212 lines
39 KiB
XML
212 lines
39 KiB
XML
<document id="9E5C8F8013CDA85E82545BCD6AA962B5" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFFEFFECBE64F8873A70F8D1" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius linang Peter J. Schwendinger 2017" docType="treatment" docVersion="7" lastPageNumber="418" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="417" updateTime="1698469770161" updateUser="ExternalLinkService">
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<mods:title id="A64FFAA447D1FAF0E2C9503BC6241751">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
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<mods:name id="D6D1485B922DEFD45BCC5AB497E9EAC0" type="personal">
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<mods:namePart id="10ADB7331C5E9C9211494D8601266709">Peter J. Schwendinger</mods:namePart>
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<mods:typeOfResource id="830BCB89A8C101825755B62DD12A226F">text</mods:typeOfResource>
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<mods:title id="98F5998098F3792CB94BFF429832E9F2">Revue suisse de Zoologie</mods:title>
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<mods:part id="9962FDF81BC73A7D776732C6AED34A5E">
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<mods:date id="7494376869D9AC6B77398467FE0195C2">2017</mods:date>
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<mods:number id="568843F675171CACB8BC6DF2E673E0DC">2017-09-30</mods:number>
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<mods:number id="682283ECBC10DEB389870D8B1BA5F8AA">124</mods:number>
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<treatment id="4C30A452FFFEFFECBE64F8873A70F8D1" ID-DOI="http://doi.org/10.5281/zenodo.6042370" ID-GBIF-Taxon="135450768" ID-Zenodo-Dep="6042370" LSID="urn:lsid:plazi:treatment:4C30A452FFFEFFECBE64F8873A70F8D1" httpUri="http://treatment.plazi.org/id/4C30A452FFFEFFECBE64F8873A70F8D1" lastPageId="27" lastPageNumber="418" pageId="26" pageNumber="417">
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<subSubSection id="8C8346CFFFFEFFEDBE64F8873CC9F86C" box="[1035,1233,1910,1935]" pageId="26" pageNumber="417" type="nomenclature">
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<paragraph id="C4261544FFFEFFEDBE64F8873CC9F86C" blockId="26.[1035,1233,1910,1935]" box="[1035,1233,1910,1935]" pageId="26" pageNumber="417">
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<heading id="9F6EA228FFFEFFEDBE64F8873CC9F86C" bold="true" box="[1035,1233,1910,1935]" centered="true" fontSize="10" level="3" pageId="26" pageNumber="417" reason="2">
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<emphasis id="F6EDC956FFFEFFEDBE64F8873CC9F86C" bold="true" box="[1035,1233,1910,1935]" pageId="26" pageNumber="417">
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<emphasis id="F6EDC956FFFEFFEDBE64F8873C9CF86E" bold="true" box="[1035,1156,1910,1933]" italics="true" pageId="26" pageNumber="417">
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The
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<taxonomicName id="03996EC7FFFEFFEDBE52F8873C9CF86E" ID-CoL="6QJ5F" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1085,1156,1910,1933]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="417" phylum="Arthropoda" rank="species" species="linang">linang</taxonomicName>
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</emphasis>
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-group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="8C8346CFFFFEFFECB950F85438F2FAFF" lastPageId="27" lastPageNumber="418" pageId="26" pageNumber="417" type="diagnosis">
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<paragraph id="C4261544FFFEFFECB950F8543A54FEF9" blockId="26.[831,1437,1957,2078]" lastBlockId="27.[151,588,257,282]" lastPageId="27" lastPageNumber="418" pageId="26" pageNumber="417">
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<emphasis id="F6EDC956FFFEFFEDB950F8543BAEF85F" bold="true" box="[831,950,1957,1980]" pageId="26" pageNumber="417">Diagnosis:</emphasis>
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Small to medium-sized spiders (carapace length of males 4.50-6.83, carapace width 3.98-6.21) with a colour pattern on carapace, limbs and opisthosomal tergites (
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<figureCitation id="5CA209C1FFFEFFEDB9CEF7F73BE3F7FD" box="[929,1019,2054,2078]" captionStart="Fig. 2" captionStartId="3.[149,184,881,902]" captionTargetBox="[166,1405,257,835]" captionTargetId="figure@3.[149,1420,257,835]" captionTargetPageId="3" captionText="Fig. 2. Habitus of two Liphistius species from peninsular Malaysia and southern Thailand. (A) Liphistius endau, female from Kota Tinggi (Malaysia), dorsal view. (B) Liphistius indra sp. nov., male paratype from the Sankalakhierie Mountains (Thailand), same view." httpUri="https://zenodo.org/record/1021941/files/figure.png" pageId="26" pageNumber="417">Fig. 2B</figureCitation>
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). Distinguished from the
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<taxonomicName id="03996EC7FFFEFFEDBF4FF7F73D8CF7FE" authorityName="Abraham" authorityYear="1923" box="[1312,1428,2054,2077]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="417" phylum="Arthropoda" rank="species" species="malayanus">
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<emphasis id="F6EDC956FFFEFFEDBF4FF7F73D8CF7FE" box="[1312,1428,2054,2077]" italics="true" pageId="26" pageNumber="417">malayanus</emphasis>
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</taxonomicName>
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-
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<emphasis id="F6EDC956FFFFFFECBAF8FEF33A54FEF9" bold="true" box="[151,588,257,282]" pageId="27" pageNumber="418">
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Key to the species of the
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<taxonomicName id="03996EC7FFFFFFECBBC0FEF039EEFEFB" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[431,502,257,280]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang">
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<emphasis id="F6EDC956FFFFFFECBBC0FEF039EEFEFB" bold="true" box="[431,502,257,280]" italics="true" pageId="27" pageNumber="418">linang</emphasis>
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</taxonomicName>
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-group:
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</emphasis>
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</paragraph>
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<paragraph id="C4261544FFFFFFECBAF8FEB33D8CFE3B" blockId="27.[151,1437,321,634]" pageId="27" pageNumber="418">
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<specimenCount id="D29FDECDFFFFFFECBAF8FEB3393CFEBB" box="[151,292,321,346]" pageId="27" pageNumber="418" type="male">1 Males</specimenCount>
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without scopula on tarsus IV; long bristle retroventrally at base of tibial apophysis thin (
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<figureCitation id="5CA209C1FFFFFFECBE9AFEB03D42FEBB" box="[1269,1370,321,344]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15A</figureCitation>
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, G-I); paracymbium quite long (
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<figureCitation id="5CA209C1FFFFFFECBB9BFE903A4EFE9B" box="[500,598,353,376]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15A</figureCitation>
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); distal edge of contrategulum with wide gap behind distinctly bent dorsal apex (
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<figureCitation id="5CA209C1FFFFFFECBA84FE703956FE7B" box="[235,334,385,408]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15B</figureCitation>
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).
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<location id="C146439FFFFFFFECBB0DFE7039DFFE7B" LSID="urn:lsid:plazi:treatment:4C30A452FFFEFFECBE64F8873A70F8D1:C146439FFFFFFFECBB0DFE7039DFFE7B" box="[354,455,385,408]" country="Malaysia" name="Poreplate" pageId="27" pageNumber="418">Poreplate</location>
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with anterior margin distinctly invaginated (
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<figureCitation id="5CA209C1FFFFFFECB9C5FE703BEFFE7B" box="[938,1015,385,408]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="27" pageNumber="418">Fig. 16</figureCitation>
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A-E); CDO a transverse slit or triangle without enlarged pores lateral to it (
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<figureCitation id="5CA209C1FFFFFFECB831FE503AD8FE5B" box="[606,704,417,440]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="27" pageNumber="418">Fig. 16A</figureCitation>
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, C-D); receptacular cluster indistinctly divided into lateral halves, its vesicles quite short (
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<figureCitation id="5CA209C1FFFFFFECBBD1FE303A22FE39" box="[446,570,449,474]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="27" pageNumber="418">Fig. 16B, E</figureCitation>
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) ............................................................................
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<taxonomicName id="03996EC7FFFFFFECBE29FE323CBEFE3B" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1094,1190,449,474]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
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<emphasis id="F6EDC956FFFFFFECBE29FE323CBEFE3B" box="[1094,1190,449,474]" italics="true" pageId="27" pageNumber="418">L. linang</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EDDE742DFFFFFFECBEC2FE323D18FE39" box="[1197,1280,451,474]" pageId="27" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
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(in
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<collectingCountry id="BC8E55D4FFFFFFECBF45FE303D95FE3B" box="[1322,1421,449,472]" name="Malaysia" pageId="27" pageNumber="418">Malaysia</collectingCountry>
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)
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</paragraph>
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<paragraph id="C4261544FFFFFFECBAF8FE123D8CFD9B" blockId="27.[151,1437,321,634]" pageId="27" pageNumber="418">
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- Males with scopula on tarsus IV; long bristle retroventrally at base of tibial apophysis thick, spine-like (
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<figureCitation id="5CA209C1FFFFFFECBF1EFE10390BFDF9" captionStart="Fig. 17" captionStartId="32.[151,186,1915,1936]" captionTargetBox="[151,1434,262,1844]" captionTargetId="figure@32.[148,1439,254,1865]" captionTargetPageId="32" captionText="Fig. 17. Liphistius indra sp. nov., details of left (A-I) and right (J-K) palp of three males; holotype (A, C-D, G, I), paratype matured 4. X. 2012 (B, F, H, K), paratype matured 24. VIII. 2011 (E, J). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view. (C) Palpal organ, retrodorsal view. (D) Same, proventral and slightly proximal view. (E-F) Tegulum, retrodorsal view. (G-H) Palpal organ, distal view (dorsal side up). (I) Paracymbium and tibial apophysis, retrolateral view. (J-K) Tibial apophysis, retrolateral and slightly proximal view. Scale lines: 1.0 mm (A, I; B-F, H; J-K), 0.5 mm (G)." httpUri="https://zenodo.org/record/1021973/files/figure.png" pageId="27" pageNumber="418">Fig. 17A</figureCitation>
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, I-K); paracymbium short (
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<figureCitation id="5CA209C1FFFFFFECB852FDF03AB9FDF9" box="[573,673,513,538]" captionStart="Fig. 17" captionStartId="32.[151,186,1915,1936]" captionTargetBox="[151,1434,262,1844]" captionTargetId="figure@32.[148,1439,254,1865]" captionTargetPageId="32" captionText="Fig. 17. Liphistius indra sp. nov., details of left (A-I) and right (J-K) palp of three males; holotype (A, C-D, G, I), paratype matured 4. X. 2012 (B, F, H, K), paratype matured 24. VIII. 2011 (E, J). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view. (C) Palpal organ, retrodorsal view. (D) Same, proventral and slightly proximal view. (E-F) Tegulum, retrodorsal view. (G-H) Palpal organ, distal view (dorsal side up). (I) Paracymbium and tibial apophysis, retrolateral view. (J-K) Tibial apophysis, retrolateral and slightly proximal view. Scale lines: 1.0 mm (A, I; B-F, H; J-K), 0.5 mm (G)." httpUri="https://zenodo.org/record/1021973/files/figure.png" pageId="27" pageNumber="418">Fig. 17A</figureCitation>
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); distal edge of contrategulum without gap behind only slightly or not bent dorsal apex (
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<figureCitation id="5CA209C1FFFFFFECBBCCFDD03A1DFDD9" box="[419,517,545,570]" captionStart="Fig. 17" captionStartId="32.[151,186,1915,1936]" captionTargetBox="[151,1434,262,1844]" captionTargetId="figure@32.[148,1439,254,1865]" captionTargetPageId="32" captionText="Fig. 17. Liphistius indra sp. nov., details of left (A-I) and right (J-K) palp of three males; holotype (A, C-D, G, I), paratype matured 4. X. 2012 (B, F, H, K), paratype matured 24. VIII. 2011 (E, J). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view. (C) Palpal organ, retrodorsal view. (D) Same, proventral and slightly proximal view. (E-F) Tegulum, retrodorsal view. (G-H) Palpal organ, distal view (dorsal side up). (I) Paracymbium and tibial apophysis, retrolateral view. (J-K) Tibial apophysis, retrolateral and slightly proximal view. Scale lines: 1.0 mm (A, I; B-F, H; J-K), 0.5 mm (G)." httpUri="https://zenodo.org/record/1021973/files/figure.png" pageId="27" pageNumber="418">Fig. 17B</figureCitation>
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, G-H). Poreplate with anterior margin only indistinctly invaginated (
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<figureCitation id="5CA209C1FFFFFFECBE84FDD03D23FDD9" box="[1259,1339,545,570]" captionStart="Fig. 18" captionStartId="33.[151,186,1968,1989]" captionTargetBox="[158,1428,576,1904]" captionTargetId="figure@33.[148,1439,544,1931]" captionTargetPageId="33" captionText="Fig. 18. Liphistius indra sp. nov., vulval plate of four female paratypes (all from exuviae): moult 15. III. 2012 (A-B), moult 10. II. 2001 (C-D), moult 15. VIII. 2002 (E-F), moult 3. XI. 2001 (G-H). (A, C, E) Entire structure, dorsal view. (B, D, F) Same, ventral view. (G) Anterior part of vulval plate, ventral view. (H) Receptacular cluster, ventral view. Scale lines: 1.0 mm (A-D, G-H; E-F)." httpUri="https://zenodo.org/record/1021975/files/figure.png" pageId="27" pageNumber="418">Fig. 18</figureCitation>
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); CDO a longitudinal slit or semicircular hole with an enlarged pore on each lateral side (
|
||
<figureCitation id="5CA209C1FFFFFFECBE2DFDB03D1DFDB9" box="[1090,1285,577,602]" captionStart="Fig. 18" captionStartId="33.[151,186,1968,1989]" captionTargetBox="[158,1428,576,1904]" captionTargetId="figure@33.[148,1439,544,1931]" captionTargetPageId="33" captionText="Fig. 18. Liphistius indra sp. nov., vulval plate of four female paratypes (all from exuviae): moult 15. III. 2012 (A-B), moult 10. II. 2001 (C-D), moult 15. VIII. 2002 (E-F), moult 3. XI. 2001 (G-H). (A, C, E) Entire structure, dorsal view. (B, D, F) Same, ventral view. (G) Anterior part of vulval plate, ventral view. (H) Receptacular cluster, ventral view. Scale lines: 1.0 mm (A-D, G-H; E-F)." httpUri="https://zenodo.org/record/1021975/files/figure.png" pageId="27" pageNumber="418">Fig. 18A, C, E, G</figureCitation>
|
||
); receptacular cluster not divided into lateral halves, its vesicles quite long (
|
||
<figureCitation id="5CA209C1FFFFFFECB91FFD903C32FD99" box="[880,1066,609,634]" captionStart="Fig. 18" captionStartId="33.[151,186,1968,1989]" captionTargetBox="[158,1428,576,1904]" captionTargetId="figure@33.[148,1439,544,1931]" captionTargetPageId="33" captionText="Fig. 18. Liphistius indra sp. nov., vulval plate of four female paratypes (all from exuviae): moult 15. III. 2012 (A-B), moult 10. II. 2001 (C-D), moult 15. VIII. 2002 (E-F), moult 3. XI. 2001 (G-H). (A, C, E) Entire structure, dorsal view. (B, D, F) Same, ventral view. (G) Anterior part of vulval plate, ventral view. (H) Receptacular cluster, ventral view. Scale lines: 1.0 mm (A-D, G-H; E-F)." httpUri="https://zenodo.org/record/1021975/files/figure.png" pageId="27" pageNumber="418">Fig. 18B, D, F, H</figureCitation>
|
||
) ....
|
||
<taxonomicName id="03996EC7FFFFFFECBE3BFD923CB2FD9B" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1108,1194,609,634]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="indra" status="sp. nov.">
|
||
<emphasis id="F6EDC956FFFFFFECBE3BFD923CB2FD9B" box="[1108,1194,609,634]" italics="true" pageId="27" pageNumber="418">L. indra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EDDE742DFFFFFFECBEDFFD923D1BFD99" box="[1200,1283,611,634]" pageId="27" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(in Thailand)
|
||
</paragraph>
|
||
<paragraph id="C4261544FFFFFFECBAF8FD4E38EFFB2D" blockId="27.[151,756,702,1230]" pageId="27" pageNumber="418">
|
||
group and the
|
||
<taxonomicName id="03996EC7FFFFFFECBB5DFD4E3964FD35" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[306,380,703,726]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFFFFFECBB5DFD4E3964FD35" box="[306,380,703,726]" italics="true" pageId="27" pageNumber="418">tioman</emphasis>
|
||
</taxonomicName>
|
||
-group by males possessing a large para-embolic plate with a coarsely serrate distal margin (
|
||
<figureCitation id="5CA209C1FFFFFFECBAF0FD0C38EFFCF7" box="[159,247,765,788]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Figs 15</figureCitation>
|
||
A-D, 17A-D), a narrow and pointed dorsal apex of the contrategulum (
|
||
<figureCitation id="5CA209C1FFFFFFECBBA5FCED3A03FCD0" box="[458,539,796,819]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Figs 15</figureCitation>
|
||
E-F, 17G-H), and a retrolateral tibial apophysis deeply divided into a long, deep retroventral part and a very short, small retrodorsal part (
|
||
<figureCitation id="5CA209C1FFFFFFECBABBFC8A3930FC71" box="[212,296,891,914]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Figs 15</figureCitation>
|
||
G-I, 17I-K) (tibial apophysis also deeply divided in
|
||
<taxonomicName id="03996EC7FFFFFFECBB67FC6D3975FC50" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[264,365,924,947]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFFFFFECBB67FC6D3975FC50" box="[264,365,924,947]" italics="true" pageId="27" pageNumber="418">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
but in that species both parts equally long;
|
||
<figureCitation id="5CA209C1FFFFFFECBAB6FC4B3924FC30" box="[217,316,954,979]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="27" pageNumber="418">Fig. 11A</figureCitation>
|
||
, G-I); three megaspines on retroventral part short and bent, single megaspine on retrodorsal part short, curved and weaker than other megaspines (
|
||
<figureCitation id="5CA209C1FFFFFFECB8AAFC0838DFFBCC" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Figs 15A</figureCitation>
|
||
, G-I; 17A, I-K). Females distinguished by a peculiar vulval plate with an only partially sclerotised poreplate widely separated from small remnant of posterior stalk situated at posterior margin of vulva (
|
||
<figureCitation id="5CA209C1FFFFFFECB856FB863A93FB6D" box="[569,651,1143,1166]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="27" pageNumber="418">Figs 16</figureCitation>
|
||
A-E, 18); CDO small, in many cases slit-like (
|
||
<figureCitation id="5CA209C1FFFFFFECB87BFB673A62FB4E" box="[532,634,1174,1197]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="27" pageNumber="418">Figs 16A</figureCitation>
|
||
, C-D; 18A, C, E, G).
|
||
</paragraph>
|
||
<paragraph id="C4261544FFFFFFECBAF8FB1538F2FAFF" blockId="27.[151,756,1252,1308]" pageId="27" pageNumber="418">
|
||
<emphasis id="F6EDC956FFFFFFECBAF8FB153979FB18" bold="true" box="[151,353,1252,1275]" pageId="27" pageNumber="418">Species included:</emphasis>
|
||
<taxonomicName id="03996EC7FFFFFFECBB03FB153A3EFB18" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[364,550,1252,1275]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
|
||
<emphasis id="F6EDC956FFFFFFECBB03FB153A3EFB18" box="[364,550,1252,1275]" italics="true" pageId="27" pageNumber="418">Liphistius linang</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EDDE742DFFFFFFECB85EFB143A92FB1F" box="[561,650,1253,1276]" pageId="27" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="03996EC7FFFFFFECB8F6FB143AECFB18" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[665,756,1252,1276]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="indra" status="sp. nov.">
|
||
<emphasis id="F6EDC956FFFFFFECB8F6FB143AECFB18" box="[665,756,1252,1276]" italics="true" pageId="27" pageNumber="418">L. indra</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EDDE742DFFFFFFECBAF8FAF438F2FAFF" box="[151,234,1285,1308]" pageId="27" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFFFFFECBAF8FAC33979F901" pageId="27" pageNumber="418" type="description">
|
||
<paragraph id="C4261544FFFFFFECBAF8FAC33979F901" blockId="27.[151,757,1330,1763]" pageId="27" pageNumber="418">
|
||
<emphasis id="F6EDC956FFFFFFECBAF8FAC33925FAAA" bold="true" box="[151,317,1330,1353]" pageId="27" pageNumber="418">Relationships:</emphasis>
|
||
The
|
||
<taxonomicName id="03996EC7FFFFFFECBBEBFAC339D1FAAA" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[388,457,1330,1353]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang">
|
||
<emphasis id="F6EDC956FFFFFFECBBEBFAC339D1FAAA" box="[388,457,1330,1353]" italics="true" pageId="27" pageNumber="418">linang</emphasis>
|
||
</taxonomicName>
|
||
-group is morphologically intermediate between the
|
||
<taxonomicName id="03996EC7FFFFFFECBB89FAA23A2AFA89" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[486,562,1363,1386]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFFFFFECBB89FAA23A2AFA89" box="[486,562,1363,1386]" italics="true" pageId="27" pageNumber="418">tioman</emphasis>
|
||
</taxonomicName>
|
||
-group and the
|
||
<taxonomicName id="03996EC7FFFFFFECBAF8FA8038E6FA6B" authorityName="Abraham" authorityYear="1923" box="[151,254,1393,1416]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="batuensis">
|
||
<emphasis id="F6EDC956FFFFFFECBAF8FA8038E6FA6B" box="[151,254,1393,1416]" italics="true" pageId="27" pageNumber="418">batuensis</emphasis>
|
||
</taxonomicName>
|
||
-group. Sharing with
|
||
<taxonomicName id="03996EC7FFFFFFECB867FA833A63FA6A" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[520,635,1394,1417]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFFFFFECB867FA833A63FA6A" box="[520,635,1394,1417]" italics="true" pageId="27" pageNumber="418">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
a deeply divided tibial apophysis with the retrodorsal-apical megaspine distinctly separated from the other three megaspines, and sharing with this and other species in the
|
||
<taxonomicName id="03996EC7FFFFFFECBAA7FA01390BF9E4" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[200,275,1520,1543]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFFFFFECBAA7FA01390BF9E4" box="[200,275,1520,1543]" italics="true" pageId="27" pageNumber="418">tioman</emphasis>
|
||
</taxonomicName>
|
||
-group a distinctly elevated para-embolic plate. Females of the
|
||
<taxonomicName id="03996EC7FFFFFFECBBD2F9FF3A1BF9C6" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[445,515,1550,1573]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang">
|
||
<emphasis id="F6EDC956FFFFFFECBBD2F9FF3A1BF9C6" box="[445,515,1550,1573]" italics="true" pageId="27" pageNumber="418">linang</emphasis>
|
||
</taxonomicName>
|
||
-group show strong resemblance with those of the
|
||
<taxonomicName id="03996EC7FFFFFFECB877F9DF3A67F9A6" authorityName="Abraham" authorityYear="1923" box="[536,639,1582,1605]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="batuensis">
|
||
<emphasis id="F6EDC956FFFFFFECB877F9DF3A67F9A6" box="[536,639,1582,1605]" italics="true" pageId="27" pageNumber="418">batuensis</emphasis>
|
||
</taxonomicName>
|
||
-group by possessing an only partially sclerotised poreplate. However, females of
|
||
<taxonomicName id="03996EC7FFFFFFECBBEAF99F39F1F967" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[389,489,1645,1669]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
|
||
<emphasis id="F6EDC956FFFFFFECBBEAF99F39F1F967" box="[389,489,1645,1669]" italics="true" pageId="27" pageNumber="418">L. linang</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EDDE742DFFFFFFECBB9CF99F3A51F966" box="[499,585,1646,1669]" pageId="27" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
|
||
have a divided receptacular cluster, as do females of all
|
||
<taxonomicName id="03996EC7FFFFFFECB80DF97F3AB4F946" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[610,684,1678,1701]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFFFFFECB80DF97F3AB4F946" box="[610,684,1678,1701]" italics="true" pageId="27" pageNumber="418">tioman</emphasis>
|
||
</taxonomicName>
|
||
-group species. The relationships of the linang-group are therefore not clear.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFFFFFECBAF8F90B3A70F8D1" pageId="27" pageNumber="418" type="distribution">
|
||
<paragraph id="C4261544FFFFFFECBAF8F90B3A70F8D1" blockId="27.[151,756,1786,1842]" pageId="27" pageNumber="418">
|
||
<emphasis id="F6EDC956FFFFFFECBAF8F90B3936F8F2" bold="true" box="[151,302,1786,1809]" pageId="27" pageNumber="418">Distribution:</emphasis>
|
||
Northeastern peninsular
|
||
<collectingCountry id="BC8E55D4FFFFFFECB835F90B3AA5F8F2" box="[602,701,1786,1809]" name="Malaysia" pageId="27" pageNumber="418">Malaysia</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="BC8E55D4FFFFFFECBA96F8E83940F8D3" box="[249,344,1817,1840]" name="Thailand" pageId="27" pageNumber="418">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="5CA209C1FFFFFFECBB08F8E839BDF8D1" box="[359,421,1817,1842]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="27" pageNumber="418">Fig. 1</figureCitation>
|
||
, localities 13-14).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |