256 lines
39 KiB
XML
256 lines
39 KiB
XML
<document id="107444D1C41E4C69B730EB030A396489" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="11755326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FFD939694D2DF925FE05FAC4" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Geostiba (Sibiota) silvigena Gusarov, sp. n." docType="treatment" docVersion="14" lastPageNumber="75" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="72" updateTime="1698188143631" updateUser="plazi">
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<mods:titleInfo id="E5A7DC4989FAC038892E2F1F9C8B2F60">
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<mods:title id="6451801CA3F79AA32F28CA4D7CF81BCF">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="1C413917162B7FC5782D8AAE1F65A3E6" type="personal">
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<mods:namePart id="C6222199E8B3FD8412096E089565F247">Gusarov, Vladimir I.</mods:namePart>
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<mods:typeOfResource id="3E754CBB06A5E1FB19B41122AE7EBEEB">text</mods:typeOfResource>
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<mods:title id="409935B4F736EDBAFD16AD9C9139CF7F">Zootaxa</mods:title>
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<mods:part id="51E8F3D30D3E6D6244D073FF2363D61A">
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<mods:date id="AD2D94619D32C1EC975DEA9B953BB4B2">2002</mods:date>
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<mods:detail id="A5FA4BCE6ED1300E6AD00B97E7B592A8" type="volume">
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<mods:number id="C02F0B134528B406F32EE765895E1792">81</mods:number>
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<mods:start id="1C3EC0F621E0EE1399BAEE2966570FD2">1</mods:start>
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<mods:classification id="3C746EDB6514F69C3066E22DC2200BBC">journal article</mods:classification>
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<mods:identifier id="5E1A929670E3D5E1E236EBFA1490F352" type="DOI">10.5281/zenodo.155701</mods:identifier>
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<mods:identifier id="EE97A1BF4CCFBED48BAB6A3EDB0AD92D" type="ISSN">11755326</mods:identifier>
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<treatment id="5B50E916FFD939694D2DF925FE05FAC4" ID-DOI="http://doi.org/10.5281/zenodo.6277587" ID-GBIF-Taxon="119330228" ID-Zenodo-Dep="6277587" LSID="urn:lsid:plazi:treatment:5B50E916FFD939694D2DF925FE05FAC4" httpUri="http://treatment.plazi.org/id/5B50E916FFD939694D2DF925FE05FAC4" lastPageId="74" lastPageNumber="75" pageId="71" pageNumber="72">
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<subSubSection id="9BE30B8BFFD939644D2DF925FBE8F969" box="[264,1024,1689,1715]" pageId="71" pageNumber="72" type="nomenclature">
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<paragraph id="D3465800FFD939644D2DF925FBE8F969" blockId="71.[264,1024,1689,1715]" box="[264,1024,1689,1715]" pageId="71" pageNumber="72">
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<heading id="880EEF6CFFD939644D2DF925FBE8F969" bold="true" box="[264,1024,1689,1715]" fontSize="11" level="1" pageId="71" pageNumber="72" reason="1">
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<emphasis id="E18D8412FFD939644D2DF925FBE8F969" bold="true" box="[264,1024,1689,1715]" pageId="71" pageNumber="72">
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14.
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<taxonomicName id="14F92383FFD939644D11F925FD1AF969" ID-CoL="3FSNK" authority="Gusarov" authorityName="Gusarov" box="[308,754,1689,1715]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="silvigena" status="sp. nov." subGenus="Sibiota">
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<emphasis id="E18D8412FFD939644D11F925FE76F969" bold="true" box="[308,414,1689,1715]" italics="true" pageId="71" pageNumber="72">Geostiba</emphasis>
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(
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<emphasis id="E18D8412FFD939644D8AF925FDEBF969" bold="true" box="[431,515,1689,1715]" italics="true" pageId="71" pageNumber="72">Sibiota</emphasis>
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)
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<emphasis id="E18D8412FFD939644E30F925FD68F969" bold="true" box="[533,640,1689,1715]" italics="true" pageId="71" pageNumber="72">silvigena</emphasis>
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Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="FABE3969FFD939644EDAF925FCA9F969" box="[767,833,1689,1715]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="4BC24485FFD939644F74F925FC10F969" box="[849,1016,1689,1715]" captionStart-0="FIGURES 321 326" captionStart-1="FIGURES 327 335" captionStartId-0="72.[264,383,1696,1720]" captionStartId-1="73.[264,383,1480,1504]" captionTargetBox-0="[286,1297,660,1656]" captionTargetBox-1="[307,1280,824,1441]" captionTargetId-0="figure@72.[275,1311,649,1670]" captionTargetId-1="figure@73.[296,1290,809,1454]" captionTargetPageId-0="72" captionTargetPageId-1="73" captionText-0="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." captionText-1="FIGURES 327 335. Genitalia of Geostiba silvigena Gusarov, sp. n. (holotype). 327 – details of internal sac retracted into median lobe, ventral view; 328 – copulatory piece of internal sac, dorsal view; 329 – distal diverticula of everted internal sac, ventral view; 330 – everted internal sac, ventral view; 331 – everted internal sac, lateral view; 332 333 – details of internal sac retracted into median lobe, lateral view; 334 – apex of left paramere, side facing median lobe; 335 – copulatory piece of internal sac, lateral view. Scale bar 0.4 mm (327, 330 333), 0.2 mm (328 329, 334 335)." httpUri-0="https://zenodo.org/record/155746/files/figure.png" httpUri-1="https://zenodo.org/record/155747/files/figure.png" pageId="71" pageNumber="72">Figs. 321335</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE30B8BFFD939644D2DF958FD75F894" pageId="71" pageNumber="72" type="materials_examined">
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<paragraph id="D3465800FFD939644D2DF958FD75F894" blockId="71.[264,1323,1764,1870]" pageId="71" pageNumber="72">
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<emphasis id="E18D8412FFD939644D2DF958FE56F924" bold="true" box="[264,446,1764,1790]" pageId="71" pageNumber="72">
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<typeStatus id="0C42E6A2FFD939644D2DF958FEADF924" box="[264,325,1764,1790]" pageId="71" pageNumber="72">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="0C42E6A2FFD939644DE8F958FDD5F924" box="[461,573,1764,1790]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
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,,
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<emphasis id="E18D8412FFD939644E4FF958FC1AF924" bold="true" box="[618,1010,1764,1790]" pageId="71" pageNumber="72">
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<collectingCountry id="ABEE1890FFD939644E4FF958FCBDF924" box="[618,853,1764,1790]" name="United States of America" pageId="71" pageNumber="72">UNITED STATES</collectingCountry>
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: California:
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</emphasis>
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Mendocino Co.: Mendocino [
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<geoCoordinate id="B6CD3EC7FFD939644D6FF8B0FE3EF8FC" box="[330,470,1804,1830]" direction="north" orientation="latitude" pageId="71" pageNumber="72" precision="15" value="39.307777">39°18'28"N</geoCoordinate>
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<geoCoordinate id="B6CD3EC7FFD939644DFAF8B0FD6BF8FC" box="[479,643,1804,1830]" direction="west" orientation="longitude" pageId="71" pageNumber="72" precision="15" value="-123.79806">123°47'53"W</geoCoordinate>
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], ex moss in bed of dried up woodland pool, Berlese funnel (J.R.Helfer),
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<date id="A7477EC0FFD939644DEBF888FDDDF894" box="[462,565,1844,1870]" pageId="71" pageNumber="72" value="1954-10-01">1.x.1954</date>
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(
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<collectionCode id="B5E8C0C5FFD939644E61F888FD78F894" box="[580,656,1844,1870]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="71" pageNumber="72">CNCI</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE30B8BFFD6396B4D1DFEA7FBD7FD94" pageId="72" pageNumber="73" type="diagnosis">
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<paragraph id="D3465800FFD6396B4D1DFEA7FBB1FE0C" blockId="72.[264,1323,283,590]" pageId="72" pageNumber="73">
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<emphasis id="E18D8412FFD6396B4D1DFEA7FE50FEEF" bold="true" box="[312,440,283,309]" pageId="72" pageNumber="73">Diagnosis.</emphasis>
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<taxonomicName id="14F92383FFD6396B4DEBFEA0FD45FEEF" box="[462,685,284,309]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="silvigena">
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<emphasis id="E18D8412FFD6396B4DEBFEA0FD45FEEF" box="[462,685,284,309]" italics="true" pageId="72" pageNumber="73">Geostiba silvigena</emphasis>
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</taxonomicName>
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can be distinguished from other Nearctic species of
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<taxonomicName id="14F92383FFD6396B4D2DFEF8FE9AFE87" box="[264,370,324,349]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="E18D8412FFD6396B4D2DFEF8FE9AFE87" box="[264,370,324,349]" italics="true" pageId="72" pageNumber="73">Geostiba</emphasis>
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</taxonomicName>
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by having large eyes (temple length to eye length ratio 2.3), pronotal pubescence of
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<typeStatus id="0C42E6A2FFD6396B4D0DFED0FEB2FE5C" box="[296,346,364,390]" pageId="72" pageNumber="73">type</typeStatus>
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V, well developed wings, long elytra (pronotum length to elytron length ratio 1.0), the presence of two short parallel carinae in the middle of male abdominal tergum
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<quantity id="1401F5E5FFD6396B48DCFE28FAC1FE74" box="[1273,1321,404,430]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="72" pageNumber="73" unit="in" value="7.0">7 in</quantity>
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front of posterior margin, and the shape of the aedeagus (
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<figureCitation id="4BC24485FFD6396B4F81FE00FBA4FE0C" box="[932,1100,444,470]" captionStart-0="FIGURES 321 326" captionStart-1="FIGURES 327 335" captionStartId-0="72.[264,383,1696,1720]" captionStartId-1="73.[264,383,1480,1504]" captionTargetBox-0="[286,1297,660,1656]" captionTargetBox-1="[307,1280,824,1441]" captionTargetId-0="figure@72.[275,1311,649,1670]" captionTargetId-1="figure@73.[296,1290,809,1454]" captionTargetPageId-0="72" captionTargetPageId-1="73" captionText-0="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." captionText-1="FIGURES 327 335. Genitalia of Geostiba silvigena Gusarov, sp. n. (holotype). 327 – details of internal sac retracted into median lobe, ventral view; 328 – copulatory piece of internal sac, dorsal view; 329 – distal diverticula of everted internal sac, ventral view; 330 – everted internal sac, ventral view; 331 – everted internal sac, lateral view; 332 333 – details of internal sac retracted into median lobe, lateral view; 334 – apex of left paramere, side facing median lobe; 335 – copulatory piece of internal sac, lateral view. Scale bar 0.4 mm (327, 330 333), 0.2 mm (328 329, 334 335)." httpUri-0="https://zenodo.org/record/155746/files/figure.png" httpUri-1="https://zenodo.org/record/155747/files/figure.png" pageId="72" pageNumber="73">Figs. 323335</figureCitation>
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).
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</paragraph>
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<paragraph id="D3465800FFD6396B4D1DFE58FBD7FD94" blockId="72.[264,1323,283,590]" pageId="72" pageNumber="73">
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<taxonomicName id="14F92383FFD6396B4D1DFE58FDFDFE27" box="[312,533,484,509]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="silvigena">
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<emphasis id="E18D8412FFD6396B4D1DFE58FDFDFE27" box="[312,533,484,509]" italics="true" pageId="72" pageNumber="73">Geostiba silvigena</emphasis>
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</taxonomicName>
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is very similar to eastern Nearctic
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<taxonomicName id="14F92383FFD6396B4FE1FE58FB70FE27" box="[964,1176,484,509]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="appalachigena">
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<emphasis id="E18D8412FFD6396B4FE1FE58FB70FE27" box="[964,1176,484,509]" italics="true" pageId="72" pageNumber="73">G. appalachigena</emphasis>
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</taxonomicName>
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, and differs only in having the apex of median lobe less bent ventrally (in lateral view) (
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<figureCitation id="4BC24485FFD6396B48A5FDB0FACFFDFC" box="[1152,1319,524,550]" captionStart="FIGURES 321 326" captionStartId="72.[264,383,1696,1720]" captionTargetBox="[286,1297,660,1656]" captionTargetId="figure@72.[275,1311,649,1670]" captionTargetPageId="72" captionText="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="72" pageNumber="73">Figs. 325326</figureCitation>
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,
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<figureCitation id="4BC24485FFD6396B4D2DFD88FEB8FD94" box="[264,336,564,590]" captionStart="FIGURES 34 37" captionStartId="21.[264,383,1148,1172]" captionTargetBox="[321,1268,571,1127]" captionTargetId="figure@21.[319,1272,569,1127]" captionTargetPageId="21" captionText="FIGURES 34 37. Aedeagus of Geostiba appalachigena Gusarov, sp. n. (paratype from Black Mountain, West Virginia). 34 – median lobe, ventral view; 35 – apex of median lobe, ventral view; 36 – median lobe, lateral view; 37 – apex of median lobe, lateral view. Scale bar 0.2 mm (34, 36), 0.1 mm (35, 37)." httpUri="https://zenodo.org/record/155708/files/figure.png" pageId="72" pageNumber="73">3637</figureCitation>
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), and longer and more narrow apex of paramere (
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<figureCitation id="4BC24485FFD6396B4FB1FD88FBEDFD94" box="[916,1029,564,590]" captionStart="FIGURES 327 335" captionStartId="73.[264,383,1480,1504]" captionTargetBox="[307,1280,824,1441]" captionTargetId="figure@73.[296,1290,809,1454]" captionTargetPageId="73" captionText="FIGURES 327 335. Genitalia of Geostiba silvigena Gusarov, sp. n. (holotype). 327 – details of internal sac retracted into median lobe, ventral view; 328 – copulatory piece of internal sac, dorsal view; 329 – distal diverticula of everted internal sac, ventral view; 330 – everted internal sac, ventral view; 331 – everted internal sac, lateral view; 332 333 – details of internal sac retracted into median lobe, lateral view; 334 – apex of left paramere, side facing median lobe; 335 – copulatory piece of internal sac, lateral view. Scale bar 0.4 mm (327, 330 333), 0.2 mm (328 329, 334 335)." httpUri="https://zenodo.org/record/155747/files/figure.png" pageId="72" pageNumber="73">Figs. 334</figureCitation>
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,
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<figureCitation id="4BC24485FFD6396B4834FD88FBDBFD94" box="[1041,1075,564,590]" captionStart="FIGURES 38 50" captionStartId="22.[264,383,1403,1427]" captionTargetBox="[306,1285,460,1385]" captionTargetId="figure@22.[294,1294,449,1393]" captionTargetPageId="22" captionText="FIGURES 38 50. Genitalia of Geostiba appalachigena Gusarov, sp. n. (paratypes from Black Mountain (38, 40 45, 47 50) and Back Allegheny Mountain (39, 46), West Virginia). 38 – details of internal sac retracted into median lobe, ventral view; 39 – copulatory piece of internal sac, dorsal view; 40 – medial lamellae of internal sac, ventral view; 41 distal diverticula of everted internal sac, ventral view; 42 – everted internal sac, ventral view; 43 – spermatheca; 44 45 – details of internal sac retracted into median lobe, lateral view; 46 – copulatory piece of internal sac, lateral view; 47 – medial lamella of internal sac, lateral view; 48 – everted internal sac, lateral view; 49 – right distal diverticulum of everted internal sac, lateral view; 50 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (38, 42, 44 45, 48), 0.1 mm (39 41, 43, 46 47, 49 50)." httpUri="https://zenodo.org/record/155709/files/figure.png" pageId="72" pageNumber="73">50</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<caption id="87860888FFD6396B4D2DF91CFC29F8C2" httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="72" pageNumber="73" targetBox="[286,1297,660,1656]" targetPageId="72">
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||
<paragraph id="D3465800FFD6396B4D2DF91CFC29F8C2" blockId="72.[264,1323,1696,1816]" pageId="72" pageNumber="73">
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<emphasis id="E18D8412FFD6396B4D2DF91CFE08F962" bold="true" box="[264,480,1696,1720]" pageId="72" pageNumber="73">FIGURES 321326</emphasis>
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. Abdominal segment 8 and aedeagus of
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<taxonomicName id="14F92383FFD6396B4F95F91DFB36F962" authority="Gusarov" authorityName="Gusarov" box="[944,1246,1697,1720]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="silvigena" status="sp. nov.">
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<emphasis id="E18D8412FFD6396B4F95F91DFB91F962" box="[944,1145,1697,1720]" italics="true" pageId="72" pageNumber="73">Geostiba silvigena</emphasis>
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||
Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="FABE3969FFD6396B48C9F91CFAC2F962" box="[1260,1322,1696,1720]" pageId="72" pageNumber="73" rank="species">
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<emphasis id="E18D8412FFD6396B48C9F91CFAC2F962" bold="true" box="[1260,1322,1696,1720]" pageId="72" pageNumber="73">sp. n.</emphasis>
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</taxonomicNameLabel>
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(holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321323, 325), 0.2 mm (324, 326).
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</paragraph>
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</caption>
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<subSubSection id="9BE30B8BFFD739694D1DFEA0FDE3FCCC" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="description">
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<paragraph id="D3465800FFD7396A4D1DFEA0FE78FE0C" blockId="73.[264,1324,284,750]" pageId="73" pageNumber="74">
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<taxonomicName id="14F92383FFD7396A4D1DFEA0FDFFFEEF" box="[312,535,284,309]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="silvigena">
|
||
<emphasis id="E18D8412FFD7396A4D1DFEA0FDFFFEEF" box="[312,535,284,309]" italics="true" pageId="73" pageNumber="74">Geostiba silvigena</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="14F92383FFD7396A4E99FEA0FC8CFEEF" box="[700,868,284,309]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="impressula">
|
||
<emphasis id="E18D8412FFD7396A4E99FEA0FC8CFEEF" box="[700,868,284,309]" italics="true" pageId="73" pageNumber="74">G. impressula</emphasis>
|
||
</taxonomicName>
|
||
, another western Nearctic species, in having pronotal pubescence of
|
||
<typeStatus id="0C42E6A2FFD7396A4E52FEF8FD41FE84" box="[631,681,324,350]" pageId="73" pageNumber="74">type</typeStatus>
|
||
V, well developed wings, longer elytra, aedeagus with more narrow apex of the median lobe (in ventral view) (
|
||
<figureCitation id="4BC24485FFD7396A4FB2FED0FBD6FE5C" box="[919,1086,364,390]" captionStart="FIGURES 321 326" captionStartId="72.[264,383,1696,1720]" captionTargetBox="[286,1297,660,1656]" captionTargetId="figure@72.[275,1311,649,1670]" captionTargetPageId="72" captionText="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="73" pageNumber="74">Figs. 323324</figureCitation>
|
||
,
|
||
<figureCitation id="4BC24485FFD7396A486CFED0FB01FE5C" box="[1097,1257,364,390]" captionStart="FIGURES 310 315" captionStartId="69.[264,383,1724,1748]" captionTargetBox="[290,1301,520,1688]" captionTargetId="figure@69.[277,1311,510,1700]" captionTargetPageId="69" captionText="FIGURES 310 315. Aedeagus of Geostiba impressula (Casey) (lectotype of Sipalia fossata Casey (310 311) and a specimen from Triangle Lake, Oregon (312 315 )). 310, 312 – median lobe, ventral view; 311, 315 – apex of median lobe, ventral view; 313 – median lobe, lateral view; 314 – apex of median lobe, lateral view. Scale bar 0.4 mm (310, 312 313), 0.2 mm (311, 314 315)." httpUri="https://zenodo.org/record/155744/files/figure.png" pageId="73" pageNumber="74">310312, 315</figureCitation>
|
||
), elytral suture slightly raised behind scutellum, and in males lacking the medial pronotal impression.
|
||
</paragraph>
|
||
<paragraph id="D3465800FFD7396A4D1DFE5FFD27FDFC" blockId="73.[264,1324,284,750]" pageId="73" pageNumber="74">
|
||
<emphasis id="E18D8412FFD7396A4D1DFE5FFE38FE27" bold="true" box="[312,464,483,509]" pageId="73" pageNumber="74">Description.</emphasis>
|
||
Length
|
||
<quantity id="1401F5E5FFD7396A4E67FE58FD4AFE24" box="[578,674,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="73" pageNumber="74" unit="mm" value="2.4">2.4 mm</quantity>
|
||
. Body reddish brown, antennae, legs and mouthparts brownish yellow. Body parallelsided.
|
||
</paragraph>
|
||
<paragraph id="D3465800FFD7396A4D1DFD88FD53FD34" blockId="73.[264,1324,284,750]" pageId="73" pageNumber="74">
|
||
Head as wide as long, surface on disk with fine isodiametric microsculpture, puncturation very fine and weak, distance between punctures equal to 23 times their diameter. Temple length to eye length ratio 2.3. Antennal article 2 longer than article 3, article 4 transverse (width to length ratio 1.6), articles 510 strongly transverse, last article as long as 9 and 10 combined (as in
|
||
<figureCitation id="4BC24485FFD7396A4E72FD68FD45FD34" box="[599,685,724,750]" captionStart="FIGURES 8 17" captionStartId="6.[264,383,1709,1733]" captionTargetBox="[307,1277,321,1687]" captionTargetId="figure@6.[297,1289,310,1697]" captionTargetPageId="6" captionText="FIGURES 8 17. Mouthparts of Geostiba circellaris (Gravenhorst) (8, Moscow Region, Russia), G. infirma (Weise) (9, IvanoFrankovsk Region, Ukraine), G. carteriensis Pace (10 15, Tennessee), G. appalachigena Gusarov, sp. n. (16, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (17, paratype from North Carolina). 8 10, – prementum; 11 – hypopharynx; 12 – left maxilla, ventral view; 13 – left galea, dorsal view; 14 – left lacinia, dorsal view; 15 – mentum; 16 17 – right antenna. Scale bar 0.1 mm (8 15), 0.2 mm (16 17). L – Length of antennal article." httpUri="https://zenodo.org/record/155703/files/figure.png" pageId="73" pageNumber="74">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="87860888FFD7396A4D2DFA74FAF1F957" httpUri="https://zenodo.org/record/155747/files/figure.png" pageId="73" pageNumber="74" targetBox="[307,1280,824,1441]" targetPageId="73">
|
||
<paragraph id="D3465800FFD7396A4D2DFA74FAF1F957" blockId="73.[264,1323,1480,1677]" pageId="73" pageNumber="74">
|
||
<emphasis id="E18D8412FFD7396A4D2DFA74FE08FA3A" bold="true" box="[264,480,1480,1504]" pageId="73" pageNumber="74">FIGURES 327335</emphasis>
|
||
. Genitalia of
|
||
<taxonomicName id="14F92383FFD7396A4E5BFA75FC43FA3A" authority="Gusarov" authorityName="Gusarov" box="[638,939,1481,1504]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="silvigena" status="sp. nov.">
|
||
<emphasis id="E18D8412FFD7396A4E5BFA75FCAEFA3A" box="[638,838,1481,1504]" italics="true" pageId="73" pageNumber="74">Geostiba silvigena</emphasis>
|
||
Gusarov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FABE3969FFD7396A4F93FA74FC1BFA3A" box="[950,1011,1480,1504]" pageId="73" pageNumber="74" rank="species">
|
||
<emphasis id="E18D8412FFD7396A4F93FA74FC1BFA3A" bold="true" box="[950,1011,1480,1504]" pageId="73" pageNumber="74">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype). 327 – details of internal sac retracted into median lobe, ventral view; 328 – copulatory piece of internal sac, dorsal view; 329 – distal diverticula of everted internal sac, ventral view; 330 – everted internal sac, ventral view; 331 – everted internal sac, lateral view; 332333 – details of internal sac retracted into median lobe, lateral view; 334 – apex of left paramere, side facing median lobe; 335 – copulatory piece of internal sac, lateral view. Scale bar 0.4 mm (327, 330333), 0.2 mm (328329, 334335).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D3465800FFD739694D1DF955FE2FFE74" blockId="73.[264,1324,1769,1875]" lastBlockId="74.[264,1324,284,1310]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
|
||
Pronotum slightly transverse, width
|
||
<quantity id="1401F5E5FFD7396A4ED4F955FC88F8D9" box="[753,864,1769,1795]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="73" pageNumber="74" unit="mm" value="0.41">0.41 mm</quantity>
|
||
, width to length ratio 1.0, wider than head (pronotal width to head width ratio 1.1); microsculpture and puncturation as on head. Pronotal pubescence of
|
||
<typeStatus id="0C42E6A2FFD7396A4E02F885FDB2F889" box="[551,602,1849,1875]" pageId="73" pageNumber="74">type</typeStatus>
|
||
V. Elytra measured from humeral angle longer than pronotum (pronotal length to elytral length ratio 1.0), wider than long (1.3), with weak and fine isodiametric microsculpture and fine, somewhat asperate puncturation, distance between punctures equals 23 times their diameter. Elytral suture behind scutellum raised. Wings fully developed.
|
||
</paragraph>
|
||
<paragraph id="D3465800FFD439694D1DFE00FB4DFDFC" blockId="74.[264,1324,284,1310]" pageId="74" pageNumber="75">Abdominal terga with fine microsculpture of transverse meshes, with fine and sparse puncturation, puncturation becoming finer towards abdomen apex, on terga 35 distance between punctures equals 35 times their diameter. Tergum 7 with white edge.</paragraph>
|
||
<paragraph id="D3465800FFD439694D1DFD88FC19FD44" blockId="74.[264,1324,284,1310]" pageId="74" pageNumber="75">
|
||
Male tergum 7 with two medial carinae in front of posterior margin. Male tergum 8 with four small carinae in front of posterior margin, posterior margin slightly convex (
|
||
<figureCitation id="4BC24485FFD4396948DEFDE0FED0FD44" captionStart="FIGURES 321 326" captionStartId="72.[264,383,1696,1720]" captionTargetBox="[286,1297,660,1656]" captionTargetId="figure@72.[275,1311,649,1670]" captionTargetPageId="72" captionText="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="74" pageNumber="75">Fig. 321</figureCitation>
|
||
). Male sternum 8 with convex posterior margin (
|
||
<figureCitation id="4BC24485FFD439694F58FD38FC0DFD44" box="[893,997,644,670]" captionStart="FIGURES 321 326" captionStartId="72.[264,383,1696,1720]" captionTargetBox="[286,1297,660,1656]" captionTargetId="figure@72.[275,1311,649,1670]" captionTargetPageId="72" captionText="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="74" pageNumber="75">Fig. 322</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFD439694D1DFD10FB14FD34" blockId="74.[264,1324,284,1310]" pageId="74" pageNumber="75">
|
||
Aedeagus as in
|
||
<figureCitation id="4BC24485FFD439694DDEFD10FD4EFD1C" box="[507,678,684,710]" captionStart-0="FIGURES 321 326" captionStart-1="FIGURES 327 335" captionStartId-0="72.[264,383,1696,1720]" captionStartId-1="73.[264,383,1480,1504]" captionTargetBox-0="[286,1297,660,1656]" captionTargetBox-1="[307,1280,824,1441]" captionTargetId-0="figure@72.[275,1311,649,1670]" captionTargetId-1="figure@73.[296,1290,809,1454]" captionTargetPageId-0="72" captionTargetPageId-1="73" captionText-0="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." captionText-1="FIGURES 327 335. Genitalia of Geostiba silvigena Gusarov, sp. n. (holotype). 327 – details of internal sac retracted into median lobe, ventral view; 328 – copulatory piece of internal sac, dorsal view; 329 – distal diverticula of everted internal sac, ventral view; 330 – everted internal sac, ventral view; 331 – everted internal sac, lateral view; 332 333 – details of internal sac retracted into median lobe, lateral view; 334 – apex of left paramere, side facing median lobe; 335 – copulatory piece of internal sac, lateral view. Scale bar 0.4 mm (327, 330 333), 0.2 mm (328 329, 334 335)." httpUri-0="https://zenodo.org/record/155746/files/figure.png" httpUri-1="https://zenodo.org/record/155747/files/figure.png" pageId="74" pageNumber="75">Figs. 323335</figureCitation>
|
||
. Apex of median lobe in lateral view bent ventrally (
|
||
<figureCitation id="4BC24485FFD439694D34FD68FE51FD34" box="[273,441,724,750]" captionStart="FIGURES 321 326" captionStartId="72.[264,383,1696,1720]" captionTargetBox="[286,1297,660,1656]" captionTargetId="figure@72.[275,1311,649,1670]" captionTargetPageId="72" captionText="FIGURES 321 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 323, 325), 0.2 mm (324, 326)." httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="74" pageNumber="75">Figs. 325326</figureCitation>
|
||
), distal diverticula of internal sac in ventral view wide (
|
||
<figureCitation id="4BC24485FFD439694863FD68FB07FD34" box="[1094,1263,724,750]" captionStart="FIGURES 327 335" captionStartId="73.[264,383,1480,1504]" captionTargetBox="[307,1280,824,1441]" captionTargetId="figure@73.[296,1290,809,1454]" captionTargetPageId="73" captionText="FIGURES 327 335. Genitalia of Geostiba silvigena Gusarov, sp. n. (holotype). 327 – details of internal sac retracted into median lobe, ventral view; 328 – copulatory piece of internal sac, dorsal view; 329 – distal diverticula of everted internal sac, ventral view; 330 – everted internal sac, ventral view; 331 – everted internal sac, lateral view; 332 333 – details of internal sac retracted into median lobe, lateral view; 334 – apex of left paramere, side facing median lobe; 335 – copulatory piece of internal sac, lateral view. Scale bar 0.4 mm (327, 330 333), 0.2 mm (328 329, 334 335)." httpUri="https://zenodo.org/record/155747/files/figure.png" pageId="74" pageNumber="75">Figs. 329330</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFD439694D1DFD40FDE3FCCC" blockId="74.[264,1324,284,1310]" box="[312,523,764,790]" pageId="74" pageNumber="75">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFD439694D1DFC9FFC0DFBF4" pageId="74" pageNumber="75" type="discussion">
|
||
<paragraph id="D3465800FFD439694D1DFC9FFC0DFBF4" blockId="74.[264,1324,284,1310]" pageId="74" pageNumber="75">
|
||
<emphasis id="E18D8412FFD439694D1DFC9FFE2BFCE7" bold="true" box="[312,451,803,829]" pageId="74" pageNumber="75">Discussion.</emphasis>
|
||
<taxonomicName id="14F92383FFD439694DFDFC98FD5EFCE7" box="[472,694,804,829]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="silvigena">
|
||
<emphasis id="E18D8412FFD439694DFDFC98FD5EFCE7" box="[472,694,804,829]" italics="true" pageId="74" pageNumber="75">Geostiba silvigena</emphasis>
|
||
</taxonomicName>
|
||
has fully developed wings and it is probably widespread along the Pacific coast of North
|
||
<collectingCountry id="ABEE1890FFD439694EF7FCF0FCD3FCBC" box="[722,827,844,870]" name="United States of America" pageId="74" pageNumber="75">America</collectingCountry>
|
||
. The species may not be active in summer when most collections in the area have been made (see the above discussion of natural history of
|
||
<taxonomicName id="14F92383FFD439694D7AFC20FDEBFC6F" box="[351,515,924,949]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="impressula">
|
||
<emphasis id="E18D8412FFD439694D7AFC20FDEBFC6F" box="[351,515,924,949]" italics="true" pageId="74" pageNumber="75">G. impressula</emphasis>
|
||
</taxonomicName>
|
||
). Although
|
||
<taxonomicName id="14F92383FFD439694EBFFC20FCC1FC6F" box="[666,809,924,949]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="silvigena">
|
||
<emphasis id="E18D8412FFD439694EBFFC20FCC1FC6F" box="[666,809,924,949]" italics="true" pageId="74" pageNumber="75">G. silvigena</emphasis>
|
||
</taxonomicName>
|
||
is closely related to the eastern Nearctic
|
||
<taxonomicName id="14F92383FFD43969492BFC20FE5EFC07" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="appalachigena">
|
||
<emphasis id="E18D8412FFD43969492BFC20FE5EFC07" italics="true" pageId="74" pageNumber="75">G. appalachigena</emphasis>
|
||
</taxonomicName>
|
||
, the westernmost locality of
|
||
<taxonomicName id="14F92383FFD439694F35FC78FC0BFC07" box="[784,995,964,989]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="appalachigena">
|
||
<emphasis id="E18D8412FFD439694F35FC78FC0BFC07" box="[784,995,964,989]" italics="true" pageId="74" pageNumber="75">G. appalachigena</emphasis>
|
||
</taxonomicName>
|
||
is in Wisconsin (
|
||
<figureCitation id="4BC24485FFD439694897FC78FED0FBDC" captionStart-0="FIGURE 336" captionStart-1="FIGURE 337" captionStartId-0="76.[264,368,1562,1586]" captionStartId-1="77.[264,367,1248,1272]" captionTargetBox-0="[315,1271,753,1543]" captionTargetBox-1="[328,1262,539,1226]" captionTargetId-0="figure@76.[311,1275,751,1546]" captionTargetId-1="figure@77.[321,1266,529,1233]" captionTargetPageId-0="76" captionTargetPageId-1="77" captionText-0="FIGURE 336. Geographical distribution of Nearctic species of Geostiba. The ellipse indicates the area where eleven Southern Appalachian species are distributed." captionText-1="FIGURE 337. Geographical distribution of Geostiba appalachigena Gusarov, sp. n. The ellipse indicates the area where eleven Southern Appalachian species are distributed." httpUri-0="https://zenodo.org/record/155748/files/figure.png" httpUri-1="https://zenodo.org/record/155749/files/figure.png" pageId="74" pageNumber="75">Figs. 336 337</figureCitation>
|
||
). Considering the gap between the ranges of the two species and the minor difference in male genitalia the separate status of the two seems justified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFD439694D1DFB87FD50FB7C" pageId="74" pageNumber="75" type="distribution">
|
||
<paragraph id="D3465800FFD439694D1DFB87FAE7FB8C" blockId="74.[264,1324,284,1310]" box="[312,1295,1083,1110]" pageId="74" pageNumber="75">
|
||
<emphasis id="E18D8412FFD439694D1DFB87FD7FFB8F" bold="true" box="[312,663,1083,1109]" pageId="74" pageNumber="75">Distribution and variability.</emphasis>
|
||
Known only from Mendocino, California (
|
||
<figureCitation id="4BC24485FFD4396948BEFB80FAEAFB8C" box="[1179,1282,1084,1110]" captionStart="FIGURE 336" captionStartId="76.[264,368,1562,1586]" captionTargetBox="[315,1271,753,1543]" captionTargetId="figure@76.[311,1275,751,1546]" captionTargetPageId="76" captionText="FIGURE 336. Geographical distribution of Nearctic species of Geostiba. The ellipse indicates the area where eleven Southern Appalachian species are distributed." httpUri="https://zenodo.org/record/155748/files/figure.png" pageId="74" pageNumber="75">Fig. 336</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D3465800FFD439694D1DFBDFFD50FB7C" blockId="74.[264,1324,284,1310]" pageId="74" pageNumber="75">
|
||
<emphasis id="E18D8412FFD439694D1DFBDFFDEDFBA7" bold="true" box="[312,517,1123,1149]" pageId="74" pageNumber="75">Natural History.</emphasis>
|
||
The only known specimen of
|
||
<taxonomicName id="14F92383FFD439694F51FBD8FBEAFBA7" box="[884,1026,1124,1149]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="silvigena">
|
||
<emphasis id="E18D8412FFD439694F51FBD8FBEAFBA7" box="[884,1026,1124,1149]" italics="true" pageId="74" pageNumber="75">G. silvigena</emphasis>
|
||
</taxonomicName>
|
||
was collected in moss in the bed of a dried up woodland pool.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BE30B8BFFD439694D1DFB0FFE05FAC4" pageId="74" pageNumber="75" type="etymology">
|
||
<paragraph id="D3465800FFD439694D1DFB0FFE05FAC4" blockId="74.[264,1324,284,1310]" pageId="74" pageNumber="75">
|
||
<emphasis id="E18D8412FFD439694D1DFB0FFE2DFB17" bold="true" box="[312,453,1203,1229]" pageId="74" pageNumber="75">Etymology.</emphasis>
|
||
The specific name is derived from the Latin noun
|
||
<emphasis id="E18D8412FFD439694865FB08FB9FFB17" box="[1088,1143,1204,1229]" italics="true" pageId="74" pageNumber="75">silva</emphasis>
|
||
(forest, woodland) and the verb
|
||
<emphasis id="E18D8412FFD439694DC9FB60FDC7FB2F" box="[492,559,1244,1269]" italics="true" pageId="74" pageNumber="75">gigno</emphasis>
|
||
(to be born, to arise). It refers to the habitat where the species presumably occurs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |