182 lines
17 KiB
XML
182 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/caucasiana.2.e101013" ID-Pensoft-Pub="2667-9809-2-29" ID-Pensoft-UUID="18D2727FF82056D4A6013DA6261236BA" ID-ZooBank="5DC5FF4391E7404F9F7B18274382D72B" ModsDocID="2667-9809-2-29" checkinTime="1679617027075" checkinUser="pensoft" docAuthor="Mumladze, Levan, Grego, Jozef & Szekeres, Miklos" docDate="2023" docId="59B517B5838A53E4A8C97D46D143057D" docLanguage="en" docName="Caucasiana 2: 29-61" docOrigin="Caucasiana 2" docPubDate="2023-03-22" docSource="http://dx.doi.org/10.3897/caucasiana.2.e101013" docTitle="Mucronaria (Mucronaria) kartvelica Mumladze & Grego & Szekeres 2023, sp. n." docType="treatment" docUuid="0A0DEA45-B225-4BFD-AD89-3A26A6D0A025" docUuidSource="ZooBank" docVersion="1" id="18D2727FF82056D4A6013DA6261236BA" lastPageNumber="29" masterDocId="18D2727FF82056D4A6013DA6261236BA" masterDocTitle="The land snail family Clausiliidae (Gastropoda, Pulmonata, Stylommatophora) in Georgia: overview, novel records and a new species" masterLastPageNumber="61" masterPageNumber="29" pageNumber="29" updateTime="1679617027075" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The land snail family Clausiliidae (Gastropoda, Pulmonata, Stylommatophora) in Georgia: overview, novel records and a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mumladze, Levan</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
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<mods:affiliation>Institute of Zoology, Ilia State University, K. Cholokashvilli Ave 3 / 5, Tbilisi, Georgia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grego, Jozef</mods:namePart>
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<mods:affiliation>Molcanska cesta 219, Horna Micina, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szekeres, Miklos</mods:namePart>
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<mods:affiliation>Institute of Plant Biology, Biological Research Centre HAS, Temesvari krt. 62, Szeged, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">szekeres.miklos@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Caucasiana</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>61</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/caucasiana.2.e101013</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/caucasiana.2.e101013</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2667-9809-2-29</mods:identifier>
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<mods:identifier type="ZooBank">5DC5FF4391E7404F9F7B18274382D72B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">18D2727FF82056D4A6013DA6261236BA</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:0A0DEA45-B225-4BFD-AD89-3A26A6D0A025" httpUri="http://treatment.plazi.org/id/59B517B5838A53E4A8C97D46D143057D" lastPageNumber="29" pageId="0" pageNumber="29">
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<subSubSection pageId="0" pageNumber="29" type="nomenclature">
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName LSID="https://zoobank.org/0A0DEA45-B225-4BFD-AD89-3A26A6D0A025" authority="Mumladze & Grego & Szekeres, 2023" authorityName="Mumladze & Grego & Szekeres" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Mucronaria (Mucronaria) kartvelica" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="kartvelica" status="sp. n." subGenus="Mucronaria">Mucronaria (Mucronaria) kartvelica</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="29">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mucronaria (Mucronaria) kartvelica sp. n., holotype (ISUTM-T 003 - H). Scale bar: 3 mm." figureDoi="10.3897/caucasiana.2.e101013.figure10" httpUri="https://binary.pensoft.net/fig/828423" pageId="0" pageNumber="29">Figures 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. The habitat (A) and a live individual (B) of Mucronaria (Mucronaria) kartvelica sp. n." figureDoi="10.3897/caucasiana.2.e101013.figure11" httpUri="https://binary.pensoft.net/fig/828424" pageId="0" pageNumber="29">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Occurrence records of Mucronaria (Mucronaria) kartvelica sp. n. (A), Mucronaria (M.) strauchi (B) and Mucronaria (Index) index (C). Symbols are as in Figure 2." figureDoi="10.3897/caucasiana.2.e101013.figure12" httpUri="https://binary.pensoft.net/fig/828425" pageId="0" pageNumber="29">, 12A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="29">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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The reduced, diffuse palatal plicae and the less elongate shell shape distinguish the new species from members of the subgenus
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<taxonomicName authority="" authorityName="index (Mousson" authorityYear="1863" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (Index)" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="subGenus" subGenus="Index">Mucronaria (Index)</taxonomicName>
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Boettger, 1877 and position it in the subgenus
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<taxonomicName authority="(Mucronaria)" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (Mucronaria)" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="subGenus" subGenus="Mucronaria">Mucronaria (Mucronaria)</taxonomicName>
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Boettger, 1877.
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<taxonomicName authorityName="Mumladze & Grego & Szekeres" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) kartvelica" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="kartvelica" subGenus="M.">Mucronaria (M.) kartvelica</taxonomicName>
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sp. n. differs from all other species of its subgenus by the slender and strongly costate shell. By contrast,
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<taxonomicName baseAuthorityName="Mousson" baseAuthorityYear="1876" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) acuminata" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="acuminata" subGenus="M.">Mucronaria (M.) acuminata</taxonomicName>
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has ventricose shell with glossy, densely striate surface and very weak lamella superior,
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<taxonomicName baseAuthorityName="Charpentier" baseAuthorityYear="1852" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) duboisi" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="duboisi" subGenus="M.">Mucronaria (M.) duboisi</taxonomicName>
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has strong, to the peristome margin extended lamella inferior and long, well outlined palatal plicae, whereas
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<taxonomicName baseAuthorityName="Boettger" baseAuthorityYear="1878" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) strauchi" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="strauchi" subGenus="M.">Mucronaria (M.) strauchi</taxonomicName>
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(Boettger, 1878) is less elongate, more densely costate, and its ribs are more widely spaced on the neck than on the rest of the last whorl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="type locality">
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<paragraph pageId="0" pageNumber="29">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Georgia, Racha-Lechkhumi and Kvemo Svaneti Region, at the Kvedi 1 Cave 2.2 km ESE of Kvedi village,
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<geoCoordinate degrees="42.5561" direction="north" orientation="latitude" precision="5" value="42.5561">N42.5561°</geoCoordinate>
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,
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<geoCoordinate degrees="43.5721" direction="east" orientation="longitude" precision="5" value="43.5721">E43.5721°</geoCoordinate>
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; 1510 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="type material">
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<paragraph pageId="0" pageNumber="29">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="29">Type locality; 14.10.2021; leg. JG-MS, holotype (ISUTM-T003-H); paratypes (FMNH 392791/1, HNHM 105342/1, MNHN-IM-2014-7037/1, NHMUK 20220496/1, NHMW-MO-113737/1, SMF 368736/1, UF 580908/1, ZMH-MOL-0141473/1, JG/4, MS/3). Georgia, Racha-Lechkhumi and Kvemo Svaneti Region, at the Kvedi 2 Cave, 2 km ESE of Kvedi village, 42.5561°, E43.5691°; 1560 m; 14.10.2021; leg. JG-MS, paratypes (ISUTM-T003-P1/1, SMF 368737/1, JG/15, MS/9).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="materials_examined">
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<paragraph pageId="0" pageNumber="29">Further material.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<materialsCitation collectingDate="2022-09-12" country="Georgia" elevation="1560" location="Cave" longitude="43.5691" municipality="Kvedi" specimenCount="1" stateProvince="Racha-Lechkhumi and Kvemo Svaneti Region">
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<collectingCountry name="Georgia">Georgia</collectingCountry>
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,
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<collectingRegion country="Georgia" name="Racha-Lechkhumi and Kvemo Svaneti">Racha-Lechkhumi and Kvemo Svaneti Region</collectingRegion>
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, at the
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<collectingMunicipality>Kvedi</collectingMunicipality>
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2
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<location LSID="urn:lsid:plazi:treatment:59B517B5838A53E4A8C97D46D143057D:EA47AEE61A48EA8A44938F2F74D2538E" country="Georgia" longitude="43.5691" municipality="Kvedi" name="Cave" stateProvince="Racha-Lechkhumi and Kvemo Svaneti Region">Cave</location>
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<locationDeviation location="Kvedi village">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
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ESE of Kvedi village
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</locationDeviation>
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, 42.5561°,
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<geoCoordinate degrees="43.5691" direction="east" orientation="longitude" precision="5" value="43.5691">E43.5691°</geoCoordinate>
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;
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.56" unit="m" value="1560.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.56" unit="m" value="1560.0">1560 m</elevation>
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</quantity>
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; leg. EC-JG-LM-MO-MS
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<collectingDate value="2022-09-12">12.09.2022</collectingDate>
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(ISU, JG, MS)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">Description.</paragraph>
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<paragraph pageId="0" pageNumber="29">The brownish-corneous, slender spindle-shaped shell consists of 12.3 to 13.5 whorls. The apical part is elongate, its tip is pointed. The teleochonch whorls have strong and sharp ribs (25 to 28 on the last whorl), which become bulkier but not more widely spaced behind the aperture. The last whorl becomes narrower toward the base, which has a prominent basal crest. The relatively small aperture is oval to somewhat diamond-shaped, its wide peristome is slightly projected. The thin, weakly emerged lamella superior reaches the margin of the peristome. Inward it converges slightly toward the lamella inferior and terminates well before reaching the lamella spiralis, which starts ventrally and ends dorsally. The lamella inferior ends high and deep, its terminal part is only barely visible in apertural view of the shell. Ending even deeper, the lamella subcolumellaris cannot be viewed through the aperture. The plica principalis spans one third of the last whorl starting from the lateral side. The well discernable, dorsolaterally positioned upper plica runs close and parallel to it. Farther toward the base there are two or three lump-like reduced plicae, and a stronger and larger but also diffuse, callus-like basal plica. The broad clausilium plate widens before ending abruptly in a short, pointed tip. In some of the specimens its parietal margin is visible in oblique view through the aperture.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">Dimensions</paragraph>
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<paragraph pageId="0" pageNumber="29">(in mm): Shell height: 12.4-13.9 (holotype 12.9), spire width: 2.8-3.1 (holotype 2.9), aperture height: 2.4-2.6 (holotype 2.5), aperture width: 1.9-2.1 (holotype 2.0).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="anatomy">
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<paragraph pageId="0" pageNumber="29">Anatomy.</paragraph>
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<paragraph pageId="0" pageNumber="29">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="habitat">
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<paragraph pageId="0" pageNumber="29">Habitat.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Live individuals of
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<taxonomicName authorityName="Mumladze & Grego & Szekeres" authorityYear="2023" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) kartvelica" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="kartvelica" subGenus="M.">Mucronaria (M.) kartvelica</taxonomicName>
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sp. n. were found on shaded limestone cliffs and among weeds (primarily
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<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Hedera" kingdom="Plantae" lsidName="Hedera helix" order="Apiales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="helix">
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<emphasis italics="true" pageId="0" pageNumber="29">Hedera helix</emphasis>
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</taxonomicName>
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) growing on those cliffs (Figure
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. The habitat (A) and a live individual (B) of Mucronaria (Mucronaria) kartvelica sp. n." figureDoi="10.3897/caucasiana.2.e101013.figure11" httpUri="https://binary.pensoft.net/fig/828424" pageId="0" pageNumber="29">11A-B</figureCitation>
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), accompanied by
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<taxonomicName authorityName="Likharev & Schileyko" authorityYear="2007" class="Gastropoda" family="Clausiliidae" genus="Acrotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrotoma (Iliamneme) baryshnikovi" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="baryshnikovi" subGenus="Iliamneme">Acrotoma (Iliamneme) baryshnikovi</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Mousson" baseAuthorityYear="1863" class="Gastropoda" family="Clausiliidae" genus="Elia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elia (Megaleuxina) derasa" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="derasa" subGenus="Megaleuxina">Elia (Megaleuxina) derasa</taxonomicName>
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,
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<taxonomicName baseAuthorityName="Mortillet" baseAuthorityYear="1854" class="Gastropoda" family="Clausiliidae" genus="Mentissoidea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mentissoidea rupicola" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="rupicola">
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<emphasis italics="true" pageId="0" pageNumber="29">Mentissoidea rupicola</emphasis>
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</taxonomicName>
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and
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<taxonomicName baseAuthorityName="Charpentier" baseAuthorityYear="1852" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) duboisi" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="duboisi" subGenus="M.">Mucronaria (M.) duboisi</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="etymology">
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<paragraph pageId="0" pageNumber="29">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="29">The name of the new species refers to its occurrence in Sa-kartvelo (native historic name for Georgia).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="remarks">
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<paragraph pageId="0" pageNumber="29">Remark.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Whereas
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<taxonomicName baseAuthorityName="Charpentier" baseAuthorityYear="1852" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (M.) duboisi" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="duboisi" subGenus="M.">Mucronaria (M.) duboisi</taxonomicName>
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is one of the widest distributed clausiliids in the Caucasus region, reaching as far as Turkey's Sinop Province toward west and northern Iran toward east, all the other species of the subgenus
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<taxonomicName authority="(Mucronaria)" class="Gastropoda" family="Clausiliidae" genus="Mucronaria" kingdom="Animalia" lsidName="Mucronaria (Mucronaria)" order="Stylommatophora" pageId="0" pageNumber="29" phylum="Mollusca" rank="subGenus" subGenus="Mucronaria">Mucronaria (Mucronaria)</taxonomicName>
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are endemic to Georgia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |