177 lines
17 KiB
XML
177 lines
17 KiB
XML
<document id="0B8DD1D8BD1A86720BC79115A71F7AAA" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FFAC4A67FF328F73FDA0E880" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole machaquila Longino 2019, new species" docType="treatment" docVersion="22" lastPageNumber="45" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="45" updateTime="1698726215157" updateUser="ExternalLinkService">
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<mods:mods id="CA4070AC8E433FFDA319DA211ED79EBF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="05F73FDDFDA222C1DD387227614E9477">
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<mods:title id="F59132D32C0081FBF2018CAEC6501908">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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<mods:name id="26362E22F9D734934C707FCF96D2581D" type="personal">
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<mods:role id="480D6670054CEBA459F414F8265A410D">
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<mods:roleTerm id="D5FFB91AC098E582C21A66E874335C52">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="95CD7E9826182BD62C58901FD18742DA">Longino, John T.</mods:namePart>
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<mods:typeOfResource id="C7CBBDAD70B6DA4397B2F637F9F44363">text</mods:typeOfResource>
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<mods:titleInfo id="9B092E5F03FFB8221DFF2887EB47AC2D">
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<mods:title id="58998A9494F569DDDF0256C993D1D331">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6E31C9048A650FF0B0F88C0B81967848">
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<mods:date id="3697E083B0D0930F09B2488EB46D16DB">2019</mods:date>
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<mods:detail id="0B1BFFF27E3FB998454AB3BF63F1D30F" type="pubDate">
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<mods:number id="FC07286FE2F417AF437DD53D550CB6B5">2019-05-06</mods:number>
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<mods:detail id="55E76BD95E41C72097AF14D489F44C3F" type="volume">
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<mods:number id="0F924ACCC0BDC102F6D1E799066F4A77">4599</mods:number>
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</mods:detail>
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<mods:detail id="5198A655E3D7B880B1CA2DDDAD6DE7F9" type="issue">
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<mods:number id="B6E936BFEAAEC2C6FBB639E4BEF12D99">1</mods:number>
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<mods:start id="520DBA60CB8F8AE9589B13325B84C1F3">1</mods:start>
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<mods:classification id="C902F024E223607C6BA835C2DE67F18E">journal article</mods:classification>
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<mods:identifier id="B7D26A108CCCFF7EDC85093B17DDF054" type="CLB-Dataset">26920</mods:identifier>
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<mods:identifier id="817D4CC585751F38A09C241F3BE68861" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
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<mods:identifier id="237A83E37C9DEC92EC476B1B75E8C741" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8D72B2C67FE4AE2D3DBD874C07014469" type="Zenodo-Dep">2668058</mods:identifier>
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<mods:identifier id="98D512B2DABAB9F96DC844EA8070656D" type="ZooBank">7CDD24FF-5BA3-4D06-898D-A3E220515D4C</mods:identifier>
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</mods:mods>
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<treatment id="5B56E872FFAC4A67FF328F73FDA0E880" ID-DOI="http://doi.org/10.5281/zenodo.5586214" ID-GBIF-Taxon="157062626" ID-Zenodo-Dep="5586214" LSID="urn:lsid:plazi:treatment:5B56E872FFAC4A67FF328F73FDA0E880" httpUri="http://treatment.plazi.org/id/5B56E872FFAC4A67FF328F73FDA0E880" lastPageNumber="45" pageId="44" pageNumber="45">
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<subSubSection id="9BE50AEFFFAC4A67FF328F73FEF1EB0C" pageId="44" pageNumber="45" type="nomenclature">
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<paragraph id="D3405964FFAC4A67FF328F73FDDAEB6D" blockId="44.[151,558,1052,1111]" box="[151,558,1052,1078]" pageId="44" pageNumber="45">
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<heading id="8808EE08FFAC4A67FF328F73FDDAEB6D" bold="true" box="[151,558,1052,1078]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
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<emphasis id="E18B8576FFAC4A67FF328F73FDDAEB6D" bold="true" box="[151,558,1052,1078]" pageId="44" pageNumber="45">
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<taxonomicName id="14FF22E7FFAC4A67FF328F73FE63EB6D" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,407,1052,1078]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="machaquila" status="sp. nov.">
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<emphasis id="E18B8576FFAC4A67FF328F73FE63EB6D" bold="true" box="[151,407,1052,1078]" italics="true" pageId="44" pageNumber="45">Pheidole machaquila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FAB8380DFFAC4A67FE3B8F73FDDAEB6D" box="[414,558,1052,1078]" pageId="44" pageNumber="45" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D3405964FFAC4A67FF328F51FEF1EB0C" blockId="44.[151,558,1052,1111]" box="[151,261,1086,1111]" pageId="44" pageNumber="45">
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(
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<figureCitation id="4BC445E1FFAC4A67FF3B8F51FF08EB0C" box="[158,252,1086,1111]" captionStart="PLATE 14" captionStartId="80.[151,238,1954,1977]" captionTargetId="figure@80.[202,1384,194,1933]" captionTargetPageId="80" captionText="PLATE 14." httpUri="https://zenodo.org/record/2668257/files/figure.png" pageId="44" pageNumber="45">Plate 14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF328FE9FC6FEBC4" box="[151,923,1158,1183]" pageId="44" pageNumber="45" type="description">
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<paragraph id="D3405964FFAC4A67FF328FE9FC6FEBC4" blockId="44.[151,1437,1158,2011]" box="[151,923,1158,1183]" pageId="44" pageNumber="45">
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<taxonomicName id="14FF22E7FFAC4A67FF328FE9FF0FEBC4" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,1158,1183]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="E18B8576FFAC4A67FF328FE9FF0FEBC4" box="[151,251,1158,1183]" italics="true" pageId="44" pageNumber="45">Pheidole</emphasis>
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</taxonomicName>
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JTL-193: morphospecies code previously used on AntWeb.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF628FC4FDD9EA08" pageId="44" pageNumber="45" type="materials_examined">
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<paragraph id="D3405964FFAC4A67FF628FC4FEA2EA74" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
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<materialsCitation id="63975339FFAC4A67FF628FC4FE87EA50" ID-GBIF-Occurrence="2237851504" collectingDate="2009-05-29" collectingDateMax="2017-06-04" collectingDateMin="2009-05-29" collectionCode="MCZC" country="Guatemala" elevation="0" location="Machaquila" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Peten" typeStatus="holotype">
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<typeStatus id="0C44E7C6FFAC4A67FF628FC4FEABEB98" box="[199,351,1195,1219]" pageId="44" pageNumber="45" type="holotype">HOLOTYPE</typeStatus>
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: 1 major worker,
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<collectingCountry id="ABE819F4FFAC4A67FD838FC5FD56EB98" box="[550,674,1194,1219]" name="Guatemala" pageId="44" pageNumber="45">Guatemala</collectingCountry>
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,
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<collectingRegion id="113B9786FFAC4A67FD0B8FC4FD04EB98" box="[686,752,1195,1219]" country="Guatemala" name="Peten" pageId="44" pageNumber="45">Petén</collectingRegion>
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:
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<quantity id="1407F481FFAC4A67FD5E8FC5FCCFEB98" box="[763,827,1194,1219]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="44" pageNumber="45" unit="km" value="13.0">13km</quantity>
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NW
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<location id="D6200FBFFFAC4A67FCDC8FC5FBF4EB98" LSID="urn:lsid:plazi:treatment:5B56E872FFAC4A67FF328F73FDA0E880:D6200FBFFFAC4A67FCDC8FC5FBF4EB98" box="[889,1024,1194,1219]" country="Guatemala" name="Machaquila" pageId="44" pageNumber="45" stateProvince="Peten">Machaquilá</location>
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, 16.44612 -89.54969 ±
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<elevation id="58D2BE57FFAC4A67FAA38FC5FA63EB99" box="[1286,1431,1194,1219]" isEnumeration="true" pageId="44" pageNumber="45">
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<quantity id="1407F481FFAC4A67FAA38FC5FAB5EB99" box="[1286,1345,1194,1219]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="44" pageNumber="45" unit="m" value="59.0">59 m</quantity>
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,
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<quantity id="1407F481FFAC4A67FAEB8FC4FA63EB99" box="[1358,1431,1195,1219]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="44" pageNumber="45" unit="m" value="400.0">400 m</quantity>
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</elevation>
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,
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<date id="A7417FA4FFAC4A67FF328FA0FEC0EBBC" box="[151,308,1231,1255]" pageId="44" pageNumber="45" value="2009-05-29">
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<collectingDate id="B705864CFFAC4A67FF328FA0FEC0EBBC" box="[151,308,1231,1255]" pageId="44" pageNumber="45" value="2009-05-29">29-May-2009</collectingDate>
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</date>
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, tropical moist forest, at bait (LLAMA, Ba-B-
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<date id="A7417FA4FFAC4A67FCCD8FA1FC17EBBC" box="[872,995,1230,1255]" pageId="44" pageNumber="45" value="2017-06-04" valueMax="2017-06-04" valueMin="2017-06-04">
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<collectingDate id="B705864CFFAC4A67FCCD8FA1FC17EBBC" box="[872,995,1230,1255]" pageId="44" pageNumber="45" value="2017-06-04" valueMax="2017-06-04" valueMin="2017-06-04">06-4-04-17</collectingDate>
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</date>
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) [
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<collectionCode id="B5EEC1A1FFAC4A67FC5A8FA0FBA0EBBC" box="[1023,1108,1231,1255]" pageId="44" pageNumber="45">MCZC</collectionCode>
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, unique specimen identifier CASENT0614273]
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</materialsCitation>
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.
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<materialsCitation id="63975339FFAC4A67FE248F9CFEA7EA74" ID-GBIF-Occurrence="2237851446" collectingDate="2009-05-29" collectingDateMax="2017-06-04" collectingDateMin="2009-05-29" collectionCode="CAS, DZUP, JTLC, MCZC, USNM, UVGC" country="Guatemala" elevation="0" location="Machaquila" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Peten" typeStatus="paratype">
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<typeStatus id="0C44E7C6FFAC4A67FE248F9CFDD4EA50" box="[385,544,1267,1291]" pageId="44" pageNumber="45" type="paratype">PARATYPES</typeStatus>
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: major, minor workers: same data as holotype [
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<collectionCode id="B5EEC1A1FFAC4A67FBF48F9CFB78EA50" box="[1105,1164,1267,1291]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="44" pageNumber="45" type="Museum">CAS</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAC4A67FB398F9CFB11EA51" box="[1180,1253,1267,1290]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="44" pageNumber="45" type="University or college">DZUP</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAC4A67FB518F9CFAC3EA50" box="[1268,1335,1267,1291]" pageId="44" pageNumber="45">JTLC</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAC4A67FAE38F9CFA6CEA50" box="[1350,1432,1267,1291]" pageId="44" pageNumber="45">MCZC</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAC4A67FF328E78FF1FEA74" box="[151,235,1303,1327]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="44" pageNumber="45" type="Museum">USNM</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAC4A67FF538E78FEB8EA74" box="[246,332,1303,1327]" country="Guatemala" name="Collecion de Artropodos" pageId="44" pageNumber="45">UVGC</collectionCode>
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]
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="D3405964FFAC4A67FF628E55FDD9EA08" blockId="44.[151,1437,1158,2011]" box="[199,557,1338,1363]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628E55FE51EA08" bold="true" box="[199,421,1338,1363]" pageId="44" pageNumber="45">Geographic range.</emphasis>
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<collectingCountry id="ABE819F4FFAC4A67FE098E55FDDDEA08" box="[428,553,1338,1363]" name="Guatemala" pageId="44" pageNumber="45">Guatemala</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF628E31FCB2E929" pageId="44" pageNumber="45" type="diagnosis">
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<paragraph id="D3405964FFAC4A67FF628E31FCB2E929" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628E31FE62EA2C" bold="true" box="[199,406,1374,1399]" pageId="44" pageNumber="45">
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Diagnosis.
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<emphasis id="E18B8576FFAC4A67FEE88E31FE62EA2C" bold="true" box="[333,406,1374,1399]" italics="true" pageId="44" pageNumber="45">Minor</emphasis>
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</emphasis>
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: face uniformly foveolate; promesonotal groove absent, promesonotum evenly convex; entire mesosoma foveolate; propodeal spines about two thirds length of posterior face of propodeum; gaster shagreened, matte; face, mesosomal dorsum, and gastral dorsum with sparse, short, erect setae; tibiae without erect setae; color orange yellow.
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<emphasis id="E18B8576FFAC4A67FE408EA5FDD8EAB8" bold="true" box="[485,556,1482,1507]" italics="true" pageId="44" pageNumber="45">Major</emphasis>
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: inner hypostomal teeth distinct, closely spaced, blunt medial tooth also prominent; scape base terete; antennal scrobes very shallow, not sharply delimited; entire face foveolate, overlain with irregular, mostly longitudinal rugulae, these strongest anteriorly, fading posteriorly; propodeal spines about two thirds length of posterior face of propodeum; first gastral tergite entirely matte; abundant erect setae on sides of head, dorsal mesosoma, and gaster; tibiae lacking erect setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF628D11FD89E858" pageId="44" pageNumber="45" type="description">
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<paragraph id="D3405964FFAC4A67FF628D11FD78E9E0" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628D11FDD9E9CC" bold="true" box="[199,557,1662,1687]" pageId="44" pageNumber="45">Measurements, minor worker</emphasis>
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: HW 0.45, HL 0.54, SL 0.62, EL 0.10, WL 0.62, PSL 0.09, PTW 0.08, PPW 0.12, CI 84, SI 136, PSLI 20, PPI 144 (n=2).
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</paragraph>
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<paragraph id="D3405964FFAC4A67FF628DA9FD89E858" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628DA9FDD9E984" bold="true" box="[199,557,1734,1759]" pageId="44" pageNumber="45">Measurements, major worker</emphasis>
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: HW 0.93, HL 1.01, SL 0.67, EL 0.12, WL 0.82, PSL 0.12, PTW 0.13, PPW 0.22, CI 92, SI 72, PSLI 13, PPI 173 (n=2).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF628C61FD5AE834" pageId="44" pageNumber="45" type="biology_ecology">
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<paragraph id="D3405964FFAC4A67FF628C61FD5AE834" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628C61FED3E87C" bold="true" box="[199,295,1806,1831]" pageId="44" pageNumber="45">Biology.</emphasis>
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This species is known from one site in the
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<collectingRegion id="113B9786FFAC4A67FC888C60FC9FE87C" box="[813,875,1807,1831]" country="Guatemala" name="Peten" pageId="44" pageNumber="45">Petén</collectingRegion>
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region of
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<collectingCountry id="ABE819F4FFAC4A67FC4C8C61FB92E87C" box="[1001,1126,1806,1831]" name="Guatemala" pageId="44" pageNumber="45">Guatemala</collectingCountry>
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. It was an area with small patches of second growth moist forest on a karst landscape. The one collection was from a ground bait, with a strong recruitment of minor and major workers.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF628C15FAF2E8EC" pageId="44" pageNumber="45" type="discussion">
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<paragraph id="D3405964FFAC4A67FF628C15FAF2E8EC" blockId="44.[151,1437,1158,2011]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628C15FEBCE8C8" bold="true" box="[199,328,1914,1939]" pageId="44" pageNumber="45">Comments</emphasis>
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. This species is similar to
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<taxonomicName id="14FF22E7FFAC4A67FD268C13FCD1E8C8" authorityName="Longino" authorityYear="2009" box="[643,805,1914,1939]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="phanigaster">
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<emphasis id="E18B8576FFAC4A67FD268C13FCD1E8C8" box="[643,805,1914,1939]" italics="true" pageId="44" pageNumber="45">P. phanigaster</emphasis>
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</taxonomicName>
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. The minor worker differs in matte gaster (shiny in
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<taxonomicName id="14FF22E7FFAC4A67FA228C13FEE9E8EC" authorityName="Longino" authorityYear="2009" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="phanigaster">
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<emphasis id="E18B8576FFAC4A67FA228C13FEE9E8EC" italics="true" pageId="44" pageNumber="45">P. phanigaster</emphasis>
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</taxonomicName>
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). The major worker differs in longer scapes (mean SI 72 vs. 61) and lighter orange color.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAC4A67FF628CADFDA0E880" box="[199,596,1986,2011]" pageId="44" pageNumber="45" type="etymology">
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<paragraph id="D3405964FFAC4A67FF628CADFDA0E880" blockId="44.[151,1437,1158,2011]" box="[199,596,1986,2011]" pageId="44" pageNumber="45">
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<emphasis id="E18B8576FFAC4A67FF628CADFEBCE880" bold="true" box="[199,328,1986,2011]" pageId="44" pageNumber="45">Etymology</emphasis>
|
|
: From the
|
|
<typeStatus id="0C44E7C6FFAC4A67FE668CACFE07E880" box="[451,499,1987,2011]" pageId="44" pageNumber="45">type</typeStatus>
|
|
locality.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |