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<document id="7145471A3A48FF5D9F185EAAECDE1B1A" ID-CLB-Dataset="26146" ID-DOI="10.11646/zootaxa.4646.1.3" ID-GBIF-Dataset="de20b639-5800-47da-9214-99a33a9c13cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3346510" ID-ZooBank="EA93232D-5C20-48FC-BA26-E5DC811CFCC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563868669035" checkinUser="plazi" docAuthor="Quéinnec, Eric &amp; Ollivier, Eric" docDate="2019" docId="5E298782FFFAFFCB3CA078FBFA4DDABB" docLanguage="en" docName="zootaxa.4646.1.3.pdf.imf" docOrigin="Zootaxa 4646 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxytrechus chioriae Quéinnec &amp; Ollivier 2019, n. sp." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="A210FFFAFFF9FFCD3C377A06FFCEDE18" masterDocTitle="Two new Oxytrechus species from Ecuadorian páramo (Coleoptera, Carabidae, Trechinae)" masterLastPageNumber="66" masterPageNumber="55" pageNumber="58" updateTime="1698747107410" updateUser="ExternalLinkService">
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<mods:title id="A8109CA779B403B8231911416C9A1422">Two new Oxytrechus species from Ecuadorian páramo (Coleoptera, Carabidae, Trechinae)</mods:title>
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<mods:namePart id="1AD73DCA0EABAA1FC565B1F7B0BACA77">Quéinnec, Eric</mods:namePart>
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<mods:namePart id="8E5FF004DD5AB6C82252F11F0DA2F5C0">Ollivier, Eric</mods:namePart>
</mods:name>
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<mods:title id="0AE65E18802F6381F87C8F5D034F20C6">Zootaxa</mods:title>
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<mods:date id="6A02AE6D65686D90E0CE26063152DD96">2019</mods:date>
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<mods:number id="B7BC01E28F2F45E49B2D77DE715A3F4F">2019-07-23</mods:number>
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<mods:detail id="CEAE07904D2A6C08AD5C44C8D79437FC" type="volume">
<mods:number id="A3CA194CAF2C3DE40D926AA9C549903B">4646</mods:number>
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<mods:classification id="B290D2E1CCD7AE63B02009408DA35CF2">journal article</mods:classification>
<mods:identifier id="0D9D1C3FA009B7AE2A5C5F86D10DCD92" type="CLB-Dataset">26146</mods:identifier>
<mods:identifier id="AF0FEEC502DB4B959003D132DD394453" type="DOI">10.11646/zootaxa.4646.1.3</mods:identifier>
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<mods:identifier id="3777FAE6CF8FB428A1F430FA4EAFBF11" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5E298782FFFAFFCB3CA078FBFA4DDABB" ID-DOI="http://doi.org/10.5281/zenodo.5943169" ID-GBIF-Taxon="158849688" ID-Zenodo-Dep="5943169" LSID="urn:lsid:plazi:treatment:5E298782FFFAFFCB3CA078FBFA4DDABB" httpUri="http://treatment.plazi.org/id/5E298782FFFAFFCB3CA078FBFA4DDABB" lastPageId="6" lastPageNumber="61" pageId="3" pageNumber="58">
<subSubSection id="9E9A651FFFFAFFCE3CA078FBFDCEDD0F" box="[151,512,765,792]" pageId="3" pageNumber="58" type="nomenclature">
<paragraph id="D63F3694FFFAFFCE3CA078FBFDCEDD0F" blockId="3.[151,512,765,827]" box="[151,512,765,792]" pageId="3" pageNumber="58">
<heading id="8D7781F8FFFAFFCE3CA078FBFDCEDD0F" bold="true" box="[151,512,765,792]" fontSize="11" level="1" pageId="3" pageNumber="58" reason="1">
<emphasis id="E4F4EA86FFFAFFCE3CA078FBFDCEDD0F" bold="true" box="[151,512,765,792]" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CA078FBFE78DD00" bold="true" box="[151,438,765,792]" italics="true" pageId="3" pageNumber="58">
(a)
<taxonomicName id="11804D17FFFAFFCE3CF678FBFE78DD00" ID-CoL="8MHY9" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[193,438,765,792]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="chioriae" status="sp. nov.">Oxytrechus chioriae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="FFC757FDFFFAFFCE3D8A78FBFDCEDD0F" box="[445,512,765,791]" pageId="3" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFAFFCE3CA07927FEDCDD23" box="[151,274,801,827]" pageId="3" pageNumber="58" type="description">
<paragraph id="D63F3694FFFAFFCE3CA07927FEDCDD23" blockId="3.[151,512,765,827]" box="[151,274,801,827]" pageId="3" pageNumber="58">
(
<figureCitation id="4EBB2A11FFFAFFCE3CA87927FF21DD23" box="[159,239,801,827]" captionStart="FIGURE 2" captionStartId="4.[151,250,1862,1887]" captionTargetBox="[192,1041,287,1783]" captionTargetId="figure@4.[264,1323,306,1834]" captionTargetPageId="4" captionText="FIGURE 2. Habitus of Oxytrechus chioriae n. sp., from the east side of the Chimborazo volcano, Ecuadorian páramo (Holotype)." httpUri="https://zenodo.org/record/3346514/files/figure.png" pageId="3" pageNumber="58">Figs. 2</figureCitation>
,
<figureCitation id="4EBB2A11FFFAFFCE3CCC7927FEC7DD23" box="[251,265,801,827]" captionStart="FIGURES 3" captionStartId="5.[151,264,1654,1679]" captionTargetBox="[378,1217,238,1577]" captionTargetId="figure@5.[347,1241,225,1585]" captionTargetPageId="5" captionText="FIGURES 3 A–C. Aedeagus of the holotype of Oxytrechus chioriae n. sp.—Left lateral view (A); —Ventral view (B); —Dorsal view (C); Scale bar: 300 µm." httpUri="https://zenodo.org/record/3346516/files/figure.png" pageId="3" pageNumber="58">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFAFFCE3CA0796FFCECDA2F" pageId="3" pageNumber="58" type="materials_examined">
<paragraph id="D63F3694FFFAFFCE3CA0796FFAD3DD9B" blockId="3.[151,1437,872,2015]" box="[151,1309,872,900]" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CA0796FFEFEDD9B" bold="true" box="[151,304,872,899]" pageId="3" pageNumber="58">
<typeStatus id="093B8836FFFAFFCE3CA0796FFF1CDD9B" box="[151,210,873,899]" pageId="3" pageNumber="58">Type</typeStatus>
locality
</emphasis>
:
<collectingCountry id="AE977604FFFAFFCE3D08796FFE50DD9B" box="[319,414,873,899]" name="Ecuador" pageId="3" pageNumber="58">Ecuador</collectingCountry>
,
<collectingRegion id="1444F876FFFAFFCE3D9E796FFDFDDD9B" box="[425,563,873,899]" country="Ecuador" name="Chimborazo" pageId="3" pageNumber="58">Chimborazo</collectingRegion>
volcano (
<collectingRegion id="1444F876FFFAFFCE3E96796FFC53DD9C" box="[673,925,873,900]" country="Ecuador" name="Chimborazo" pageId="3" pageNumber="58">Chimborazo Province</collectingRegion>
),
<location id="D35F604FFFFAFFCE3F87796FFBE5DD9B" LSID="urn:lsid:plazi:treatment:5E298782FFFAFFCB3CA078FBFA4DDABB:D35F604FFFFAFFCE3F87796FFBE5DD9B" box="[944,1067,873,899]" country="Ecuador" name="North-East" pageId="3" pageNumber="58" stateProvince="Chimborazo">North-East</location>
side, around
<quantity id="11789B71FFFAFFCE38F6796FFAD7DD9B" box="[1217,1305,873,899]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="58" unit="m" value="4400.0">
<elevation id="5DADD1A7FFFAFFCE38F6796FFAD7DD9B" box="[1217,1305,873,899]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="58" unit="m" value="4400.0">4400 m</elevation>
</quantity>
.
</paragraph>
<paragraph id="D63F3694FFFAFFCE3CF0798BFCECDA2F" blockId="3.[151,1437,872,2015]" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CF0798BFE85DDBF" bold="true" box="[199,331,908,935]" pageId="3" pageNumber="58">Type series</emphasis>
:
<materialsCitation id="66E83CC9FFFAFFCE3D6E798BFA52DDF7" ID-GBIF-Occurrence="2294309104" collectingDate="2018-07-28" collectionCode="CEQ" collectorName="E. Queinnec &amp; R. Chiori" country="Ecuador" determinerName="Queinnec &amp; Ollivier" latitude="-1.4602778" location="Chimborazo volcano, north-eastern slope, Portal Andino, Polylepis sp. grove" longLatPrecision="21" longitude="-78.770836" pageId="3" pageNumber="58" specimenCount="4401" specimenCount-male="4401" stateProvince="Chimborazo Province" typeStatus="holotype">
<specimenCount id="C086FD1DFFFAFFCE3D6E798BFE5DDDBF" box="[345,403,909,935]" pageId="3" pageNumber="58" type="male">Male</specimenCount>
<typeStatus id="093B8836FFFAFFCE3DAE798BFE35DDBF" box="[409,507,909,935]" pageId="3" pageNumber="58" type="holotype">holotype</typeStatus>
labelled: „
<collectingCountry id="AE977604FFFAFFCE3E45798BFD1ADDBF" box="[626,724,909,935]" name="Ecuador" pageId="3" pageNumber="58">Ecuador</collectingCountry>
,
<collectingRegion id="1444F876FFFAFFCE3EE8798BFC18DDBF" box="[735,982,909,935]" country="Ecuador" name="Chimborazo" pageId="3" pageNumber="58">Chimborazo Province</collectingRegion>
,
<location id="D35F604FFFFAFFCE3FD6798BFDF1DDD3" LSID="urn:lsid:plazi:treatment:5E298782FFFAFFCB3CA078FBFA4DDABB:D35F604FFFFAFFCE3FD6798BFDF1DDD3" country="Ecuador" latitude="-1.4602778" longLatPrecision="21" longitude="-78.770836" name="Chimborazo volcano, north-eastern slope, Portal Andino, Polylepis sp. grove" pageId="3" pageNumber="58" stateProvince="Chimborazo Province">
Chimborazo volcano, north-eastern slope, Portal Andino,
<taxonomicName id="11804D17FFFAFFCE3D5F79B7FE1EDDD3" box="[360,464,945,971]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="E4F4EA86FFFAFFCE3D5F79B7FE1EDDD3" box="[360,464,945,971]" italics="true" pageId="3" pageNumber="58">Polylepis</emphasis>
</taxonomicName>
sp. grove
</location>
, alt.
<specimenCount id="C086FD1DFFFAFFCE3E4579B7FD07DDD3" box="[626,713,945,971]" pageId="3" pageNumber="58" type="male">
<quantity id="11789B71FFFAFFCE3E4579B7FD07DDD3" box="[626,713,945,971]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="58" unit="m" value="4400.0">4400 m</quantity>
</specimenCount>
,
<geoCoordinate id="B3B45053FFFAFFCE3EE379B7FC81DDD3" box="[724,847,945,971]" degrees="1" direction="south" minutes="27" orientation="latitude" pageId="3" pageNumber="58" precision="15" seconds="37" value="-1.4602778">1°27’37’’S</geoCoordinate>
<geoCoordinate id="B3B45053FFFAFFCE3F6279B6FC26DDD3" box="[853,1000,944,971]" degrees="78" direction="west" minutes="46" orientation="longitude" pageId="3" pageNumber="58" precision="15" seconds="15" value="-78.770836">78°46’15’’W</geoCoordinate>
,
<date id="A23E1054FFFAFFCE3FC479B7FBB0DDD3" box="[1011,1150,945,971]" pageId="3" pageNumber="58" value="2018-07-28">
<collectingDate id="B27AE9BCFFFAFFCE3FC479B7FBB0DDD3" box="[1011,1150,945,971]" pageId="3" pageNumber="58" value="2018-07-28">28.VII.2018</collectingDate>
</date>
,
<collectorName id="7B755342FFFAFFCE38BE79B7FADCDDD3" box="[1161,1298,945,971]" pageId="3" pageNumber="58">E. Quéinnec</collectorName>
&amp;
<collectorName id="7B755342FFFAFFCE390279B7FA53DDD3" box="[1333,1437,945,971]" pageId="3" pageNumber="58">R. Chiori</collectorName>
leg.“ (white label, p) / „Holotype
<taxonomicName id="11804D17FFFAFFCE3E2D79D3FD30DDF7" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[538,766,981,1007]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="chioriae" status="sp. nov.">
<emphasis id="E4F4EA86FFFAFFCE3E2D79D3FD30DDF7" box="[538,766,981,1007]" italics="true" pageId="3" pageNumber="58">Oxytrechus chioriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFC757FDFFFAFFCE3F3079D3FC91DDF7" box="[775,863,981,1007]" pageId="3" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="23E3DA50FFFAFFCE3F5079D3FC1CDDF7" box="[871,978,981,1007]" pageId="3" pageNumber="58">Quéinnec</determinerName>
&amp;
<determinerName id="23E3DA50FFFAFFCE3FCD79D3FB9FDDF7" box="[1018,1105,981,1007]" pageId="3" pageNumber="58">Ollivier</determinerName>
des.“ (red label, p), in
<collectionCode id="B091AE51FFFAFFCE396B79D3FA56DDF7" box="[1372,1432,981,1007]" pageId="3" pageNumber="58">CEQ</collectionCode>
;
</materialsCitation>
<materialsCitation id="66E83CC9FFFAFFCE3CA079FFFCD0DA2F" ID-GBIF-Occurrence="2294309105" collectingDate="2018-07-28" collectorName="E. Queinnec &amp; R. Chiori" country="Ecuador" elevation="4400" latitude="-1.4602778" location="Portal Andino" longLatPrecision="21" longitude="-78.770836" pageId="3" pageNumber="58" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chimborazo" typeStatus="paratype">
<typeStatus id="093B8836FFFAFFCE3CA079FFFEC6DA0B" box="[151,264,1017,1043]" pageId="3" pageNumber="58" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C086FD1DFFFAFFCE3D2379FFFEA9DA0B" box="[276,359,1017,1043]" pageId="3" pageNumber="58" type="male">1 Male</specimenCount>
,
<specimenCount id="C086FD1DFFFAFFCE3D4379FFFE2EDA0B" box="[372,480,1017,1043]" pageId="3" pageNumber="58" type="female">1 Female</specimenCount>
, same data as for the holotype, but labelled: „
<typeStatus id="093B8836FFFAFFCE3FDD79FFFB81DA0B" box="[1002,1103,1017,1043]" pageId="3" pageNumber="58" type="paratype">Paratype</typeStatus>
<taxonomicName id="11804D17FFFAFFCE386F79FFFAF2DA0B" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[1112,1340,1017,1043]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="chioriae" status="sp. nov.">
<emphasis id="E4F4EA86FFFAFFCE386F79FFFAF2DA0B" box="[1112,1340,1017,1043]" italics="true" pageId="3" pageNumber="58">Oxytrechus chioriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FFC757FDFFFAFFCE397279FFFA53DA0B" box="[1349,1437,1017,1043]" pageId="3" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
Quéinnec &amp; Ollivier des. “ (red label, p) in MNHN, CEO
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFAFFCE3CF07E46FC19DB4F" pageId="3" pageNumber="58" type="diagnosis">
<paragraph id="D63F3694FFFAFFCE3CF07E46FC19DB4F" blockId="3.[151,1437,872,2015]" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CF07E46FEF4DA43" bold="true" box="[199,314,1088,1115]" pageId="3" pageNumber="58">Diagnosis</emphasis>
. Externally similar to
<taxonomicName id="11804D17FFFAFFCE3E027E47FBE7DA43" authority="Allegro, Giachino &amp; Sciaky, 2008" authorityName="Allegro, Giachino &amp; Sciaky" authorityYear="2008" box="[565,1065,1089,1115]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="mateui">
<emphasis id="E4F4EA86FFFAFFCE3E027E47FD6ADA43" box="[565,676,1089,1115]" italics="true" pageId="3" pageNumber="58">O. mateui</emphasis>
<bibRefCitation id="B2114B65FFFAFFCE3E9C7E47FBE7DA43" author="Allegro, G. &amp; Giachino, P. M. &amp; Sciaky, R." box="[683,1065,1089,1115]" pageId="3" pageNumber="58" pagination="131 - 171" refId="ref6494" refString="Allegro, G., Giachino, P. M. &amp; Sciaky, R. (2008) Notes on the Trechinae (Coleoptera Carabidae) of South America with description of new species from Chile, Ecuador and Peru. In: Giachino, P. M. (Ed.), Biodiversity of South America. Memoirs on Biodiversity, I, pp. 131 - 171." type="journal article" year="2008">Allegro, Giachino &amp; Sciaky, 2008</bibRefCitation>
</taxonomicName>
from Atacazo volcano due to the general shape of the body but easily recognized at first sight on its smaller size: 3.0–
<quantity id="11789B71FFFAFFCE38657E63FB68DA67" box="[1106,1190,1125,1151]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="58" unit="mm" value="3.1">3.1 mm</quantity>
(instead of
<quantity id="11789B71FFFAFFCE391B7E62FF0DDABB" metricMagnitude="-3" metricUnit="m" metricValue="3.41" metricValueMax="3.51" metricValueMin="3.31" pageId="3" pageNumber="58" unit="mm" value="3.41" valueMax="3.51" valueMin="3.31">3.31–3.51 mm</quantity>
in
<taxonomicName id="11804D17FFFAFFCE3CDF7E8FFE99DABB" authorityName="Allegro, Giachino &amp; Sciaky" authorityYear="2008" box="[232,343,1161,1187]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="mateui">
<emphasis id="E4F4EA86FFFAFFCE3CDF7E8FFE99DABB" box="[232,343,1161,1187]" italics="true" pageId="3" pageNumber="58">O. mateui</emphasis>
</taxonomicName>
). However, it differs markedly by the shape of the aedeagal median lobe and the structure of the endophallus. Also, very similar to
<taxonomicName id="11804D17FFFAFFCE3E157EABFC92DADF" authority="Mateu, 1991" authorityName="Mateu" authorityYear="1991" box="[546,860,1197,1223]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="guaguanus">
<emphasis id="E4F4EA86FFFAFFCE3E157EABFD0CDADF" box="[546,706,1197,1223]" italics="true" pageId="3" pageNumber="58">O. guaguanus</emphasis>
<bibRefCitation id="B2114B65FFFAFFCE3EFC7EABFC92DADF" author="Mateu, J." box="[715,860,1197,1223]" pageId="3" pageNumber="58" pagination="71 - 83" refId="ref7053" refString="Mateu, J. (1991) Sur le genre Oxytrechus Jeannel, 1927, avec la description de nouvelles especes de l'Equateur et de la Colombie (Coleoptera, Carabidae, Trechinae). Eos, 67, 71 - 83." type="journal article" year="1991">Mateu, 1991</bibRefCitation>
</taxonomicName>
, from Guagua
<collectingRegion id="1444F876FFFAFFCE383D7EABFBB6DADF" box="[1034,1144,1197,1223]" country="Ecuador" name="Pichincha" pageId="3" pageNumber="58">Pichincha</collectingRegion>
volcano, by the aedeagal shape but, in lateral view, the median lobe is more convex in the middle, less elongated and thick, more narrowed towards apex; with a helicoidal-shaped endophallic copulatory piece and with an extensive sclerotinization of its anterior portion. Though similar to each other at first sight, the three species differ in many details of external morphology and besides, quite different in conformation of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFAFFCB3CF07F66FE9DDFEF" lastPageId="6" lastPageNumber="61" pageId="3" pageNumber="58" type="description">
<paragraph id="D63F3694FFFAFFCE3CF07F66FEC8DBDB" blockId="3.[151,1437,872,2015]" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CF07F66FE97DB63" bold="true" box="[199,345,1376,1403]" pageId="3" pageNumber="58">Description.</emphasis>
A medium-sized
<taxonomicName id="11804D17FFFAFFCE3E1F7F67FD68DB63" authorityName="Jeannel" authorityYear="1927" box="[552,678,1377,1403]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="E4F4EA86FFFAFFCE3E1F7F67FD68DB63" box="[552,678,1377,1403]" italics="true" pageId="3" pageNumber="58">Oxytrechus</emphasis>
</taxonomicName>
species (LT = 3.0–
<quantity id="11789B71FFFAFFCE3FBB7F67FC26DB63" box="[908,1000,1377,1403]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="58" unit="mm" value="3.1">3.1 mm</quantity>
). Head, pronotum and elytra blackish brown, markedly shiny; elytra slightly iridescent. Clypeus, labrum, palpi, antenna and legs bright reddish brown. Apterous.
</paragraph>
<paragraph id="D63F3694FFFAFFCE3CF07FCBFB45D91F" blockId="3.[151,1437,872,2015]" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CF07FCBFEC9DBFF" bold="true" box="[199,263,1485,1511]" pageId="3" pageNumber="58">Head</emphasis>
subquadrate (HW/HL = 0.55/0.55 =
<quantity id="11789B71FFFAFFCE3E827FCBFD2DDBFF" box="[693,739,1485,1511]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="58" unit="in" value="1.0">1 in</quantity>
male
<typeStatus id="093B8836FFFAFFCE3F1A7FCBFC5CDBFF" box="[813,914,1485,1511]" pageId="3" pageNumber="58" type="holotype">holotype</typeStatus>
, and in average), large, transverse, with deep frontal furrows not angulate at the middle and widely divergent posteriad; frons and supraorbital areas moderately convex, bearing a foveola at the root of each supraorbital seta; two pairs of supraorbital setae lying on lines diverging backwards; eyes small but fairly protruding, more or less longer than genae, which are gently convex and completely glabrous; neck very wide, with constriction distinct; labrum transverse, with the apical margin shallowly but widely emarginate; mandible short, stout, feebly arcuate in apical third and obtusely tridentate; antennae short and fairly stout, reaching basal third of the elytra, scape thick, shorter but broader than terminal segment, segments 2 and 4–10 subequal in length, each subcylindrical and fully twice as long as large, segment 3 slightly longer than the neighboring ones and about five-sevenths as long as the terminal, which is the longest.
</paragraph>
<paragraph id="D63F3694FFFAFFC93CF07D17FC10DF07" blockId="3.[151,1437,872,2015]" lastBlockId="4.[151,1436,153,287]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
<emphasis id="E4F4EA86FFFAFFCE3CF07D17FE8FD933" bold="true" box="[199,321,1809,1835]" pageId="3" pageNumber="58">Prothorax</emphasis>
. Pronotum cordate, wider than head, strongly transverse (PW/PL = 0.75/0.55 =
<quantity id="11789B71FFFAFFCE38FB7D17FAD5D933" box="[1228,1307,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.4544" pageId="3" pageNumber="58" unit="in" value="1.36">1.36 in</quantity>
male
<typeStatus id="093B8836FFFAFFCE39567D17FF05D957" pageId="3" pageNumber="58" type="holotype">holotype</typeStatus>
;
<quantity id="11789B71FFFAFFCE3CE17D32FEEDD957" box="[214,291,1844,1871]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" pageId="3" pageNumber="58" unit="in" value="1.35">1.35 in</quantity>
average), widest at about five-sevenths from base, and more gradually but strongly contracted towards base than towards apex, with the sides narrowly but entirely bordered; sides strongly rounded near the widest part, rather gently so behind middle and briefly sinuate just a little before basal angles; front angles not projecting forwards; base strongly arcuate, with hind angles very obtuse, apex nearly straight, with front angles obtuse; base much larger than apex; dorsum well convex, with sharply impressed median line reaching base and usually with a small round depression on each side at the level of the widest part; apical transverse impression well distinct, usually uneven; basal impressions wide, laterally joining basal fovea, which are small and narrow; basal area sharply striated; two pairs of marginal setae on each side, of which the anterior one lies just before the widest part and the posterior one not on hind angles but slightly advanced in front of them.
</paragraph>
<caption id="82FF661CFFFDFFC93CA07D40FF1CD99A" httpUri="https://zenodo.org/record/3346514/files/figure.png" pageId="4" pageNumber="59" startId="4.[151,250,1862,1887]" targetBox="[192,1041,287,1783]" targetPageId="4">
<paragraph id="D63F3694FFFDFFC93CA07D40FF1CD99A" blockId="4.[151,1437,1862,1922]" pageId="4" pageNumber="59">
<emphasis id="E4F4EA86FFFDFFC93CA07D40FEDAD947" bold="true" box="[151,276,1862,1887]" pageId="4" pageNumber="59">FIGURE 2.</emphasis>
Habitus of
<taxonomicName id="11804D17FFFDFFC93DBA7D40FD95D946" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[397,603,1862,1886]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="chioriae" status="sp. nov.">
<emphasis id="E4F4EA86FFFDFFC93DBA7D40FD95D946" box="[397,603,1862,1886]" italics="true" pageId="4" pageNumber="59">Oxytrechus chioriae</emphasis>
</taxonomicName>
<emphasis id="E4F4EA86FFFDFFC93E557D40FD52D946" bold="true" box="[610,668,1862,1886]" pageId="4" pageNumber="59">
<taxonomicNameLabel id="FFC757FDFFFDFFC93E557D40FD52D946" box="[610,668,1862,1886]" pageId="4" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, from the east side of the Chimborazo volcano, Ecuadorian páramo (Holotype).
</paragraph>
</caption>
<caption id="82FF661CFFFCFFC83CA07C70FE2FD8AB" httpUri="https://zenodo.org/record/3346516/files/figure.png" pageId="5" pageNumber="60" startId="5.[151,264,1654,1679]" targetBox="[378,1217,238,1577]" targetPageId="5">
<paragraph id="D63F3694FFFCFFC83CA07C70FE2FD8AB" blockId="5.[151,1436,1654,1715]" pageId="5" pageNumber="60">
<emphasis id="E4F4EA86FFFCFFC83CA07C70FE96D897" bold="true" box="[151,344,1654,1679]" pageId="5" pageNumber="60">FIGURES 3 A–C.</emphasis>
Aedeagus of the holotype of
<taxonomicName id="11804D17FFFCFFC83EB67C71FC83D897" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[641,845,1655,1679]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="chioriae" status="sp. nov.">
<emphasis id="E4F4EA86FFFCFFC83EB67C71FC83D897" box="[641,845,1655,1679]" italics="true" pageId="5" pageNumber="60">Oxytrechus chioriae</emphasis>
</taxonomicName>
<emphasis id="E4F4EA86FFFCFFC83F637C71FC43D897" bold="true" box="[852,909,1655,1679]" pageId="5" pageNumber="60">
<taxonomicNameLabel id="FFC757FDFFFCFFC83F637C71FC43D897" box="[852,909,1655,1679]" pageId="5" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
—Left lateral view (
<emphasis id="E4F4EA86FFFCFFC8386B7C70FBA0D897" bold="true" box="[1116,1134,1654,1679]" pageId="5" pageNumber="60">A</emphasis>
); —Ventral view (B); —Dorsal view (C); Scale bar: 300 µm.
</paragraph>
</caption>
<paragraph id="D63F3694FFFCFFCB3CF07CE7FE4ADECF" blockId="5.[151,1437,1760,2040]" lastBlockId="6.[151,1437,153,1187]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">
<emphasis id="E4F4EA86FFFCFFC83CF07CE7FE90D8E3" bold="true" box="[199,350,1761,1787]" pageId="5" pageNumber="60">Pterothorax.</emphasis>
Elytra oval, oblong (EW/EL = 1.25/1.60 =
<quantity id="11789B71FFFCFFC83F097CE7FC45D8E3" box="[830,907,1761,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.9812" pageId="5" pageNumber="60" unit="in" value="0.78">0.78 in</quantity>
male
<typeStatus id="093B8836FFFCFFC83FFA7CE7FBFDD8E3" box="[973,1075,1761,1787]" pageId="5" pageNumber="60" type="holotype">holotype</typeStatus>
;
<quantity id="11789B71FFFCFFC8380A7CE7FBB2D8E3" box="[1085,1148,1761,1787]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="60" unit="in" value="0.8">0.8 in</quantity>
average), widest at about four-ninths from bases, and equally narrowed toward bases and toward apices, dorsum strongly convex; shoulders rounded, thinly distinct, sides entirely bordered, rather strongly arcuate in basal third, less so in median parts, conjointly rounded at apices, each margin forming a distinct terminal point at the apical part; striae vestigial excepted stria 1 which is always sharply impressed throughout, striae 2–3 not traceable on the disc though very shallow, stria 8 impressed only behind the middle set of marginal umbilicate pores; scutellar striole well defined, bordered; apical striole short but distinct, divergent anteriorly, and free at anterior end; apical carina straight; two setiferous dorsal pores, usually foveolate, present on the site of stria 3 at 1/6 and about 2/3 from base respectively; preapical pore present and located at the 1/10 from apex; humeral set of marginal umbilicate pores nearly regular though not perfectly equidistant.
</paragraph>
<paragraph id="D63F3694FFFFFFCB3CF07AE7FD1EDF07" blockId="6.[151,1437,153,1187]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07AE7FF31DEE3" bold="true" box="[199,255,225,251]" pageId="6" pageNumber="61">Legs</emphasis>
short but robust; protibial furrow superficial. Meso- and metatibias short and straight. First two male protarsomeres distinctly and asymmetrically widened.
</paragraph>
<paragraph id="D63F3694FFFFFFCB3CF07B2EFE9DDFEF" blockId="6.[151,1437,153,1187]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07B2EFEF4DF5B" bold="true" box="[199,314,296,323]" pageId="6" pageNumber="61">Aedeagus</emphasis>
(fig. 3) robust, elongate. Median lobe, in lateral view, sharply curved at base, sublinear and regular along the central part, tapering from apical third toward apex; apex long, regularly thinned dorso-ventrally, sharply thinned in ventral or dorsal view, hook shaped at apical extremity. Basal bulb very long and enlarged, sagittal carina thin and long, extended to the dorsal part. Inner sac provided with a long, sclerified, twisted copulatory piece which is progressively enlarged at apex. Styles long, reaching the apical third of the median lobe, each provided with four long distal setae.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFFFFCB3CF07807FBA5DC27" pageId="6" pageNumber="61" type="etymology">
<paragraph id="D63F3694FFFFFFCB3CF07807FBA5DC27" blockId="6.[151,1437,153,1187]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07807FE51DC03" box="[199,415,513,539]" italics="true" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF07807FE56DC03" bold="true" box="[199,408,513,539]" italics="true" pageId="6" pageNumber="61">Derivatio nominis</emphasis>
.
</emphasis>
It is a real pleasure to dedicate this new species to one of its collectors, our friend Dr. R. Chiori for her longstanding support to entomological researches in numerous countries.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFFFFCB3CA0784EFCE6DCB3" pageId="6" pageNumber="61" type="distribution">
<paragraph id="D63F3694FFFFFFCB3CA0784EFCE6DCB3" blockId="6.[151,1437,153,1187]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CA0784EFEE7DC7B" bold="true" box="[151,297,584,611]" pageId="6" pageNumber="61">Distribution</emphasis>
. Known only from the northeastern slope of the
<collectingRegion id="1444F876FFFFFFCB3F60784FFC2FDC7B" box="[855,993,585,611]" country="Ecuador" name="Chimborazo" pageId="6" pageNumber="61">Chimborazo</collectingRegion>
volcano (Western Cordillera). Located within patches of
<taxonomicName id="11804D17FFFFFFCB3D56786BFE07DC9F" box="[353,457,621,647]" class="Diplopoda" family="Platyrhacidae" genus="Polylepis" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="E4F4EA86FFFFFFCB3D56786BFE07DC9F" box="[353,457,621,647]" italics="true" pageId="6" pageNumber="61">Polylepis</emphasis>
</taxonomicName>
sp. forest, residual at elevations between 4300 and
<quantity id="11789B71FFFFFFCB3824786AFBA6DC9F" box="[1043,1128,620,647]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="61" unit="m" value="4500.0">4500 m</quantity>
along the ecotone between the grazed páramo and the lower part of the super-páramo.
</paragraph>
</subSubSection>
<subSubSection id="9E9A651FFFFFFFCB3CF078B2FA4DDABB" pageId="6" pageNumber="61" type="diagnosis">
<paragraph id="D63F3694FFFFFFCB3CF078B2FA4DDABB" blockId="6.[151,1437,153,1187]" pageId="6" pageNumber="61">
<emphasis id="E4F4EA86FFFFFFCB3CF078B2FE05DCD7" bold="true" box="[199,459,692,719]" pageId="6" pageNumber="61">Diagnostic characters</emphasis>
: The general shape of the habitus relates this species to
<taxonomicName id="11804D17FFFFFFCB387E78B3FA52DCD7" authority="Allegro et al., 2008" authorityName="Allegro" authorityYear="2008" box="[1097,1436,693,719]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="mateui">
<emphasis id="E4F4EA86FFFFFFCB387E78B3FB76DCD7" box="[1097,1208,693,719]" italics="true" pageId="6" pageNumber="61">O. mateui</emphasis>
<bibRefCitation id="B2114B65FFFFFFCB388878B3FA52DCD7" author="Allegro, G. &amp; Giachino, P. M. &amp; Sciaky, R." box="[1215,1436,693,719]" pageId="6" pageNumber="61" pagination="131 - 171" refId="ref6494" refString="Allegro, G., Giachino, P. M. &amp; Sciaky, R. (2008) Notes on the Trechinae (Coleoptera Carabidae) of South America with description of new species from Chile, Ecuador and Peru. In: Giachino, P. M. (Ed.), Biodiversity of South America. Memoirs on Biodiversity, I, pp. 131 - 171." type="journal article" year="2008">
Allegro
<emphasis id="E4F4EA86FFFFFFCB392B78B0FA98DCD7" box="[1308,1366,693,719]" italics="true" pageId="6" pageNumber="61">et al.</emphasis>
, 2008
</bibRefCitation>
</taxonomicName>
even though the latter have a far larger size (
<quantity id="11789B71FFFFFFCB3EB478DEFCE5DCEB" box="[643,811,728,755]" metricMagnitude="-3" metricUnit="m" metricValue="3.41" metricValueMax="3.51" metricValueMin="3.31" pageId="6" pageNumber="61" unit="mm" value="3.41" valueMax="3.51" valueMin="3.31">3.31–3.51 mm</quantity>
). Specimens of
<taxonomicName id="11804D17FFFFFFCB3FEC78DFFB97DCEB" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[987,1113,729,755]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae">
<emphasis id="E4F4EA86FFFFFFCB3FEC78DFFB97DCEB" box="[987,1113,729,755]" italics="true" pageId="6" pageNumber="61">O. chioriae</emphasis>
</taxonomicName>
studied were not longer than
<quantity id="11789B71FFFFFFCB3CA078FBFF3EDD0F" box="[151,240,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="61" unit="mm" value="3.1">3.1 mm</quantity>
. Nevertheless, the aedeagus of the two species are very different. Aedeagus of
<taxonomicName id="11804D17FFFFFFCB385678FBFB2EDD0F" authority="Quéinnec &amp; Ollivier, 2019" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[1121,1248,765,791]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae" status="sp. nov.">
<emphasis id="E4F4EA86FFFFFFCB385678FBFB2EDD0F" box="[1121,1248,765,791]" italics="true" pageId="6" pageNumber="61">O. chioriae</emphasis>
</taxonomicName>
<emphasis id="E4F4EA86FFFFFFCB38D078FBFAE8DD0F" bold="true" box="[1255,1318,765,791]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="FFC757FDFFFFFFCB38D078FBFAE8DD0F" box="[1255,1318,765,791]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
presents a general conformation close to
<taxonomicName id="11804D17FFFFFFCB3DD87927FD40DD23" authorityName="Mateu" authorityYear="1991" box="[495,654,801,827]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guaguanus">
<emphasis id="E4F4EA86FFFFFFCB3DD87927FD40DD23" box="[495,654,801,827]" italics="true" pageId="6" pageNumber="61">O. guaguanus</emphasis>
</taxonomicName>
: the median lobe is stretched out, the tip progressively slender, regularly reducing to the apex and ending in a hook. However, the aedeagus’s apex presents a sickle extremity clearly not as developed as in
<taxonomicName id="11804D17FFFFFFCB3DAE796CFDDADD9B" authorityName="Mateu" authorityYear="1991" box="[409,532,874,899]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guaguanus">
<emphasis id="E4F4EA86FFFFFFCB3DAE796CFDDADD9B" box="[409,532,874,899]" italics="true" pageId="6" pageNumber="61">guaguanus</emphasis>
</taxonomicName>
. Nevertheless,
<taxonomicName id="11804D17FFFFFFCB3EF5796FFCD1DD9B" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[706,799,873,899]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae">
<emphasis id="E4F4EA86FFFFFFCB3EF5796FFCD1DD9B" box="[706,799,873,899]" italics="true" pageId="6" pageNumber="61">chioriae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="11804D17FFFFFFCB3F82796CFBFEDD9B" authorityName="Mateu" authorityYear="1991" box="[949,1072,874,899]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guaguanus">
<emphasis id="E4F4EA86FFFFFFCB3F82796CFBFEDD9B" box="[949,1072,874,899]" italics="true" pageId="6" pageNumber="61">guaguanus</emphasis>
</taxonomicName>
with a larger median lobe and a less ventrally straightened apex; the sagittal aileron is far less developed and is proportionally very small in
<taxonomicName id="11804D17FFFFFFCB3954798BFF0ADDD3" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae">
<emphasis id="E4F4EA86FFFFFFCB3954798BFF0ADDD3" italics="true" pageId="6" pageNumber="61">chioriae</emphasis>
</taxonomicName>
. Concerning the habitus,
<taxonomicName id="11804D17FFFFFFCB3DD779B7FD91DDD3" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[480,607,945,971]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae">
<emphasis id="E4F4EA86FFFFFFCB3DD779B7FD91DDD3" box="[480,607,945,971]" italics="true" pageId="6" pageNumber="61">O. chioriae</emphasis>
</taxonomicName>
’s elytra are more oblong, more stretched out than for
<taxonomicName id="11804D17FFFFFFCB388579B7FA81DDD3" authorityName="Mateu" authorityYear="1991" box="[1202,1359,945,971]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guaguanus">
<emphasis id="E4F4EA86FFFFFFCB388579B7FA81DDD3" box="[1202,1359,945,971]" italics="true" pageId="6" pageNumber="61">O. guaguanus</emphasis>
</taxonomicName>
whose general size is markedly smaller (
<quantity id="11789B71FFFFFFCB3E3879D3FD55DDF7" box="[527,667,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="6" pageNumber="61" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
). It is likely that
<taxonomicName id="11804D17FFFFFFCB3F6F79D3FC19DDF7" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[856,983,981,1007]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae">
<emphasis id="E4F4EA86FFFFFFCB3F6F79D3FC19DDF7" box="[856,983,981,1007]" italics="true" pageId="6" pageNumber="61">O. chioriae</emphasis>
</taxonomicName>
constitutes, with different species of the Occidental Cordillera,
<emphasis id="E4F4EA86FFFFFFCB3DAE79FCFD57DA0B" box="[409,665,1018,1043]" italics="true" pageId="6" pageNumber="61">
i.e.
<taxonomicName id="11804D17FFFFFFCB3DF479FCFDDCDA0B" box="[451,530,1018,1043]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="mateui">mateui</taxonomicName>
,
<taxonomicName id="11804D17FFFFFFCB3E2979FCFD57DA0B" authorityName="Mateu" authorityYear="1991" box="[542,665,1018,1043]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="guaguanus">guaguanus</taxonomicName>
</emphasis>
and most likely
<taxonomicName id="11804D17FFFFFFCB3F6F79FCFC2DDA0B" box="[856,995,1018,1043]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="pierremoreti">
<emphasis id="E4F4EA86FFFFFFCB3F6F79FCFC2DDA0B" box="[856,995,1018,1043]" italics="true" pageId="6" pageNumber="61">pierremoreti</emphasis>
</taxonomicName>
(even though the aedeagus representation of this species in
<bibRefCitation id="B2114B65FFFFFFCB3DBA7E1BFDA7DA2F" author="Allegro, G. &amp; Giachino, P. M. &amp; Sciaky, R." box="[397,617,1053,1079]" pageId="6" pageNumber="61" pagination="131 - 171" refId="ref6494" refString="Allegro, G., Giachino, P. M. &amp; Sciaky, R. (2008) Notes on the Trechinae (Coleoptera Carabidae) of South America with description of new species from Chile, Ecuador and Peru. In: Giachino, P. M. (Ed.), Biodiversity of South America. Memoirs on Biodiversity, I, pp. 131 - 171." type="journal article" year="2008">
Allegro
<emphasis id="E4F4EA86FFFFFFCB3DDD7E18FDD5DA2F" box="[490,539,1053,1079]" italics="true" pageId="6" pageNumber="61">et al</emphasis>
., 2008
</bibRefCitation>
probably implies a damaged median lobe), a related-group of species by the aedeagus’s conformation and also by the shape of the habitus. Aedeagus of
<taxonomicName id="11804D17FFFFFFCB383C7E47FB44DA43" authorityName="Quéinnec &amp; Ollivier" authorityYear="2019" box="[1035,1162,1089,1115]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="chioriae">
<emphasis id="E4F4EA86FFFFFFCB383C7E47FB44DA43" box="[1035,1162,1089,1115]" italics="true" pageId="6" pageNumber="61">O. chioriae</emphasis>
</taxonomicName>
share, as well, a median lobe conformation analogous to that of
<taxonomicName id="11804D17FFFFFFCB3E627E63FD0EDA67" authorityName="Mateu" authorityYear="1988" box="[597,704,1125,1151]" class="Insecta" family="Carabidae" genus="Oxytrechus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="moreti">
<emphasis id="E4F4EA86FFFFFFCB3E627E63FD0EDA67" box="[597,704,1125,1151]" italics="true" pageId="6" pageNumber="61">O. moreti</emphasis>
</taxonomicName>
with an apical blade longer and slimmer at the apex. The base is widened, robust, and offers a thin sagittal blade but quite long which ventrally reduces at the level of the ostium.
</paragraph>
</subSubSection>
</treatment>
</document>
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