treatments-xml/data/5B/56/E8/5B56E872FFA64A6DFF328A00FBB1E8E2.xml
2024-06-21 12:37:31 +02:00

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<document id="C40A307EDF0D3CC088FF8E64E9C2DC95" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FFA64A6DFF328A00FBB1E8E2" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole indagarama Longino 2019, new species" docType="treatment" docVersion="22" lastPageNumber="39" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="39" updateTime="1698726215157" updateUser="ExternalLinkService">
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<mods:title id="790FA8E2456D2711E91E05F83FC39CEB">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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<mods:namePart id="7C6B2E20BD0352DC0CDB57414F5B1E43">Longino, John T.</mods:namePart>
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<mods:date id="873E86F79DA9E9CFD588721AECBBA36F">2019</mods:date>
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<mods:number id="04612975F92447ADFDE4FC592CF1EF05">2019-05-06</mods:number>
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<treatment id="5B56E872FFA64A6DFF328A00FBB1E8E2" ID-DOI="http://doi.org/10.5281/zenodo.5586196" ID-GBIF-Taxon="157062612" ID-Zenodo-Dep="5586196" LSID="urn:lsid:plazi:treatment:5B56E872FFA64A6DFF328A00FBB1E8E2" httpUri="http://treatment.plazi.org/id/5B56E872FFA64A6DFF328A00FBB1E8E2" lastPageNumber="39" pageId="38" pageNumber="39">
<subSubSection id="9BE50AEFFFA64A6DFF328A00FEF0EEF0" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="D3405964FFA64A6DFF328A00FDC6EED1" blockId="38.[151,562,367,427]" box="[151,562,367,394]" pageId="38" pageNumber="39">
<heading id="8808EE08FFA64A6DFF328A00FDC6EED1" bold="true" box="[151,562,367,394]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="E18B8576FFA64A6DFF328A00FDC6EED1" bold="true" box="[151,562,367,394]" pageId="38" pageNumber="39">
<taxonomicName id="14FF22E7FFA64A6DFF328A00FE6FEED2" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,411,367,393]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagarama" status="sp. nov.">
<emphasis id="E18B8576FFA64A6DFF328A00FE6FEED2" bold="true" box="[151,411,367,393]" italics="true" pageId="38" pageNumber="39">Pheidole indagarama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAB8380DFFA64A6DFE078A1FFDC6EED1" box="[418,562,368,394]" pageId="38" pageNumber="39" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D3405964FFA64A6DFF328AFDFEF0EEF0" blockId="38.[151,562,367,427]" box="[151,260,402,427]" pageId="38" pageNumber="39">
(
<figureCitation id="4BC445E1FFA64A6DFF3B8AFDFF08EEF0" box="[158,252,402,427]" captionStart="PLATE 45" captionStartId="111.[151,238,1954,1977]" captionTargetId="figure@111.[202,1384,194,1933]" captionTargetPageId="111" captionText="PLATE 45." httpUri="https://zenodo.org/record/2668323/files/figure.png" pageId="38" pageNumber="39">Plate 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF328AB5FC6FEEA8" box="[151,923,474,499]" pageId="38" pageNumber="39" type="description">
<paragraph id="D3405964FFA64A6DFF328AB5FC6FEEA8" blockId="38.[151,1437,474,1978]" box="[151,923,474,499]" pageId="38" pageNumber="39">
<taxonomicName id="14FF22E7FFA64A6DFF328AB5FF0FEEA8" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,474,499]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="E18B8576FFA64A6DFF328AB5FF0FEEA8" box="[151,251,474,499]" italics="true" pageId="38" pageNumber="39">Pheidole</emphasis>
</taxonomicName>
JTL-153: morphospecies code previously used on AntWeb.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF628A90FDDAEDFC" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="D3405964FFA64A6DFF628A90FDF0EDD8" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<materialsCitation id="63975339FFA64A6DFF628A90FDA7ED04" ID-GBIF-Occurrence="2237851545" collectingDate="2000-05-16" collectionCode="MUCR" country="Costa Rica" elevation="0" location="La Selva Biological Station" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="0C44E7C6FFA64A6DFF628A90FEABED4C" box="[199,351,511,535]" pageId="38" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
: 1 major worker,
<collectingCountry id="ABE819F4FFA64A6DFD828A90FD51ED4C" box="[551,677,510,535]" name="Costa Rica" pageId="38" pageNumber="39">Costa Rica</collectingCountry>
,
<collectingRegion id="113B9786FFA64A6DFD178A91FCFBED4C" box="[690,783,510,535]" country="Costa Rica" name="Heredia" pageId="38" pageNumber="39">Heredia</collectingRegion>
:
<location id="D6200FBFFFA64A6DFCBF8A90FBA1ED4C" LSID="urn:lsid:plazi:treatment:5B56E872FFA64A6DFF328A00FBB1E8E2:D6200FBFFFA64A6DFCBF8A90FBA1ED4C" box="[794,1109,510,535]" country="Costa Rica" name="La Selva Biological Station" pageId="38" pageNumber="39" stateProvince="Heredia">La Selva Biological Station</location>
, 10.43215 -84.01511 ±
<elevation id="58D2BE57FFA64A6DFAFE8A90FF2CED61" isEnumeration="true" pageId="38" pageNumber="39">
<quantity id="1407F481FFA64A6DFAFE8A90FA63ED4D" box="[1371,1431,511,535]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="39" unit="m" value="20.0">20 m</quantity>
,
<quantity id="1407F481FFA64A6DFF32894DFF2CED61" box="[151,216,546,571]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="38" pageNumber="39" unit="m" value="50.0">50 m</quantity>
</elevation>
,
<date id="A7417FA4FFA64A6DFF4F894DFE73ED60" box="[234,391,546,571]" pageId="38" pageNumber="39" value="2000-05-16">
<collectingDate id="B705864CFFA64A6DFF4F894DFE73ED60" box="[234,391,546,571]" pageId="38" pageNumber="39" value="2000-05-16">16-May-2000</collectingDate>
</date>
, secondgrowth rainforest, ex
<taxonomicName id="14FF22E7FFA64A6DFD4C894DFC2DED60" authorityName="Burret" authorityYear="1926" box="[745,985,546,571]" class="Aves" family="Trochilidae" genus="Goethalsia" kingdom="Animalia" order="Apodiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="meiantha">
<emphasis id="E18B8576FFA64A6DFD4C894DFC2DED60" box="[745,985,546,571]" italics="true" pageId="38" pageNumber="39">Goethalsia meiantha</emphasis>
</taxonomicName>
(ALAS, FOT/51/03) [
<collectionCode id="B5EEC1A1FFA64A6DFB43894CFACAED60" box="[1254,1342,547,571]" country="Costa Rica" name="Museo de Insectos" pageId="38" pageNumber="39">MUCR</collectionCode>
, unique specimen identifier CASENT0636566]
</materialsCitation>
.
<materialsCitation id="63975339FFA64A6DFDFA8928FDF5EDD8" ID-GBIF-Occurrence="2237851434" collectingDate="2000-05-16" collectionCode="CAS, JTLC, MCZC, MUCR, USNM" country="Costa Rica" elevation="0" location="La Selva Biological Station" pageId="38" pageNumber="39" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<typeStatus id="0C44E7C6FFA64A6DFDFA8928FD0AED04" box="[607,766,583,607]" pageId="38" pageNumber="39" type="paratype">PARATYPES</typeStatus>
: major, minor worker, queen: same data as holotype [
<collectionCode id="B5EEC1A1FFA64A6DFAF98928FA63ED04" box="[1372,1431,583,607]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="39" type="Museum">CAS</collectionCode>
,
<collectionCode id="B5EEC1A1FFA64A6DFF328904FF2EEDD8" box="[151,218,619,643]" pageId="38" pageNumber="39">JTLC</collectionCode>
,
<collectionCode id="B5EEC1A1FFA64A6DFF418904FEC2EDD8" box="[228,310,619,643]" pageId="38" pageNumber="39">MCZC</collectionCode>
,
<collectionCode id="B5EEC1A1FFA64A6DFEE78904FE63EDD8" box="[322,407,619,643]" country="Costa Rica" name="Museo de Insectos" pageId="38" pageNumber="39">MUCR</collectionCode>
,
<collectionCode id="B5EEC1A1FFA64A6DFE078904FE0DEDD8" box="[418,505,619,643]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="39" type="Museum">USNM</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="D3405964FFA64A6DFF6289E1FDDAEDFC" blockId="38.[151,1437,474,1978]" box="[199,558,654,679]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF6289E1FE51EDFC" bold="true" box="[199,421,654,679]" pageId="38" pageNumber="39">Geographic range.</emphasis>
<collectingCountry id="ABE819F4FFA64A6DFE0989E0FDDEEDFC" box="[428,554,654,679]" name="Costa Rica" pageId="38" pageNumber="39">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF6289DDFCE4ECB0" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="D3405964FFA64A6DFF6289DDFCE4ECB0" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF6289DDFE61ED90" bold="true" box="[199,405,690,715]" pageId="38" pageNumber="39">
Diagnosis.
<emphasis id="E18B8576FFA64A6DFEE989DDFE61ED90" bold="true" box="[332,405,690,715]" italics="true" pageId="38" pageNumber="39">Minor</emphasis>
</emphasis>
: face smooth and shiny; head tapering behind to moderately developed occipital carina; promesonotal groove present, distinctly impressed; promesonotum entirely smooth and shining; katepisternum uniformly foveolate; propodeal spines spiniform, about one third length of posterior face of propodeum; gaster smooth and shining; mesosomal and gastral dorsum with abundant flexuous erect setae; tibiae with erect setae; color red brown.
<emphasis id="E18B8576FFA64A6DFEC9882DFE47EC00" bold="true" box="[364,435,834,859]" italics="true" pageId="38" pageNumber="39">Major</emphasis>
: inner hypostomal teeth widely spaced, acicular; scape base terete; face uniformly foveolate, with longitudinal carinulae between eye and frontal carinae; propodeal spines about one fourth length of posterior face of propodeum; gastral dorsum very faintly foveolate on anterior half; side of head with abundant erect setae; mesosomal and gastral dorsum with abundant erect setae; tibiae with erect setae; head bright yellow, contrasting with darker red brown mesosoma and gaster.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF628899FDDCEB98" pageId="38" pageNumber="39" type="description">
<paragraph id="D3405964FFA64A6DFF628899FD7EEB68" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF628899FDD9EB54" bold="true" box="[199,557,1014,1039]" pageId="38" pageNumber="39">Measurements, minor worker</emphasis>
: HW 0.62, HL 0.79, SL 1.06, EL 0.17, WL 1.00, PSL 0.07, PTW 0.11, PPW 0.16, CI 78, SI 172, PSLI 11, PPI 143 (n=3).
</paragraph>
<paragraph id="D3405964FFA64A6DFF628F51FD9BEB20" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF628F51FDD9EB0C" bold="true" box="[199,557,1086,1111]" pageId="38" pageNumber="39">Measurements, major worker</emphasis>
: HW 1.23, HL 1.27, SL 1.04, EL 0.20, WL 1.25, PSL 0.09, PTW 0.18, PPW 0.24, CI 97, SI 85, PSLI 7, PPI 136 (n=2).
</paragraph>
<paragraph id="D3405964FFA64A6DFF628FE9FDDCEB98" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF628FE9FE38EBC4" bold="true" box="[199,460,1158,1183]" pageId="38" pageNumber="39">Measurements, queen</emphasis>
: HW 1.48, HL 1.30, SL 0.82, EL 0.37, WL 2.20, PSL 0.19, PTW 0.47, PPW 0.74, CI 114, SI 55, PSLI 13, PPI 157 (n=1).
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF628FA1FECDEA74" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="D3405964FFA64A6DFF628FA1FECDEA74" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF628FA1FED3EBBC" bold="true" box="[199,295,1230,1255]" pageId="38" pageNumber="39">Biology.</emphasis>
This species occurs in lowland rainforest. It is arboreal, known from canopy fogging samples and specimens collected in a fresh treefall. Major workers, minor workers, males, and dealate queens have occurred in these samples.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF628E55FE78E8CD" pageId="38" pageNumber="39" type="discussion">
<paragraph id="D3405964FFA64A6DFF628E55FD09E928" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF628E55FEBCEA08" bold="true" box="[199,328,1338,1363]" pageId="38" pageNumber="39">Comments</emphasis>
. This species is known only from La Selva Biological Station, from two canopy fogging samples (taken from different trees on different dates) and in a hand collection in a new treefall. It appears to be a local variant of the widespread
<taxonomicName id="14FF22E7FFA64A6DFE1C8EEBFDBDEAC0" authorityName="Wilson" authorityYear="2003" box="[441,585,1410,1435]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFE1C8EEBFDBDEAC0" box="[441,585,1410,1435]" italics="true" pageId="38" pageNumber="39">P. indagatrix</emphasis>
</taxonomicName>
, specialized for the high canopy. The only consistent character difference in workers is the color of the major worker. Throughout the range of
<taxonomicName id="14FF22E7FFA64A6DFCD98EC8FBF9EAE4" authorityName="Wilson" authorityYear="2003" box="[892,1037,1446,1471]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFCD98EC8FBF9EAE4" box="[892,1037,1446,1471]" italics="true" pageId="38" pageNumber="39">P. indagatrix</emphasis>
</taxonomicName>
, from
<collectingCountry id="ABE819F4FFA64A6DFBF28EC9FB24EAE4" box="[1111,1232,1446,1471]" name="Guatemala" pageId="38" pageNumber="39">Guatemala</collectingCountry>
to
<collectingCountry id="ABE819F4FFA64A6DFB518EC8FAA4EAE5" box="[1268,1360,1447,1470]" name="Panama" pageId="38" pageNumber="39">Panama</collectingCountry>
, major workers are dark red brown.
<taxonomicName id="14FF22E7FFA64A6DFE458EA5FD35EAB8" authorityName="Wilson" authorityYear="2003" box="[480,705,1482,1507]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFE458EA5FD35EAB8" box="[480,705,1482,1507]" italics="true" pageId="38" pageNumber="39">Pheidole indagatrix</emphasis>
</taxonomicName>
is very common at ground baits and nests in leaf litter, in rotten wood, and in dead stems in the low arboreal zone. At La Selva Biological Station,
<taxonomicName id="14FF22E7FFA64A6DFBCD8E80FB08E95C" authorityName="Wilson" authorityYear="2003" box="[1128,1276,1518,1543]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFBCD8E80FB08E95C" box="[1128,1276,1518,1543]" italics="true" pageId="38" pageNumber="39">P. indagatrix</emphasis>
</taxonomicName>
is a common ground ant, while
<taxonomicName id="14FF22E7FFA64A6DFEC78D7BFDF2E970" authorityName="Longino" authorityYear="2019" box="[354,518,1554,1579]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagarama">
<emphasis id="E18B8576FFA64A6DFEC78D7BFDF2E970" box="[354,518,1554,1579]" italics="true" pageId="38" pageNumber="39">P. indagarama</emphasis>
</taxonomicName>
is known only from the canopy. There is a tendency for majors of
<taxonomicName id="14FF22E7FFA64A6DFB4A8D7BFA8BE970" authorityName="Wilson" authorityYear="2003" box="[1263,1407,1554,1579]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFB4A8D7BFA8BE970" box="[1263,1407,1554,1579]" italics="true" pageId="38" pageNumber="39">P. indagatrix</emphasis>
</taxonomicName>
to have heavier sculpture on the gaster. Major workers of
<taxonomicName id="14FF22E7FFA64A6DFCAC8D58FC5BE914" authorityName="Longino" authorityYear="2019" box="[777,943,1590,1615]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagarama">
<emphasis id="E18B8576FFA64A6DFCAC8D58FC5BE914" box="[777,943,1590,1615]" italics="true" pageId="38" pageNumber="39">P. indagarama</emphasis>
</taxonomicName>
have the first gastral tergite nearly smooth, with very faint foveolate sculpture on the anterior half.
</paragraph>
<paragraph id="D3405964FFA64A6DFF628DEDFE78E8CD" blockId="38.[151,1437,474,1978]" pageId="38" pageNumber="39">
Two dealate queens and some
<taxonomicName id="14FF22E7FFA64A6DFD888DEEFD65E9C1" authorityName="Biolleyi Forel" authorityYear="1908" box="[557,657,1665,1690]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="E18B8576FFA64A6DFD888DEEFD65E9C1" box="[557,657,1665,1690]" italics="true" pageId="38" pageNumber="39">Pheidole</emphasis>
</taxonomicName>
males were found in the same
<quantity id="1407F481FFA64A6DFBA08DEDFBC7E9C2" box="[1029,1075,1666,1690]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1.0">1 m</quantity>
<superScript id="248AF42CFFA64A6DFB968D11FBCFE9D7" attach="left" box="[1075,1083,1662,1676]" fontSize="6" pageId="38" pageNumber="39">2</superScript>
fogging funnel with abundant minor and major workers of
<emphasis id="E18B8576FFA64A6DFE4E8DC9FD6FE9E5" box="[491,667,1701,1726]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="14FF22E7FFA64A6DFE4E8DC9FD63E9E5" authorityName="Longino" authorityYear="2019" box="[491,663,1701,1726]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagarama">P. indagarama</taxonomicName>
.
</emphasis>
These were assumed to be from one colony, and thus the queens conspecific with the workers. The queens have the mesoscutum and scutellum completely smooth and shining, and the first gastral tergite is almost entirely smooth and shining.
<taxonomicName id="14FF22E7FFA64A6DFCFE8D82FBC8E85D" authorityName="Wilson" authorityYear="2003" box="[859,1084,1773,1798]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFCFE8D82FBC8E85D" box="[859,1084,1773,1798]" italics="true" pageId="38" pageNumber="39">Pheidole indagatrix</emphasis>
</taxonomicName>
queens from the
<collectingRegion id="113B9786FFA64A6DFAA28D82FA8DE85D" box="[1287,1401,1773,1798]" country="Paraguay" name="Cordillera" pageId="38" pageNumber="39">Cordillera</collectingRegion>
de Tilarán,
<quantity id="1407F481FFA64A6DFF5D8C7EFEB6E871" box="[248,322,1809,1834]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="38" pageNumber="39" unit="km" value="85.0">85 km</quantity>
west of La Selva, have the mesoscutum and scutellum coarsely sculptured, and the first gastral tergite is strongly shagreened and opaque. An isolated queen from a different fogging sample from La Selva is tentatively identified as
<taxonomicName id="14FF22E7FFA64A6DFE038C35FDC3E829" authorityName="Wilson" authorityYear="2003" box="[422,567,1881,1906]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFE038C35FDC3E829" box="[422,567,1881,1906]" italics="true" pageId="38" pageNumber="39">P. indagatrix</emphasis>
</taxonomicName>
. It is lighter colored but otherwise has the heavier sculpture of the
<collectingRegion id="113B9786FFA64A6DFA8F8C36FA68E829" box="[1322,1436,1881,1906]" country="Paraguay" name="Cordillera" pageId="38" pageNumber="39">Cordillera</collectingRegion>
de Tilarán specimens.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFFA64A6DFF628CCEFBB1E8E2" box="[199,1093,1953,1978]" pageId="38" pageNumber="39" type="etymology">
<paragraph id="D3405964FFA64A6DFF628CCEFBB1E8E2" blockId="38.[151,1437,474,1978]" box="[199,1093,1953,1978]" pageId="38" pageNumber="39">
<emphasis id="E18B8576FFA64A6DFF628CCEFEBCE8E1" bold="true" box="[199,328,1953,1978]" pageId="38" pageNumber="39">Etymology</emphasis>
: Referring to this species being an arboreal version of
<taxonomicName id="14FF22E7FFA64A6DFC0B8CCDFBCAE8E1" authorityName="Wilson" authorityYear="2003" box="[942,1086,1953,1978]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="indagatrix">
<emphasis id="E18B8576FFA64A6DFC0B8CCDFBCAE8E1" box="[942,1086,1953,1978]" italics="true" pageId="38" pageNumber="39">P. indagatrix</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>