treatments-xml/data/5E/29/87/5E298782FFFFFFC43CA07EF2FEFDDADF.xml

<document id="35040C4DB1026A5071D583034B45578E" ID-CLB-Dataset="26146" ID-DOI="10.11646/zootaxa.4646.1.3" ID-GBIF-Dataset="de20b639-5800-47da-9214-99a33a9c13cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3346510" ID-ZooBank="EA93232D-5C20-48FC-BA26-E5DC811CFCC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563868669035" checkinUser="plazi" docAuthor="Quéinnec, Eric &amp; Ollivier, Eric" docDate="2019" docId="5E298782FFFFFFC43CA07EF2FEFDDADF" docLanguage="en" docName="zootaxa.4646.1.3.pdf.imf" docOrigin="Zootaxa 4646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxytrechus cayambeensis Quéinnec &amp; Ollivier 2019, n. sp." docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="A210FFFAFFF9FFCD3C377A06FFCEDE18" masterDocTitle="Two new Oxytrechus species from Ecuadorian páramo (Coleoptera, Carabidae, Trechinae)" masterLastPageNumber="66" masterPageNumber="55" pageNumber="61" updateTime="1698747107410" updateUser="ExternalLinkService">
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<mods:title id="79DABC0DEE087D4BCFC80DBEF2F5CB8F">Two new Oxytrechus species from Ecuadorian páramo (Coleoptera, Carabidae, Trechinae)</mods:title>
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<mods:namePart id="30317A8692771758259310CB91DD69A7">Quéinnec, Eric</mods:namePart>
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<mods:namePart id="06BB7F5FD12205B00AB9DC9E92DD2046">Ollivier, Eric</mods:namePart>
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<paragraph id="D63F3694FFFFFFCB3CA07EF2FDFEDB17" blockId="6.[151,560,1268,1331]" box="[151,560,1268,1295]" pageId="6" pageNumber="61">
<heading id="8D7781F8FFFFFFCB3CA07EF2FDFEDB17" bold="true" box="[151,560,1268,1295]" fontSize="11" level="1" pageId="6" pageNumber="61" reason="1">
<emphasis id="E4F4EA86FFFFFFCB3CA07EF2FDFEDB17" bold="true" box="[151,560,1268,1295]" pageId="6" pageNumber="61">
(b) 
<taxonomicName id="11804D17FFFFFFCB3CF77EF2FE25DB17" ID-CoL="8M636" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[192,491,1268,1295]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="cayambeensis" status="sp. nov.">
<emphasis id="E4F4EA86FFFFFFCB3CF77EF2FE25DB17" bold="true" box="[192,491,1268,1295]" italics="true" pageId="6" pageNumber="61">Oxytrechus cayambeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFC757FDFFFFFFCB3DC67EF3FDFEDB17" box="[497,560,1269,1295]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="9E9A651FFFFFFFCB3CA07F1FFEC2DB2B" box="[151,268,1304,1331]" pageId="6" pageNumber="61" type="description">
<paragraph id="D63F3694FFFFFFCB3CA07F1FFEC2DB2B" blockId="6.[151,560,1268,1331]" box="[151,268,1304,1331]" pageId="6" pageNumber="61">(figs. 4, 5)</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFFFFCB3CA07F67FDE9D837" pageId="6" pageNumber="61" type="materials_examined">
<paragraph id="D63F3694FFFFFFCB3CA07F67FA83DB63" blockId="6.[151,1437,1376,2015]" box="[151,1357,1376,1404]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CA07F67FEFEDB63" bold="true" box="[151,304,1376,1403]" pageId="6" pageNumber="61">
<typeStatus id="093B8836FFFFFFCB3CA07F67FF1CDB63" box="[151,210,1377,1403]" pageId="6" pageNumber="61">Type</typeStatus>
locality
</emphasis>
: 
<collectingCountry id="AE977604FFFFFFCB3D087F67FE50DB63" box="[319,414,1377,1403]" name="Ecuador" pageId="6" pageNumber="61">Ecuador</collectingCountry>
, 
<collectingMunicipality id="365BACEEFFFFFFCB3D9E7F67FD8FDB63" box="[425,577,1377,1403]" pageId="6" pageNumber="61">Southwestern</collectingMunicipality>
slope of the 
<location id="D35F604FFFFFFFCB3EE57F67FCF2DB63" LSID="urn:lsid:plazi:treatment:5E298782FFFFFFC43CA07EF2FEFDDADF:D35F604FFFFFFFCB3EE57F67FCF2DB63" box="[722,828,1377,1403]" country="Ecuador" municipality="Southwestern" name="Cayambe" pageId="6" pageNumber="61" stateProvince="Pichincha">Cayambe</location>
volcano (
<collectingRegion id="1444F876FFFFFFCB3F9D7F67FB47DB64" box="[938,1161,1377,1404]" country="Ecuador" name="Pichincha" pageId="6" pageNumber="61">Pichincha Province</collectingRegion>
), around 
<quantity id="11789B71FFFFFFCB38C67F67FA87DB63" box="[1265,1353,1377,1403]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="61" unit="m" value="4400.0">
<elevation id="5DADD1A7FFFFFFCB38C67F67FA87DB63" box="[1265,1353,1377,1403]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="61" unit="m" value="4400.0">4400 m</elevation>
</quantity>
.
</paragraph>
<paragraph id="D63F3694FFFFFFCB3CF07F83FDE9D837" blockId="6.[151,1437,1376,2015]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07F83FE9FDB87" bold="true" box="[199,337,1412,1439]" pageId="6" pageNumber="61">Type series</emphasis>
: 
<materialsCitation id="66E83CC9FFFFFFCB3D527F83FB70DBFF" ID-GBIF-Occurrence="2294309102" collectingDate="2018-07-22" collectionCode="CEQ" collectorName="E. Queinnec &amp; R. Chiori" country="Ecuador" elevation="4400" latitude="0.010277778" location="Cayambe volcano, south-west slope, Polylepis sp. grove" longLatPrecision="21" longitude="-78.024445" pageId="6" pageNumber="61" specimenCount="1" stateProvince="Pichincha Province" typeStatus="holotype">
<specimenCount id="C086FD1DFFFFFFCB3D527F83FE51DB87" box="[357,415,1413,1439]" pageId="6" pageNumber="61" type="male">Male</specimenCount>
<typeStatus id="093B8836FFFFFFCB3D9C7F83FDC3DB87" box="[427,525,1413,1439]" pageId="6" pageNumber="61" type="holotype">holotype</typeStatus>
labelled: „ 
<collectingCountry id="AE977604FFFFFFCB3EB87F83FD3FDB87" box="[655,753,1413,1439]" name="Ecuador" pageId="6" pageNumber="61">Ecuador</collectingCountry>
, 
<collectingRegion id="1444F876FFFFFFCB3F367F83FC2CDB87" box="[769,994,1413,1439]" country="Ecuador" name="Pichincha" pageId="6" pageNumber="61">Pichincha Province</collectingRegion>
, 
<location id="D35F604FFFFFFFCB3FC47F83FEA1DBDB" LSID="urn:lsid:plazi:treatment:5E298782FFFFFFC43CA07EF2FEFDDADF:D35F604FFFFFFFCB3FC47F83FEA1DBDB" country="Ecuador" latitude="0.010277778" longLatPrecision="21" longitude="-78.024445" name="Cayambe volcano, south-west slope, Polylepis sp. grove" pageId="6" pageNumber="61" stateProvince="Pichincha Province">
Cayambe volcano, south-west slope, 
<taxonomicName id="11804D17FFFFFFCB3CA07FAFFF31DBDB" box="[151,255,1449,1475]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="E4F4EA86FFFFFFCB3CA07FAFFF31DBDB" box="[151,255,1449,1475]" italics="true" pageId="6" pageNumber="61">Polylepis</emphasis>
</taxonomicName>
sp. grove
</location>
, alt. 
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<elevation id="5DADD1A7FFFFFFCB3D957FAFFE36DBDB" box="[418,504,1449,1475]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="61" unit="m" value="4400.0">4400 m</elevation>
</quantity>
, 
<geoCoordinate id="B3B45053FFFFFFCB3E347FAFFD4FDBDB" box="[515,641,1449,1475]" degrees="0" direction="north" minutes="00" orientation="latitude" pageId="6" pageNumber="61" precision="15" seconds="37" value="0.010277778">0°00’37’’N</geoCoordinate>
<geoCoordinate id="B3B45053FFFFFFCB3EBF7FAFFCD5DBDB" box="[648,795,1449,1475]" degrees="78" direction="west" minutes="01" orientation="longitude" pageId="6" pageNumber="61" precision="15" seconds="28" value="-78.024445">78°01’28’’W</geoCoordinate>
, 
<date id="A23E1054FFFFFFCB3F117FAFFC7CDBDB" box="[806,946,1449,1475]" pageId="6" pageNumber="61" value="2018-07-22">
<collectingDate id="B27AE9BCFFFFFFCB3F117FAFFC7CDBDB" box="[806,946,1449,1475]" pageId="6" pageNumber="61" value="2018-07-22">22.VII.2018</collectingDate>
</date>
, 
<collectorName id="7B755342FFFFFFCB3F8B7FAFFB8BDBDB" box="[956,1093,1449,1475]" pageId="6" pageNumber="61">E. Quéinnec</collectorName>
&amp; 
<collectorName id="7B755342FFFFFFCB385F7FAFFB1EDBDB" box="[1128,1232,1449,1475]" pageId="6" pageNumber="61">R. Chiori</collectorName>
leg.“ (white label, p)/„Holotype 
<taxonomicName id="11804D17FFFFFFCB3D057FCBFD9DDBFF" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[306,595,1485,1511]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="cayambeensis" status="sp. nov.">
<emphasis id="E4F4EA86FFFFFFCB3D057FCBFD9DDBFF" box="[306,595,1485,1511]" italics="true" pageId="6" pageNumber="61">Oxytrechus cayambeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFC757FDFFFFFFCB3E6D7FCBFD7FDBFF" box="[602,689,1485,1511]" pageId="6" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
Quéinnec &amp; Ollivier des.“ (red label, p), 
<collectionCode id="B091AE51FFFFFFCB38B57FCBFB70DBFF" box="[1154,1214,1485,1511]" pageId="6" pageNumber="61">CEQ</collectionCode>
</materialsCitation>
; 
<materialsCitation id="66E83CC9FFFFFFCB38FE7FCBFDEDD837" ID-GBIF-Occurrence="2294309103" collectingDate="2018-07-22" collectorName="E. Queinnec &amp; R. Chiori" country="Ecuador" elevation="4400" latitude="0.010277778" location="Cayambe" longLatPrecision="21" longitude="-78.024445" pageId="6" pageNumber="61" specimenCount="2" stateProvince="Pichincha" typeStatus="paratype">
<typeStatus id="093B8836FFFFFFCB38FE7FCBFAE1DBFF" box="[1225,1327,1485,1511]" pageId="6" pageNumber="61" type="paratype">Paratype</typeStatus>
: 1F, same data as holotype (white label, p) but labelled: „ 
<typeStatus id="093B8836FFFFFFCB3E837FF7FCD7D813" box="[692,793,1521,1547]" pageId="6" pageNumber="61" type="paratype">Paratype</typeStatus>
<taxonomicName id="11804D17FFFFFFCB3F147FF7FB88D813" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[803,1094,1521,1547]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="cayambeensis" status="sp. nov.">
<emphasis id="E4F4EA86FFFFFFCB3F147FF7FB88D813" box="[803,1094,1521,1547]" italics="true" pageId="6" pageNumber="61">Oxytrechus cayambeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFC757FDFFFFFFCB38787FF7FB66D813" box="[1103,1192,1521,1547]" pageId="6" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
Quéinnec &amp; Ollivier des.“ (red label, p), in coll. MNHN
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFFFFCB3CF07C3EFBBBD883" pageId="6" pageNumber="61" type="diagnosis">
<paragraph id="D63F3694FFFFFFCB3CF07C3EFBBBD883" blockId="6.[151,1437,1376,2015]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07C3EFEF4D84B" bold="true" box="[199,314,1592,1619]" pageId="6" pageNumber="61">Diagnosis</emphasis>
: An 
<taxonomicName id="11804D17FFFFFFCB3D437C3FFE3CD84B" authorityName="Jeannel" authorityYear="1927" box="[372,498,1593,1619]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="E4F4EA86FFFFFFCB3D437C3FFE3CD84B" box="[372,498,1593,1619]" italics="true" pageId="6" pageNumber="61">Oxytrechus</emphasis>
</taxonomicName>
of large size (LT = 
<quantity id="11789B71FFFFFFCB3EE57C3EFC93D84B" box="[722,861,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="6" pageNumber="61" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
; 
<quantity id="11789B71FFFFFFCB3F5E7C3EFC71D84B" box="[873,959,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="61" unit="mm" value="3.5">3.5 mm</quantity>
for male 
<typeStatus id="093B8836FFFFFFCB381A7C3FFB5AD84C" box="[1069,1172,1593,1620]" pageId="6" pageNumber="61" type="holotype">holotype</typeStatus>
). Body fairly elongated and broad, with relatively small head and long legs. Apterous. Head, pronotum and elytra shiny blackish brown; elytra slightly iridescent. Clypeus, labrum, palpi, antenna and legs bright reddish brown.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFFFFC43CF07CA3FAADDC7B" lastPageId="9" lastPageNumber="64" pageId="6" pageNumber="61" type="description">
<paragraph id="D63F3694FFFFFFCB3CF07CA3FADFD9C7" blockId="6.[151,1437,1376,2015]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07CA3FEC9D8A7" bold="true" box="[199,263,1701,1727]" pageId="6" pageNumber="61">Head</emphasis>
small, subquadrate (HW/HL = 0.67/0.67 = 
<quantity id="11789B71FFFFFFCB3EC37CA3FCD1D8A7" box="[756,799,1701,1727]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="61" unit="in" value="1.0">1 in</quantity>
male 
<typeStatus id="093B8836FFFFFFCB3F527CA3FC09D8A7" box="[869,967,1701,1727]" pageId="6" pageNumber="61" type="holotype">holotype</typeStatus>
and female 
<typeStatus id="093B8836FFFFFFCB38657CA3FB7AD8A7" box="[1106,1204,1701,1727]" pageId="6" pageNumber="61" type="paratype">paratype</typeStatus>
) and moderately depressed above, with deep entire frontal furrows which are sub-angulate at middle and moderately divergent in front; both frons and supraorbital areas gently convex, the latter with a foveola on each side at the root of supraorbital seta; microsculpture distinct, consisting of coarse polygonal meshes which are mostly isodiametric on frons and supraorbital areas but transverse on vertex; eyes small but gently protruding though convex; genae glabrous, curved, about as long as eyes; neck wide; mentum gently emarginate at apex; mandibles long and slender; palpi short though fairly slender; antennae fairly long (LA = 
<quantity id="11789B71FFFFFFCB3E157D7BFD4BD98F" box="[546,645,1917,1943]" metricMagnitude="-3" metricUnit="m" metricValue="1.93" pageId="6" pageNumber="61" unit="mm" value="1.93">1.93 mm</quantity>
in both sex) for a direct relative of 
<taxonomicName id="11804D17FFFFFFCB38307D7BFB42D98F" authorityName="Mateu" authorityYear="1991" box="[1031,1164,1917,1943]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="E4F4EA86FFFFFFCB38307D7BFB42D98F" box="[1031,1164,1917,1943]" italics="true" pageId="6" pageNumber="61">O. globosus</emphasis>
</taxonomicName>
, reaching or extending a little beyond basal one-sixth of elytra, with segment two slightly longer than segments 3 and 4 each, as long segment 5 and only shorter than segment 6; antennal segments 7–10 each about three-sevenths as wide as long.
</paragraph>
<caption id="82FF661CFFFEFFCA3CA07CBCFEC0D8EF" httpUri="https://zenodo.org/record/3346518/files/figure.png" pageId="7" pageNumber="62" startId="7.[151,250,1722,1747]" targetBox="[438,1156,221,1650]" targetPageId="7">
<paragraph id="D63F3694FFFEFFCA3CA07CBCFEC0D8EF" blockId="7.[151,1436,1722,1783]" pageId="7" pageNumber="62">
<emphasis id="E4F4EA86FFFEFFCA3CA07CBCFEDDD8CB" bold="true" box="[151,275,1722,1747]" pageId="7" pageNumber="62">FIGURE 4.</emphasis>
Habitus of 
<taxonomicName id="11804D17FFFEFFCA3DBC7CBDFD5CD8CB" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[395,658,1723,1747]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cayambeensis" status="sp. nov.">
<emphasis id="E4F4EA86FFFEFFCA3DBC7CBDFD5CD8CB" box="[395,658,1723,1747]" italics="true" pageId="7" pageNumber="62">Oxytrechus cayambeensis</emphasis>
</taxonomicName>
<emphasis id="E4F4EA86FFFEFFCA3EAF7CBDFD1CD8CB" bold="true" box="[664,722,1723,1747]" pageId="7" pageNumber="62">
<taxonomicNameLabel id="FFC757FDFFFEFFCA3EAF7CBDFD1CD8CB" box="[664,722,1723,1747]" pageId="7" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from the western slopes of the Cayambe volcano, Ecuadorian páramo (Holotype).
</paragraph>
</caption>
<paragraph id="D63F3694FFFEFFC53CF07D23FDE6DF07" blockId="7.[151,1437,1828,2036]" lastBlockId="8.[151,1437,153,683]" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62">
<emphasis id="E4F4EA86FFFEFFCA3CF07D23FE8FD927" bold="true" box="[199,321,1829,1855]" pageId="7" pageNumber="62">Prothorax</emphasis>
. Pronotum transverse (PW/PL = 0.92/0.67 = 
<quantity id="11789B71FFFEFFCA3F797D23FC50D927" box="[846,926,1829,1855]" metricMagnitude="-2" metricUnit="m" metricValue="3.4798" pageId="7" pageNumber="62" unit="in" value="1.37">1.37 in</quantity>
male 
<typeStatus id="093B8836FFFEFFCA3FD07D23FB83D927" box="[999,1101,1829,1855]" pageId="7" pageNumber="62" type="holotype">holotype</typeStatus>
; 
<quantity id="11789B71FFFEFFCA386D7D22FB65D927" box="[1114,1195,1828,1855]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" pageId="7" pageNumber="62" unit="in" value="1.35">1.35 in</quantity>
female) and convex, wider at about three-fifths from base and more strongly contracted posteriad than anteriad; microsculpture composed of very fine transverse lines partially forming very transverse meshes; sides sharply bordered and reflexed, particularly behind middle, moderately rounded in front, much so behind but not sinuate before hind angles, which are scarcely marked; anterior marginal seta situated before the widest part, the posterior one close to hind angle; apex either straight or slightly emarginate, with front angle slightly advanced though narrowly rounded; base nar- rower than apex, straight at middle but oblique on each side inside hind angle; median line sharply impressed, somewhat widening near base; apical transverse impression usually continuous though shallow at the median part and continuing on each side to near basal border; basal fovea not large and rather mal-defined; basal area uneven with marked longitudinal striations.
</paragraph>
<paragraph id="D63F3694FFF1FFC53CF07B2FFC0BDCB3" blockId="8.[151,1437,153,683]" pageId="8" pageNumber="63">
<emphasis id="E4F4EA86FFF1FFC53CF07B2FFE90DF5B" bold="true" box="[199,350,297,323]" pageId="8" pageNumber="63">Pterothorax.</emphasis>
Elytra ovate (EW/EL = 1.67/2.21 = 
<quantity id="11789B71FFF1FFC53F337B2EFC9ADF5B" box="[772,852,296,323]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="8" pageNumber="63" unit="in" value="0.75">0.75 in</quantity>
male 
<typeStatus id="093B8836FFF1FFC53FAB7B2FFBCCDF5B" box="[924,1026,297,323]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
; 
<quantity id="11789B71FFF1FFC538397B2FFB90DF5B" box="[1038,1118,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="1.9558" pageId="8" pageNumber="63" unit="in" value="0.77">0.77 in</quantity>
female), widest about fourninths from base, well convex at the sides and in apical part but more or less depressed on the disk; the ratio of the width of the elytra to that of pronotum 1.79–1.81 (mean 1.80); microsculpture sharply impressed, consisting of polygonal meshes which are usually wide; shoulders distinct though rounded; prehumeral borders fairly long and moderately curved; sides rather widely reflexed, moderately arcuate from shoulders to apices which are rounded, each with a slight preapical emargination; striae completely obsolete, even in the basal part and apical one; sutural stria scarcely evident. Scutellar striole slightly reduced but discernable, very short, nearly straight; apical carina hardly visible; two setiferous dorsal pores, fovea-like, on the putative site of stria 3, the anterior one at the 3/10–3/8 from base, the posterior one just on the middle; humeral group of umbilicate series with the four pores equidistant, first pore of the middle group behind the level of the second discal pore; preapical pore close to the apex, at the level of the 8th pore of the umbilicate series; internal apical pore not detected.
</paragraph>
<caption id="82FF661CFFF1FFC53CA07DA8FDC6D9F3" httpUri="https://zenodo.org/record/3346520/files/figure.png" pageId="8" pageNumber="63" startId="8.[151,264,1966,1991]" targetBox="[334,1260,729,1918]" targetPageId="8">
<paragraph id="D63F3694FFF1FFC53CA07DA8FDC6D9F3" blockId="8.[151,1436,1966,2027]" pageId="8" pageNumber="63">
<emphasis id="E4F4EA86FFF1FFC53CA07DA8FE96D9DF" bold="true" box="[151,344,1966,1991]" pageId="8" pageNumber="63">FIGURES 5 A–C.</emphasis>
Aedeagus of the holotype of 
<taxonomicName id="11804D17FFF1FFC53EB47DA9FC47D9DF" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[643,905,1967,1991]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="cayambeensis" status="sp. nov.">
<emphasis id="E4F4EA86FFF1FFC53EB47DA9FC47D9DF" box="[643,905,1967,1991]" italics="true" pageId="8" pageNumber="63">Oxytrechus cayambeensis</emphasis>
</taxonomicName>
<emphasis id="E4F4EA86FFF1FFC53FB87DA9FC06D9DF" bold="true" box="[911,968,1967,1991]" pageId="8" pageNumber="63">
<taxonomicNameLabel id="FFC757FDFFF1FFC53FB87DA9FC06D9DF" box="[911,968,1967,1991]" pageId="8" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
—Left lateral view (
<emphasis id="E4F4EA86FFF1FFC538AF7DA8FB64D9DF" bold="true" box="[1176,1194,1966,1991]" pageId="8" pageNumber="63">A</emphasis>
);—Ventral view (B);— Dorsal view (C); Scale bar: 300 µm.
</paragraph>
</caption>
<paragraph id="D63F3694FFF0FFC43CF07A9FFB1ADF07" blockId="9.[151,1437,153,1223]" pageId="9" pageNumber="64">
<emphasis id="E4F4EA86FFF0FFC43CF07A9FFF31DEAB" bold="true" box="[199,255,153,179]" pageId="9" pageNumber="64">Legs</emphasis>
relatively long though fairly stout; protibiae straight, moderately dilated towards apices, scarcely pubescent on the anterior face, each with a groove on the external face and also with a very shallow groove along the inner side of the external one; tarsi fairly slender, tarsal segment 4 with a long ventral apophysis in pro- and mesotarsi; two proximal segment of each protarsus widely dilated in male, and well produced inwards at apices.
</paragraph>
<paragraph id="D63F3694FFF0FFC43CF07B2EFB61DF7F" blockId="9.[151,1437,153,1223]" pageId="9" pageNumber="64">
<emphasis id="E4F4EA86FFF0FFC43CF07B2EFEFDDF5B" bold="true" box="[199,307,296,323]" pageId="9" pageNumber="64">Ventrites</emphasis>
. Ventral surface smooth, but covered with microscopic pubescence perceptible only under high magnification; anal sternite provided with a pair of sexual setae in male, with two pairs in female.
</paragraph>
<paragraph id="D63F3694FFF0FFC43CF07B76FAADDC7B" blockId="9.[151,1437,153,1223]" pageId="9" pageNumber="64">
<emphasis id="E4F4EA86FFF0FFC43CF07B76FEF4DF93" bold="true" box="[199,314,368,395]" pageId="9" pageNumber="64">Aedeagus</emphasis>
(fig. 5). Male genital organ fairly large, elongate but slightly sclerotized. Aedeagus fairly slender and hardly arcuate behind middle, with rather large basal part, which is straight, heavily sclerotized ventral side, and then bent towards the dorsal side; lateral sides of basal orifice semi-circularly emarginated; sagittal aileron very large, elongate and roundly dilated at apex; viewed laterally, apical part regularly attenuated, slightly produced and turn up at the extremity, which is rounded; viewed ventrally (fig. 5B), median lobe subparallel, apical part slightly blunt, with conical extremity; viewed dorsally (fig. 5C) apical part seems to be slightly widening and then attenuated towards apex. Inner sac armed with an elongated than the right, each provided four, rather short setae at apex.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFF0FFC43CF0786BFAB1DC9F" box="[199,1407,621,647]" pageId="9" pageNumber="64" type="etymology">
<paragraph id="D63F3694FFF0FFC43CF0786BFAB1DC9F" blockId="9.[151,1437,153,1223]" box="[199,1407,621,647]" pageId="9" pageNumber="64">
<emphasis id="E4F4EA86FFF0FFC43CF0786BFE59DC9F" bold="true" box="[199,407,621,647]" italics="true" pageId="9" pageNumber="64">Derivatio nominis</emphasis>
: The specific epithet refers to the 
<typeStatus id="093B8836FFF0FFC43F21786BFC88DC9F" box="[790,838,621,647]" pageId="9" pageNumber="64">type</typeStatus>
locality of the new species, the Cayambe volcano.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFF0FFC43CF07896FAD4DCB3" box="[199,1306,656,683]" pageId="9" pageNumber="64" type="distribution">
<paragraph id="D63F3694FFF0FFC43CF07896FAD4DCB3" blockId="9.[151,1437,153,1223]" box="[199,1306,656,683]" pageId="9" pageNumber="64">
<emphasis id="E4F4EA86FFF0FFC43CF07896FE97DCB3" bold="true" box="[199,345,656,683]" pageId="9" pageNumber="64">Distribution</emphasis>
: Localized on the Eastern Cordillera on the Southwestern slope of Cayambe volcano.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFF0FFC43CF078B2FEFDDADF" pageId="9" pageNumber="64" type="diagnosis">
<paragraph id="D63F3694FFF0FFC43CF078B2FEFDDADF" blockId="9.[151,1437,153,1223]" pageId="9" pageNumber="64">
<emphasis id="E4F4EA86FFF0FFC43CF078B2FE02DCD7" bold="true" box="[199,460,692,719]" pageId="9" pageNumber="64">Diagnostic characters</emphasis>
: The 
<taxonomicName id="11804D17FFF0FFC43E2578B3FCFBDCD7" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[530,821,693,719]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="cayambeensis" status="sp. nov.">
<emphasis id="E4F4EA86FFF0FFC43E2578B3FCFBDCD7" box="[530,821,693,719]" italics="true" pageId="9" pageNumber="64">Oxytrechus cayambeensis</emphasis>
</taxonomicName>
<emphasis id="E4F4EA86FFF0FFC43F0978B3FCB1DCD7" bold="true" box="[830,895,693,719]" pageId="9" pageNumber="64">
<taxonomicNameLabel id="FFC757FDFFF0FFC43F0978B3FCB1DCD7" box="[830,895,693,719]" pageId="9" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is morphologically distinct from 
<taxonomicName id="11804D17FFF0FFC4393478B3FE95DCEB" authority="Allegro et al., 2008" authorityName="Allegro" authorityYear="2008" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="onorei">
<emphasis id="E4F4EA86FFF0FFC4393478B3FAA1DCD7" box="[1283,1391,693,719]" italics="true" pageId="9" pageNumber="64">O. onorei</emphasis>
<bibRefCitation id="B2114B65FFF0FFC4394078B3FE95DCEB" author="Allegro, G. &amp; Giachino, P. M. &amp; Sciaky, R." pageId="9" pageNumber="64" pagination="131 - 171" refId="ref6494" refString="Allegro, G., Giachino, P. M. &amp; Sciaky, R. (2008) Notes on the Trechinae (Coleoptera Carabidae) of South America with description of new species from Chile, Ecuador and Peru. In: Giachino, P. M. (Ed.), Biodiversity of South America. Memoirs on Biodiversity, I, pp. 131 - 171." type="journal article" year="2008">
Allegro 
<emphasis id="E4F4EA86FFF0FFC43CEE78DCFEC2DCEB" box="[217,268,729,755]" italics="true" pageId="9" pageNumber="64">et al</emphasis>
., 2008
</bibRefCitation>
</taxonomicName>
with whom it lives in sympatry. It differentiates itself from this species by its much larger size (
<quantity id="11789B71FFF0FFC43CA878FAFEE7DD0F" box="[159,297,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="9" pageNumber="64" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
<emphasis id="E4F4EA86FFF0FFC43D0778F8FEB6DD0F" box="[304,376,766,791]" italics="true" pageId="9" pageNumber="64">versus</emphasis>
<quantity id="11789B71FFF0FFC43DB778FBFDEADD0F" box="[384,548,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="3.01" metricValueMax="3.14" metricValueMin="2.88" pageId="9" pageNumber="64" unit="mm" value="3.01" valueMax="3.14" valueMin="2.88">2.88–3.14 mm</quantity>
for 
<taxonomicName id="11804D17FFF0FFC43E6278F8FD53DD0F" box="[597,669,766,791]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="onorei">
<emphasis id="E4F4EA86FFF0FFC43E6278F8FD53DD0F" box="[597,669,766,791]" italics="true" pageId="9" pageNumber="64">onorei</emphasis>
</taxonomicName>
), its very different morphology with wider elytra, in a regular oval, and by its conformation to the aedeagus. 
<taxonomicName id="11804D17FFF0FFC43E5D7927FCE7DD23" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[618,809,801,827]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="cayambeensis">
<emphasis id="E4F4EA86FFF0FFC43E5D7927FCE7DD23" box="[618,809,801,827]" italics="true" pageId="9" pageNumber="64">O. cayambeensis</emphasis>
</taxonomicName>
is also divergent from 
<taxonomicName id="11804D17FFF0FFC438187927FB04DD23" authorityName="Jeannel" authorityYear="1927" box="[1071,1226,801,827]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="lallemandi">
<emphasis id="E4F4EA86FFF0FFC438187927FB04DD23" box="[1071,1226,801,827]" italics="true" pageId="9" pageNumber="64">O. lallemandi</emphasis>
</taxonomicName>
shares similarities with 
<taxonomicName id="11804D17FFF0FFC43CF87943FEF6DD47" box="[207,312,837,863]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="onorei">
<emphasis id="E4F4EA86FFF0FFC43CF87943FEF6DD47" box="[207,312,837,863]" italics="true" pageId="9" pageNumber="64">O. onorei</emphasis>
</taxonomicName>
. 
<taxonomicName id="11804D17FFF0FFC43D727943FDCDDD47" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[325,515,837,863]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="cayambeensis">
<emphasis id="E4F4EA86FFF0FFC43D727943FDCDDD47" box="[325,515,837,863]" italics="true" pageId="9" pageNumber="64">O. cayambeensis</emphasis>
</taxonomicName>
species is also close to 
<taxonomicName id="11804D17FFF0FFC43F317943FC45DD47" authorityName="Mateu" authorityYear="1991" box="[774,907,837,863]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="E4F4EA86FFF0FFC43F317943FC45DD47" box="[774,907,837,863]" italics="true" pageId="9" pageNumber="64">O. globosus</emphasis>
</taxonomicName>
by its general conformation, its size and its elytral shape. Nevertheless, 
<emphasis id="E4F4EA86FFF0FFC43D86796FFDB6DD9B" box="[433,632,873,899]" italics="true" pageId="9" pageNumber="64">O. cayambeensis’</emphasis>
head is smaller than 
<taxonomicName id="11804D17FFF0FFC43F50796FFC05DD9B" authorityName="Mateu" authorityYear="1991" box="[871,971,873,899]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="E4F4EA86FFF0FFC43F50796FFC05DD9B" box="[871,971,873,899]" italics="true" pageId="9" pageNumber="64">globosus</emphasis>
</taxonomicName>
’, the eyes are smaller and not prominent, the pronotum is more transverse, with a less convex surface; its sides present a maximal width at the 3/5 
<superScript id="21F59BDCFFF0FFC4392C798AFAE6DD82" attach="left" box="[1307,1320,908,922]" fontSize="6" pageId="9" pageNumber="64">th</superScript>
from base (by the middle for 
<taxonomicName id="11804D17FFF0FFC43D5379B7FE24DDD3" authorityName="Mateu" authorityYear="1991" box="[356,490,945,971]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="E4F4EA86FFF0FFC43D5379B7FE24DDD3" box="[356,490,945,971]" italics="true" pageId="9" pageNumber="64">O. globosus</emphasis>
</taxonomicName>
). The elytral striation is far more pronounced for 
<taxonomicName id="11804D17FFF0FFC4383B79B7FB5CDDD3" authorityName="Mateu" authorityYear="1991" box="[1036,1170,945,971]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="E4F4EA86FFF0FFC4383B79B7FB5CDDD3" box="[1036,1170,945,971]" italics="true" pageId="9" pageNumber="64">O. globosus</emphasis>
</taxonomicName>
in which the location of the three first striations can be distinguished (the elytral surface is smooth for 
<taxonomicName id="11804D17FFF0FFC43FCE79D3FB79DDF7" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[1017,1207,981,1007]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="cayambeensis">
<emphasis id="E4F4EA86FFF0FFC43FCE79D3FB79DDF7" box="[1017,1207,981,1007]" italics="true" pageId="9" pageNumber="64">O. cayambeensis</emphasis>
</taxonomicName>
). On the other hand, the aedeagus of the two species is of substantially different general shape (
<emphasis id="E4F4EA86FFF0FFC43FE579FFFC20DA0B" box="[978,1006,1017,1043]" italics="true" pageId="9" pageNumber="64">cf.</emphasis>
<bibRefCitation id="B2114B65FFF0FFC43FC379FFFB4EDA0B" author="Moret, P." box="[1012,1152,1016,1043]" pageId="9" pageNumber="64" refId="ref7201" refString="Moret, P. (2005) Los coleopteros Carabidae del paramo en los Andes del Ecuador. Sistematica, ecologia y biogeografia. Quito, Pontificia Universidad Catolica del Ecuador, Centro de Biodiversidad y Ambiente, Monografia 2. Gruppo Editoriale il Capitello, Torino, 306 pp." type="book" year="2005">Moret, 2005</bibRefCitation>
, fig. 91, for 
<taxonomicName id="11804D17FFF0FFC4393079FFFA42DA0B" authorityName="Mateu" authorityYear="1991" box="[1287,1420,1017,1043]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="E4F4EA86FFF0FFC4393079FFFA42DA0B" box="[1287,1420,1017,1043]" italics="true" pageId="9" pageNumber="64">O. globosus</emphasis>
</taxonomicName>
). What is quite surprising is that the aedeagus of 
<taxonomicName id="11804D17FFF0FFC43E837E1BFCBADA2F" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[692,884,1053,1079]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="cayambeensis">
<emphasis id="E4F4EA86FFF0FFC43E837E1BFCBADA2F" box="[692,884,1053,1079]" italics="true" pageId="9" pageNumber="64">O. cayambeensis</emphasis>
</taxonomicName>
presents some analogies with those of 
<taxonomicName id="11804D17FFF0FFC439067E1BFA52DA2F" authorityName="Mateu" authorityYear="1988" box="[1329,1436,1053,1079]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="moreti">
<emphasis id="E4F4EA86FFF0FFC439067E1BFA52DA2F" box="[1329,1436,1053,1079]" italics="true" pageId="9" pageNumber="64">O. moreti</emphasis>
</taxonomicName>
and 
<taxonomicName id="11804D17FFF0FFC43CF07E47FEEEDA43" authorityName="Casale &amp; Sciaky" authorityYear="1986" box="[199,288,1089,1115]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="zoiai">
<emphasis id="E4F4EA86FFF0FFC43CF07E47FEEEDA43" box="[199,288,1089,1115]" italics="true" pageId="9" pageNumber="64">O. zoiai</emphasis>
</taxonomicName>
but is far more robust: the sagittal blade is wider and stretched out; the median lobe progressively diminishes around the apex. The habitus of 
<taxonomicName id="11804D17FFF0FFC43E5C7E63FCE7DA67" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[619,809,1125,1151]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="cayambeensis">
<emphasis id="E4F4EA86FFF0FFC43E5C7E63FCE7DA67" box="[619,809,1125,1151]" italics="true" pageId="9" pageNumber="64">O. cayambeensis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="11804D17FFF0FFC43F8A7E63FBDBDA67" authorityName="Casale &amp; Sciaky" authorityYear="1986" box="[957,1045,1125,1151]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="zoiai">
<emphasis id="E4F4EA86FFF0FFC43F8A7E63FBDBDA67" box="[957,1045,1125,1151]" italics="true" pageId="9" pageNumber="64">O. zoiai</emphasis>
</taxonomicName>
and 
<taxonomicName id="11804D17FFF0FFC4387B7E63FB78DA67" authorityName="Mateu" authorityYear="1988" box="[1100,1206,1125,1151]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="moreti">
<emphasis id="E4F4EA86FFF0FFC4387B7E63FB78DA67" box="[1100,1206,1125,1151]" italics="true" pageId="9" pageNumber="64">O. moreti</emphasis>
</taxonomicName>
by its imposing size (
<quantity id="11789B71FFF0FFC43CA87E8EFEE6DABB" box="[159,296,1160,1187]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="9" pageNumber="64" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
<emphasis id="E4F4EA86FFF0FFC43D197E8CFEB8DABB" box="[302,374,1162,1187]" italics="true" pageId="9" pageNumber="64">versus</emphasis>
2.9–3.0 mm for 
<taxonomicName id="11804D17FFF0FFC43E047E8CFDB5DABB" authorityName="Mateu" authorityYear="1988" box="[563,635,1162,1187]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="moreti">
<emphasis id="E4F4EA86FFF0FFC43E047E8CFDB5DABB" box="[563,635,1162,1187]" italics="true" pageId="9" pageNumber="64">moreti</emphasis>
</taxonomicName>
and 3.0– 
<quantity id="11789B71FFF0FFC43ED37E8FFCF7DABB" box="[740,825,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="64" unit="mm" value="3.3">3.3 mm</quantity>
for 
<taxonomicName id="11804D17FFF0FFC43F5F7E8CFC50DABB" authorityName="Casale &amp; Sciaky" authorityYear="1986" box="[872,926,1162,1187]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="zoiai">
<emphasis id="E4F4EA86FFF0FFC43F5F7E8CFC50DABB" box="[872,926,1162,1187]" italics="true" pageId="9" pageNumber="64">zoiai</emphasis>
</taxonomicName>
) and by its quite different, broader, and more globose form.
</paragraph>
</subSubSection>
</treatment>
</document>