height slightly more than twice width (Figs 137); aedeagus in lateral view (Fig. 140) very elongate, narrow, with length of sclerotized, basal portion about 10
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<normalizedToken originalValue="×">x</normalizedToken>
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or more width, apical, membranous portion not greatly expanded or ballooned apically, at most slightly swollen, in ventral view (Fig. 139) also narrow and very elongate, almost parallel sided; phallapodeme in lateral view (Fig. 140) narrow, elongate, unevenly clavate toward hypandrium, keel slightly developed, apical width, including keel, slightly more than greatest aedeagal width, in ventral view (Fig. 139) with hypandrial end narrowly as a double T (2 bars), thereafter toward base of aedeagus bar-like, parallel sided; gonite in lateral view (Fig. 140) narrow, elongate, very shallowly sinuous, in ventral view (Fig. 139) narrow, bar-like with apices slightly curved in opposite directions; hypandrium in lateral view (Fig. 140) elongate, moderately shallow, width slightly more than 1/4 length, very shallowly curved, in ventral view (Fig. 139) generally V-shaped with posterior margin very deeply V-shaped, anterior margin moderately narrow, truncate.
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</paragraph>
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<caption pageId="73" pageNumber="74">
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<paragraph pageId="73" pageNumber="74">
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Figures 137-140.
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
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sp. n. (USA. Utah. Grand: Green River) 137 epandrium and cerci, posterior view 138 same, lateral view 139 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 140 same, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="73" pageNumber="74" type="type material">
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<paragraph pageId="73" pageNumber="74">Type material.</paragraph>
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<paragraph pageId="73" pageNumber="74">
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The holotype male of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
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is labeled "USA [UNITED STATES]. UT. Grand: Green River (15.3 km N;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.116665">39°7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.11">110°6.6'W</geoCoordinate>
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; 1255 m), 30 Jul 2007, D.&W.N.Mathis/USNM ENT 00117970 [plastic bar code label]/HOLOTYPE ♂
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
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Mathis & Zatwarnicki, USNM [red]." The holotype is double mounted (minuten in a block of plastic), is in excellent condition, and is deposited in the USNM. Seven paratypes (3♂, 4♀; USNM) bear the same label data as the holotype. Other paratypes are as follows: UTAH. Emery: Green River (3 km N;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.983334">38°59'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-110.15">110°09'W</geoCoordinate>
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), 30 Jul-5 Aug 1992, 2007, D. and W. N. Mathis (1♂, 3♀; USNM); Green River (Green River;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.993332">38°59.6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.14167">110°08.5'W</geoCoordinate>
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; 1240 m), 19-31 Jul 1988, 2007, D. and W. N. Mathis (1♂, 2♀; USNM). Grand: Thompson Spring (8.9 km N Thompson Springs;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.038334">39°02.3'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-109.723335">109°43.4'W</geoCoordinate>
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; 1740 m), 1-16 Aug 2007, 2008, D. and W. N. Mathis (3♂, 4♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="73" pageNumber="74" type="type locality">
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<paragraph pageId="73" pageNumber="74">Type locality.</paragraph>
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<paragraph pageId="73" pageNumber="74">
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United States. Utah. Grand: Swasey Beach (15.3 km N Green River;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="39.116665">39°07'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-110.11">110°06.6'W</geoCoordinate>
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; Green River; 1255 m).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="75" lastPageNumber="76" pageId="73" pageNumber="74" type="other specimens examined">
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<paragraph pageId="73" pageNumber="74">Other specimens examined.</paragraph>
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<paragraph pageId="73" pageNumber="74">
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Nearctic. UNITED STATES. CALIFORNIA. Imperial: Yuma (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="32.733334">32°44'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-114.61667">114°37'W</geoCoordinate>
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), 3-5 May 1918, J. C. Bradley (1♂, 2♀; CU).
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</paragraph>
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<paragraph pageId="73" pageNumber="74">
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COLORADO. Montezuma: McPhee Reservoir (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.495">37°29.7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-108.55">108°33'W</geoCoordinate>
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; 2115 m), 3 Aug 2007, D. and W. N. Mathis (1♀; USNM).
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</paragraph>
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<paragraph pageId="74" pageNumber="75">
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<pageBreakToken pageId="74" pageNumber="75" start="start">MISSOURI</pageBreakToken>
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. Boone: Columbia (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.951668">38°57.1'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-92.333336">92°20'W</geoCoordinate>
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), 26 May-8 Jun 1906, C. R. Crosby (1♂, 6♀; ANSP). Jackson: Atherton (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.18667">39°11.2'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-94.305">94°18.3'W</geoCoordinate>
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), (1♀; ANSP).
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</paragraph>
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<paragraph pageId="74" pageNumber="75">
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NEW MEXICO. Catron: Gila River (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="33.226665">33°13.6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.25166">106°15.1'W</geoCoordinate>
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; 1750 m), 15 Aug 2007, D. and W. N. Mathis (1♂; USNM). Grant: Mimbres River (NM Highway 61 and Royal John Mine Road;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="32.73">32°43.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-107.86667">107°52'W</geoCoordinate>
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; 1665 m), 1-22 Aug 2008, 2009, D. and W. N. Mathis, T. Zatwarnicki (33♂, 13♀; USNM). Lincoln: Capitan (3.2 km E; Salado Creek;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="33.543335">33°32.6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-105.53833">105°32.3'W</geoCoordinate>
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; 1890 m), 10 Aug 2007, D. and W. N. Mathis (1♂; USNM). Sandoval: La Cueva (Junction of Highways 126 & 4;
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="35.866665">35°52'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.64">106°38.4'W</geoCoordinate>
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; 2342 m), 15 Jun 2011, D. and W. N. Mathis (2♂; USNM). Valencia:
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Puerco (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="34.796665">34°47.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-106.99167">106°59.5'W</geoCoordinate>
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; 1575 m), 9 Aug 2007, D. and W. N. Mathis (1♂; USNM).
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</paragraph>
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<paragraph pageId="74" pageNumber="75">
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SOUTH DAKOTA. Bon Homme: Springfield (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.853333">42°51.2'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-97.89667">97°53.8'W</geoCoordinate>
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), 26 Jun 1924 (1♂, 1♀; ANSP).
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</paragraph>
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<paragraph pageId="74" pageNumber="75">
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TEXAS. Galveston: Galveston Island (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="25.166666">25°10'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-95.083336">95°05'W</geoCoordinate>
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), 14 May 1993, D. and W. N. Mathis (2♀; USNM). St. Augustine: Rayburn Park (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="31.066668">31°04'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-94.083336">94°05'W</geoCoordinate>
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), 15 May 1993, D. and W. N. Mathis (3♂, 2♀; USNM).
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</paragraph>
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<paragraph pageId="75" pageNumber="76">
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<pageBreakToken pageId="75" pageNumber="76" start="start">WASHINGTON</pageBreakToken>
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. Franklin: Columbia River N of Pasco (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.358334">46°21.5'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.25833">119°15.5'W</geoCoordinate>
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), 29 Jul 1998, W. N. Mathis (7♂, 1♀; USNM); Ringold (4.8 km N;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="46.515">46°30.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-119.255">119°15.3'W</geoCoordinate>
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), 2 Jul 1988, D. and W. N. Mathis (1♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="75" pageNumber="76" type="distribution">
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<paragraph pageId="75" pageNumber="76">Distribution</paragraph>
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<paragraph pageId="75" pageNumber="76">(Fig. 141). Nearctic: United States (California, Colorado, Missouri, New Mexico, South Dakota, Texas, Utah, Washington).</paragraph>
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<caption pageId="75" pageNumber="76">
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<paragraph pageId="75" pageNumber="76">
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Figure 141. Distribution of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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<caption pageId="75" pageNumber="76">
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<paragraph pageId="75" pageNumber="76">
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Figures 142-143. Abdomens of males, dorsal view. 142
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma dolabrutum" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="dolabrutum">Hydrochasma dolabrutum</taxonomicName>
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sp. n. (Jamaica. Clarendon: Barnswell Beach) 143
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma urnulum" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="urnulum">Hydrochasma urnulum</taxonomicName>
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sp. n. (Dominican Republic. LaVega: Jarabacoa). Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="75" pageNumber="76" type="etymology">
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<paragraph pageId="75" pageNumber="76">Etymology.</paragraph>
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<paragraph pageId="75" pageNumber="76">The species epithet, annae, is a feminine genitive patronym to recognize Ms. AnnaLee Thayn who inspired us while collecting in the Southwest.</paragraph>
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</subSubSection>
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<subSubSection pageId="75" pageNumber="76" type="remarks">
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<paragraph pageId="75" pageNumber="76">Remarks.</paragraph>
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<paragraph pageId="75" pageNumber="76">
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Externally, this species is very similar and evidently closely related to
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma andeum" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="andeum">Hydrochasma andeum</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae</taxonomicName>
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but can be distinguished from either of these two species by having a hindtibia with only yellowish setulae ventrally near the apex, less sinuous margins of the epandrium, and by having a more delicately developed hypandrium that is deeply V-shaped in ventral view (Fig. 139).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |