309 lines
61 KiB
XML
309 lines
61 KiB
XML
<document id="70C07EBCFB37CBD6C0ABABDFD56CC41A" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFF7FFE3B91EFC4A3D4CF7E7" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius tioman Platnick & Sedgwick 1984" docType="treatment" docVersion="9" lastPageNumber="411" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="410" updateTime="1698469770161" updateUser="ExternalLinkService">
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<mods:title id="4946389CE29CED31BD097326A0621D23">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
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<mods:namePart id="7AF819F0D3FDF8CEEE4E05697ED7AF52">Peter J. Schwendinger</mods:namePart>
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<mods:title id="BD77314F6B5253D95DA1FD731AB2241D">Revue suisse de Zoologie</mods:title>
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<mods:part id="BF71F77731EC08F1C6F7927678643619">
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<mods:date id="1A3527E9FD49B0139455253DC1E59A93">2017</mods:date>
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<mods:number id="0901D90D842F62596632471C0E7F859C">2017-09-30</mods:number>
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<treatment id="4C30A452FFF7FFE3B91EFC4A3D4CF7E7" ID-DOI="http://doi.org/10.5281/zenodo.6042361" ID-GBIF-Taxon="135450774" ID-Zenodo-Dep="6042361" LSID="urn:lsid:plazi:treatment:4C30A452FFF7FFE3B91EFC4A3D4CF7E7" httpUri="http://treatment.plazi.org/id/4C30A452FFF7FFE3B91EFC4A3D4CF7E7" lastPageId="20" lastPageNumber="411" pageId="19" pageNumber="410">
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<subSubSection id="8C8346CFFFF7FFE4B91EFC4A3D72FC31" box="[881,1386,955,978]" pageId="19" pageNumber="410" type="nomenclature">
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<paragraph id="C4261544FFF7FFE4B91EFC4A3D72FC31" blockId="19.[881,1386,955,1012]" box="[881,1386,955,978]" pageId="19" pageNumber="410">
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<taxonomicName id="03996EC7FFF7FFE4B91EFC4A3D72FC31" ID-CoL="72J77" authority="Platnick & Sedgwick, 1984" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[881,1386,955,978]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="410" phylum="Arthropoda" rank="species" species="tioman">
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<emphasis id="F6EDC956FFF7FFE4B91EFC4A3D72FC31" bold="true" box="[881,1386,955,978]" pageId="19" pageNumber="410">
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<emphasis id="F6EDC956FFF7FFE4B91EFC4A3C2BFC31" bold="true" box="[881,1075,955,978]" italics="true" pageId="19" pageNumber="410">Liphistius tioman</emphasis>
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<bibRefCitation id="A00868B5FFF7FFE4BE56FC4A3D72FC31" author="Platnick N. & Sedgwick W. C." box="[1081,1386,955,978]" journalOrPublisher="American Museum Novitates" pageId="19" pageNumber="410" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick, 1984</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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||
<paragraph id="C4261544FFF7FFE4BE5AFC2D3CBFFC17" blockId="19.[881,1386,955,1012]" box="[1077,1191,988,1012]" pageId="19" pageNumber="410">
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<figureCitation id="5CA209C1FFF7FFE4BE5AFC2D3CBFFC17" box="[1077,1191,988,1012]" captionStart-0="Fig. 11" captionStart-1="Fig. 12" captionStartId-0="21.[151,186,1705,1726]" captionStartId-1="22.[151,186,1724,1745]" captionTargetBox-0="[158,1428,255,1634]" captionTargetBox-1="[151,1421,259,1654]" captionTargetId-0="figure@21.[148,1439,254,1658]" captionTargetId-1="figure@22.[148,1439,234,1682]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." captionText-1="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri-0="https://zenodo.org/record/1021959/files/figure.png" httpUri-1="https://zenodo.org/record/1021961/files/figure.png" pageId="19" pageNumber="410">Figs 11-12</figureCitation>
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</paragraph>
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<paragraph id="C4261544FFF7FFE4B950FBFB3D84FBC3" blockId="19.[831,1436,1034,1085]" box="[831,1436,1034,1056]" pageId="19" pageNumber="410">
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<treatmentCitationGroup id="E489326AFFF7FFE4B950FBFB3D84FBC3" box="[831,1436,1034,1056]" pageId="19" pageNumber="410">
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<taxonomicName id="03996EC7FFF7FFE4B950FBFB3D3DFBC3" ID-CoL="72J77" authority="Platnick & Sedgwick, 1984: 28" authorityName="Platnick & Sedgwick" authorityPageNumber="28" authorityYear="1984" box="[831,1317,1034,1056]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="410" phylum="Arthropoda" rank="species" species="tioman">
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<emphasis id="F6EDC956FFF7FFE4B950FBFB3BF1FBC3" box="[831,1001,1034,1056]" italics="true" pageId="19" pageNumber="410">Liphistius tioman</emphasis>
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<treatmentCitation id="45383355FFF7FFE4B99FFBFB3D3DFBC3" author="Platnick N. & Sedgwick W. C." box="[1008,1317,1034,1056]" page="28" pageId="19" pageNumber="410" year="1984">
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<bibRefCitation id="A00868B5FFF7FFE4B99FFBFB3D1AFBC3" author="Platnick N. & Sedgwick W. C." box="[1008,1282,1034,1056]" journalOrPublisher="American Museum Novitates" pageId="19" pageNumber="410" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick, 1984</bibRefCitation>
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: 28
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</treatmentCitation>
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</taxonomicName>
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-29, figs 81-
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="C4261544FFF7FFE4B9E4FBD93CF1FBDF" blockId="19.[831,1436,1034,1085]" box="[907,1257,1063,1085]" pageId="19" pageNumber="410">87 (description of male and female).</paragraph>
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<subSubSection id="8C8346CFFFF7FFE3B950FBA7392EFEB9" lastPageId="20" lastPageNumber="411" pageId="19" pageNumber="410" type="materials_examined">
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||
<subSubSection id="8C8346CFFFF7FFE4B950FBA73CB3FB2D" pageId="19" pageNumber="410" type="materials_examined">
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||
<paragraph id="C4261544FFF7FFE4B950FBA73CB3FB2D" blockId="19.[831,1436,1109,1232]" pageId="19" pageNumber="410">
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||
<emphasis id="F6EDC956FFF7FFE4B950FBA73BEAFB8F" bold="true" box="[831,1010,1109,1134]" pageId="19" pageNumber="410">
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<typeStatus id="1B22ABE6FFF7FFE4B950FBA73B6FFB8D" box="[831,887,1110,1134]" pageId="19" pageNumber="410">Type</typeStatus>
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material:
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</emphasis>
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<materialsCitation id="74F11F19FFF7FFE4BE6EFBA73CB3FB2D" ID-GBIF-Occurrence="1675315706" collectingDate="1982-07-29" collectionCode="AMHN" collectorName="W. C. Sedgwick" country="Malaysia" elevation="792" location="Gua Sinah and Gua Panah" municipality="Tioman" pageId="19" pageNumber="410" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
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||
<collectionCode id="A2888D81FFF7FFE4BE6EFBA73C4FFB8D" box="[1025,1111,1110,1134]" collectionName="AMHN" pageId="19" pageNumber="410">AMHN</collectionCode>
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;
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||
<specimenCount id="D29FDECDFFF7FFE4BE05FBA43C87FB8F" box="[1130,1183,1109,1132]" pageId="19" pageNumber="410" type="male">male</specimenCount>
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||
<typeStatus id="1B22ABE6FFF7FFE4BEC1FBA43D15FB8F" box="[1198,1293,1109,1132]" pageId="19" pageNumber="410" type="holotype">holotype</typeStatus>
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and
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||
<specimenCount id="D29FDECDFFF7FFE4BF3CFBA43D84FB8F" box="[1363,1436,1109,1132]" pageId="19" pageNumber="410" type="female">female</specimenCount>
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||
<typeStatus id="1B22ABE6FFF7FFE4B950FB863B84FB6C" box="[831,924,1143,1167]" pageId="19" pageNumber="410" type="paratype">paratype</typeStatus>
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(not examined);
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||
<collectingCountry id="BC8E55D4FFF7FFE4BE02FB873CCBFB6E" box="[1133,1235,1142,1165]" name="Malaysia" pageId="19" pageNumber="410">Malaysia</collectingCountry>
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||
,
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||
<collectingRegion id="065DDBA6FFF7FFE4BE8AFB873D2FFB6E" box="[1253,1335,1142,1165]" country="Malaysia" name="Pahang" pageId="19" pageNumber="410">Pahang</collectingRegion>
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||
,
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<collectingMunicipality id="24428F3EFFF7FFE4BF26FB873D84FB6E" box="[1353,1436,1142,1165]" pageId="19" pageNumber="410">Tioman</collectingMunicipality>
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||
<collectingCountry id="BC8E55D4FFF7FFE4B950FB673B9CFB4E" box="[831,900,1174,1197]" name="Iceland" pageId="19" pageNumber="410">Island</collectingCountry>
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,
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||
<location id="C146439FFFF7FFE4B9FAFB663CD4FB4E" LSID="urn:lsid:plazi:treatment:4C30A452FFF7FFE3B91EFC4A3D4CF7E7:C146439FFFF7FFE4B9FAFB663CD4FB4E" box="[917,1228,1174,1199]" country="Malaysia" municipality="Tioman" name="Gua Sinah and Gua Panah" pageId="19" pageNumber="410" stateProvince="Pahang">Gua Sinah and Gua Panah</location>
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,
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<elevation id="4FB4F277FFF7FFE4BEB2FB673D2BFB4E" box="[1245,1331,1174,1197]" metricMagnitude="2" metricUnit="m" metricValue="7.9248" pageId="19" pageNumber="410" unit="ft" value="2600.0">
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<quantity id="0361B8A1FFF7FFE4BEB2FB673D2BFB4E" box="[1245,1331,1174,1197]" metricMagnitude="2" metricUnit="m" metricValue="7.9248" pageId="19" pageNumber="410" unit="ft" value="2600.0">2600 ft</quantity>
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</elevation>
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. altitude;
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<collectingDate id="A063CA6CFFF7FFE4B950FB493BDDFB33" box="[831,965,1208,1232]" pageId="19" pageNumber="410" value="1982-07-29">
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<date id="B0273384FFF7FFE4B950FB493BDDFB33" box="[831,965,1208,1232]" pageId="19" pageNumber="410" value="1982-07-29">29.VII.1982</date>
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</collectingDate>
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; leg.
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<collectorName id="696C7092FFF7FFE4B992FB493CBFFB2D" box="[1021,1191,1207,1232]" pageId="19" pageNumber="410">W.C. Sedgwick</collectorName>
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||
.
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||
</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<paragraph id="C4261544FFF7FFE4B950FB163D84FABC" blockId="19.[831,1436,1255,1377]" pageId="19" pageNumber="410">
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||
<emphasis id="F6EDC956FFF7FFE4B950FB163C2CFB1D" bold="true" box="[831,1076,1255,1278]" pageId="19" pageNumber="410">Material examined:</emphasis>
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MHNG, sample SIM-01/09; 1 male (matured
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<date id="B0273384FFF7FFE4BE47FAF93C9EFAC3" box="[1064,1158,1288,1312]" pageId="19" pageNumber="410" value="2002-01-03">3.I.2002</date>
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), 7 females (moulted
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<date id="B0273384FFF7FFE4B950FAD83BDFFAA2" box="[831,967,1321,1345]" pageId="19" pageNumber="410" value="2001-12-17">17.XII.2001</date>
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||
;
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||
<date id="B0273384FFF7FFE4B9B4FAD83C53FAA2" box="[987,1099,1321,1345]" pageId="19" pageNumber="410" value="2001-09-02">2.IX.2001</date>
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||
, 6.III.,
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||
<date id="B0273384FFF7FFE4BEC3FAD83D0BFAA2" box="[1196,1299,1321,1345]" pageId="19" pageNumber="410" value="2002-10-04">4.X.2002</date>
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||
,
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||
<date id="B0273384FFF7FFE4BF48FAD83D80FAA2" box="[1319,1432,1321,1345]" pageId="19" pageNumber="410" value="2003-06-07">7.VI.2003</date>
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||
;
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||
<date id="B0273384FFF7FFE4B950FAB83BA8FA82" box="[831,944,1353,1377]" pageId="19" pageNumber="410" value="2001-09-08">8.IX.2001</date>
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||
, 16.II.,
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||
<date id="B0273384FFF7FFE4BE78FAB83C8EFA82" box="[1047,1174,1353,1377]" pageId="19" pageNumber="410" value="2002-06-23">23.VI.2002</date>
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||
), 1 penultimate male;
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||
</paragraph>
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||
<paragraph id="C4261544FFF0FFE3BAF8FEF3392EFEB9" blockId="20.[151,756,258,346]" pageId="20" pageNumber="411">
|
||
<materialsCitation id="74F11F19FFF0FFE3BAF8FEF3392EFEB9" ID-GBIF-Occurrence="1675315707" collectingDate="2001-06-30" collectorName="P. J. Schwendinger." elevation="835" latitude="2.7625" location="Gunung Kajang" longLatPrecision="21" longitude="104.14528" municipality="Tioman Island" pageId="20" pageNumber="411" specimenCount="1" stateProvince="Pahang">
|
||
<collectingRegion id="065DDBA6FFF0FFE3BAF8FEF338F1FEFA" box="[151,233,258,281]" country="Malaysia" name="Pahang" pageId="20" pageNumber="411">Pahang</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="24428F3EFFF0FFE3BA96FEF33985FEFA" box="[249,413,258,281]" pageId="20" pageNumber="411">Tioman Island</collectingMunicipality>
|
||
,
|
||
<location id="C146439FFFF0FFE3BBC2FEF23A46FEFA" LSID="urn:lsid:plazi:treatment:4C30A452FFF7FFE3B91EFC4A3D4CF7E7:C146439FFFF0FFE3BBC2FEF23A46FEFA" box="[429,606,258,283]" latitude="2.7625" longLatPrecision="21" longitude="104.14528" municipality="Tioman Island" name="Gunung Kajang" pageId="20" pageNumber="411" stateProvince="Pahang">Gunung Kajang</location>
|
||
(
|
||
<geoCoordinate id="A1AD7383FFF0FFE3B81DFEF33AF7FEFA" box="[626,751,258,281]" direction="north" orientation="latitude" pageId="20" pageNumber="411" precision="15" value="2.7625">2°45’45’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A1AD7383FFF0FFE3BAF8FED23936FED8" box="[151,302,291,315]" direction="east" orientation="longitude" pageId="20" pageNumber="411" precision="15" value="104.14528">104°08’43’’E</geoCoordinate>
|
||
),
|
||
<elevation id="4FB4F277FFF0FFE3BB3DFED339C3FED9" box="[338,475,290,314]" metricMagnitude="2" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="20" pageNumber="411" unit="m" value="835.0" valueMax="850.0" valueMin="820.0">
|
||
<quantity id="0361B8A1FFF0FFE3BB3DFED339C3FED9" box="[338,475,290,314]" metricMagnitude="2" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="20" pageNumber="411" unit="m" value="835.0" valueMax="850.0" valueMin="820.0">820-850 m</quantity>
|
||
</elevation>
|
||
;
|
||
<collectingDate id="A063CA6CFFF0FFE3BB99FED23A6EFED8" box="[502,630,291,315]" pageId="20" pageNumber="411" value="2001-06-30">
|
||
<date id="B0273384FFF0FFE3BB99FED23A6EFED8" box="[502,630,291,315]" pageId="20" pageNumber="411" value="2001-06-30">30.VI.2001</date>
|
||
</collectingDate>
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||
; leg.
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||
<collectorName id="696C7092FFF0FFE3B8A0FED2392EFEB9" pageId="20" pageNumber="411">P.J. Schwendinger.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFF0FFE3BAF8FE833ABCFC5C" pageId="20" pageNumber="411" type="diagnosis">
|
||
<paragraph id="C4261544FFF0FFE3BAF8FE833975FD13" blockId="20.[151,756,370,752]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3BAF8FE833916FE6A" bold="true" box="[151,270,370,393]" pageId="20" pageNumber="411">Diagnosis:</emphasis>
|
||
Males distinguished from those of all other
|
||
<taxonomicName id="03996EC7FFF0FFE3BAF8FE62391AFE49" authorityName="Schiodte" authorityYear="1849" box="[151,258,403,426]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F6EDC956FFF0FFE3BAF8FE62391AFE49" box="[151,258,403,426]" italics="true" pageId="20" pageNumber="411">Liphistius</emphasis>
|
||
</taxonomicName>
|
||
species by a deeply and widely divided retrolateral apophysis on palpal tibia, both parts being equally long (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB40FE22398BFE0F" box="[303,403,467,492]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11A</figureCitation>
|
||
, H-I). Males distinguished from those of other species in the
|
||
<taxonomicName id="03996EC7FFF0FFE3BBB5FE043A3CFDEF" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[474,548,501,524]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFF0FFE3BBB5FE043A3CFDEF" box="[474,548,501,524]" italics="true" pageId="20" pageNumber="411">tioman</emphasis>
|
||
</taxonomicName>
|
||
-group additionally by a long and pointed para-embolic plate (
|
||
<figureCitation id="5CA209C1FFF0FFE3B81EFDE53AA5FDCE" box="[625,701,532,557]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11</figureCitation>
|
||
C-E) and by a very narrow tegulum with few denticles on proximal edge, situated very close to dorsal apex of contrategulum (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB27FD8739CAFD6D" box="[328,466,630,654]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11B, F</figureCitation>
|
||
). Females with poreplate widely fused with posterior stalk; receptacular cluster medially more or less distinctly divided; CDO undivided (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB7FFD263978FD13" box="[272,352,727,752]" captionStart="Fig. 12" captionStartId="22.[151,186,1724,1745]" captionTargetBox="[151,1421,259,1654]" captionTargetId="figure@22.[148,1439,234,1682]" captionTargetPageId="22" captionText="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021961/files/figure.png" pageId="20" pageNumber="411">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C4261544FFF0FFE3BAF8FCF73ABCFC5C" blockId="20.[151,757,774,1901]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3BAF8FCF73A3BFCFC" bold="true" box="[151,547,774,799]" pageId="20" pageNumber="411">Additions to description of male:</emphasis>
|
||
<emphasis id="F6EDC956FFF0FFE3B841FCF73A9DFCFE" box="[558,645,774,797]" italics="true" pageId="20" pageNumber="411">Scopula</emphasis>
|
||
: In distal two-thirds of ventral side of tarsus I, in distal threequarters of tarsus II, in distal two-thirds of tarsus III, in distal half of tarsus IV (narrower than on other tarsi). All scopulae thin and only distally (behind the claws) divided by a glabrous longitudinal median stripe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFF0FFE3BAF8FC383D08FD7D" pageId="20" pageNumber="411" type="description">
|
||
<paragraph id="C4261544FFF0FFE3BAF8FC383914F88E" blockId="20.[151,757,774,1901]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3BAF8FC3838D2FC03" box="[151,202,969,992]" italics="true" pageId="20" pageNumber="411">Palp</emphasis>
|
||
: Tibial apophysis situated apically, not set back from anterior margin of palp (
|
||
<figureCitation id="5CA209C1FFF0FFE3B874FC183A94FBE1" box="[539,652,1001,1026]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11G</figureCitation>
|
||
), deeply divided: retrolateral part deeper, aligned with axis of tibia and carrying three very short pointed megaspines; retrodorsal part much narrower, pointing away from axis of tibia, carrying a single, slightly longer pointed magaspine (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB73FB7A3999FB47" box="[284,385,1163,1188]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11A</figureCitation>
|
||
, G-I). Distal margin of cymbium with very indistinct lobes (
|
||
<figureCitation id="5CA209C1FFF0FFE3BBB6FB5D3A5BFB27" box="[473,579,1196,1220]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11C</figureCitation>
|
||
). Paracymbium relatively short and moderately deep (
|
||
<figureCitation id="5CA209C1FFF0FFE3B819FB3D3AFFFB06" box="[630,743,1228,1253]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11A</figureCitation>
|
||
); cumulus indistinct, carrying a fairly compact group of long stiff bristles (
|
||
<figureCitation id="5CA209C1FFF0FFE3BBFEFAFC39E2FAC5" box="[401,506,1293,1318]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11A</figureCitation>
|
||
). Subtegulum without apophysis. Tegulum very narrow, its proximal edge carrying few overlapping scale-like denticles, situated close to dorsal apex of contrategulum (
|
||
<figureCitation id="5CA209C1FFF0FFE3B831FA9F3AFFFA64" box="[606,743,1390,1415]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11B, F</figureCitation>
|
||
). Contrategulum with large, conical ventral process being slightly constricted at base (
|
||
<figureCitation id="5CA209C1FFF0FFE3BBB5FA5E3A27FA2B" box="[474,575,1455,1480]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11F</figureCitation>
|
||
); distal edge of contrategulum distinctly elevated and composed of numerous parallel ribs instead of teeth (
|
||
<figureCitation id="5CA209C1FFF0FFE3B807FA013AA0F9EA" box="[616,696,1520,1545]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11</figureCitation>
|
||
B-F), dorsal apex relatively small and narrowly rounded, only slighly elevated above underlying surface (
|
||
<figureCitation id="5CA209C1FFF0FFE3B807F9C03AF0F9AA" box="[616,744,1585,1609]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11B, F</figureCitation>
|
||
). Para-embolic plate long, narrow and asymmetrically triangular (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB70F9833962F969" box="[287,378,1650,1674]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11</figureCitation>
|
||
B-E); embolus proper strongly inclined prolaterad (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB1AF96339CFF948" box="[373,471,1682,1707]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11B</figureCitation>
|
||
), distally widened; dorsal wall of sclerotised part distinctly wider than ventral wall, ending in a quadrangular lobe (
|
||
<figureCitation id="5CA209C1FFF0FFE3B82BF9223A8CF90F" box="[580,660,1747,1772]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11</figureCitation>
|
||
B-C, F), retrolateral wall not enforced by ridges but only with an indistinct, short distal keel (
|
||
<figureCitation id="5CA209C1FFF0FFE3BBA1F8E53A4AF8CF" box="[462,594,1812,1836]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11B, F</figureCitation>
|
||
); membranous part of embolus proper slightly widened distally (
|
||
<figureCitation id="5CA209C1FFF0FFE3BAF0F8A538E7F88E" box="[159,255,1876,1901]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C4261544FFF0FFE3BAF8F8753D08FD7D" blockId="20.[151,756,1924,2078]" lastBlockId="20.[831,1437,258,670]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3BAF8F8753A5BF878" bold="true" box="[151,579,1924,1949]" pageId="20" pageNumber="411">Additions to description of female:</emphasis>
|
||
Genital atrium of vulval plate (
|
||
<figureCitation id="5CA209C1FFF0FFE3BB23F8553986F85E" box="[332,414,1956,1981]" captionStart="Fig. 12" captionStartId="22.[151,186,1724,1745]" captionTargetBox="[151,1421,259,1654]" captionTargetId="figure@22.[148,1439,234,1682]" captionTargetPageId="22" captionText="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021961/files/figure.png" pageId="20" pageNumber="411">Fig. 12</figureCitation>
|
||
) without median hairs, few to several lateral hairs present in some females; lateral margin of genital atrium not forming flaps bending ventrad and inwards (as usual in
|
||
<taxonomicName id="03996EC7FFF0FFE3B878F7F73A99F7FE" authorityName="Schiodte" authorityYear="1849" box="[535,641,2054,2077]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F6EDC956FFF0FFE3B878F7F73A99F7FE" box="[535,641,2054,2077]" italics="true" pageId="20" pageNumber="411">Liphistius</emphasis>
|
||
</taxonomicName>
|
||
) but only indistinct mounds on the dorsal (not the ventral) side (
|
||
<figureCitation id="5CA209C1FFF0FFE3B910FED33C37FED8" box="[895,1071,290,315]" captionStart="Fig. 12" captionStartId="22.[151,186,1724,1745]" captionTargetBox="[151,1421,259,1654]" captionTargetId="figure@22.[148,1439,234,1682]" captionTargetPageId="22" captionText="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021961/files/figure.png" pageId="20" pageNumber="411">Fig. 12A, C, F</figureCitation>
|
||
). Posterior stalk relatively short and wide, fused with poreplate without recognizable transition, reaching lateral margins of vulval plate in some females. Poreplate of variable shape, in most females examined anteriorly slightly narrower than posteriorly, its anterior margin indistinctly invaginated, without lobes; CDO quite large, undivided, anteriorly deep, posteriorly level with rest of poreplate (
|
||
<figureCitation id="5CA209C1FFF0FFE3BF1FFDF53BACFDDF" captionStart="Fig. 12" captionStartId="22.[151,186,1724,1745]" captionTargetBox="[151,1421,259,1654]" captionTargetId="figure@22.[148,1439,234,1682]" captionTargetPageId="22" captionText="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021961/files/figure.png" pageId="20" pageNumber="411">Fig. 12A, C, F</figureCitation>
|
||
; but see illustration of
|
||
<typeStatus id="1B22ABE6FFF0FFE3BED1FDD43D02FDDE" box="[1214,1306,549,573]" pageId="20" pageNumber="411" type="paratype">paratype</typeStatus>
|
||
in
|
||
<bibRefCitation id="A00868B5FFF0FFE3BF2BFDD53C0CFDBE" author="Platnick N. & Sedgwick W. C." journalOrPublisher="American Museum Novitates" pageId="20" pageNumber="411" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick, 1984</bibRefCitation>
|
||
: fig. 86); receptacular cluster quite large and wide, more or less distinctly divided into two paramedian subclusters (
|
||
<figureCitation id="5CA209C1FFF0FFE3BE27FD743CB1FD7D" box="[1096,1193,645,670]" captionStart="Fig. 12" captionStartId="22.[151,186,1724,1745]" captionTargetBox="[151,1421,259,1654]" captionTargetId="figure@22.[148,1439,234,1682]" captionTargetPageId="22" captionText="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021961/files/figure.png" pageId="20" pageNumber="411">Fig. 12B</figureCitation>
|
||
, D-E, G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFF0FFE3B950FD443BDEFB1C" pageId="20" pageNumber="411" type="discussion">
|
||
<paragraph id="C4261544FFF0FFE3B950FD443D77FC4C" blockId="20.[831,1437,693,943]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3B950FD443C37FD2E" bold="true" box="[831,1071,693,717]" pageId="20" pageNumber="411">Taxonomic remarks:</emphasis>
|
||
The tibial apophysis of the male palp of
|
||
<taxonomicName id="03996EC7FFF0FFE3B9FCFD273BE1FD0E" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[915,1017,726,749]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="F6EDC956FFF0FFE3B9FCFD273BE1FD0E" box="[915,1017,726,749]" italics="true" pageId="20" pageNumber="411">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
is unique and distinctly autapomorhic. A similar divided tibial apophysis is found in males of the
|
||
<taxonomicName id="03996EC7FFF0FFE3B907FCE43BB4FCCF" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[872,940,789,812]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="species" species="linang">
|
||
<emphasis id="F6EDC956FFF0FFE3B907FCE43BB4FCCF" box="[872,940,789,812]" italics="true" pageId="20" pageNumber="411">linang</emphasis>
|
||
</taxonomicName>
|
||
-group, but that is considered to have evolved convergently (see also Discussion). The wide ventral process of the contrategulum, the wide posterior stalk and the divided receptacular cluster indicate a close relationship with
|
||
<taxonomicName id="03996EC7FFF0FFE3B995FC693C61FC4E" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[1018,1145,918,943]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="species" species="panching">
|
||
<emphasis id="F6EDC956FFF0FFE3B995FC693C61FC4E" box="[1018,1145,918,943]" italics="true" pageId="20" pageNumber="411">L. panching</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="03996EC7FFF0FFE3BEC2FC693D0EFC4C" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1197,1302,920,943]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="species" species="negara" status="sp. nov.">
|
||
<emphasis id="F6EDC956FFF0FFE3BEC2FC693D0EFC4C" box="[1197,1302,920,943]" italics="true" pageId="20" pageNumber="411">L. negara</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EDDE742DFFF0FFE3BF73FC693D77FC4C" box="[1308,1391,920,943]" pageId="20" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="C4261544FFF0FFE3B950FC373BDEFB1C" blockId="20.[831,1437,966,1279]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3B950FC373BAEFC3E" bold="true" box="[831,950,966,989]" pageId="20" pageNumber="411">Variation:</emphasis>
|
||
Carapace lengths in males (n=2; values of the
|
||
<typeStatus id="1B22ABE6FFF0FFE3B9E2FC173BF3FC1E" box="[909,1003,998,1021]" pageId="20" pageNumber="411" type="holotype">holotype</typeStatus>
|
||
are taken from
|
||
<bibRefCitation id="A00868B5FFF0FFE3BEC7FC173B61FBFC" author="Platnick N. & Sedgwick W. C." journalOrPublisher="American Museum Novitates" pageId="20" pageNumber="411" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick, 1984</bibRefCitation>
|
||
: 29) 6.23-6.93, carapace widths 6.01-6.34. In females with fully developed copulatory organs (n=7) the carapace length is 5.54-7.92 and the carapace width 5.15-7.38. Variation in the shape of poreplates of the females examined is shown in
|
||
<figureCitation id="5CA209C1FFF0FFE3BF66FB763D78FB43" box="[1289,1376,1159,1184]" captionStart="Fig. 12" captionStartId="22.[151,186,1724,1745]" captionTargetBox="[151,1421,259,1654]" captionTargetId="figure@22.[148,1439,234,1682]" captionTargetPageId="22" captionText="Fig. 12. Liphistius tioman, vulval plate of three females from the type locality: specimen moulted 17. XII. 2001 (A-B); exuvia, moult 7. VI. 2003 (C-E); exuvia, moult 16. II. 2002 (F-G). (A, C, F) Entire structure, dorsal view. (B, D, G) Same, ventral view (vesicles in G omitted). (E) Receptacular cluster, ventral and slightly posterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021961/files/figure.png" pageId="20" pageNumber="411">Fig. 12</figureCitation>
|
||
. The retrolateral tibial apophyses of the left and right palp of the examined male are shown in
|
||
<figureCitation id="5CA209C1FFF0FFE3BEC5FB393D10FB02" box="[1194,1288,1224,1249]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">Fig. 11A</figureCitation>
|
||
and
|
||
<figureCitation id="5CA209C1FFF0FFE3BF2FFB393D80FB02" box="[1344,1432,1224,1249]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="20" pageNumber="411">
|
||
Fig.
|
||
<date id="B0273384FFF0FFE3BF1CFB383D80FB02" box="[1395,1432,1225,1249]" pageId="20" pageNumber="411" value="2011-01">11I</date>
|
||
</figureCitation>
|
||
, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFF0FFE3B950FAE93CAEFAB2" pageId="20" pageNumber="411" type="distribution">
|
||
<paragraph id="C4261544FFF0FFE3B950FAE93CAEFAB2" blockId="20.[831,1436,1304,1361]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3B950FAE93BCEFACC" bold="true" box="[831,982,1304,1327]" pageId="20" pageNumber="411">Distribution:</emphasis>
|
||
Known only from higher altitudes on Tioman Island (
|
||
<figureCitation id="5CA209C1FFF0FFE3B987FAC93C3FFAB2" box="[1000,1063,1336,1361]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="20" pageNumber="411">Fig. 1</figureCitation>
|
||
, locality 10).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C8346CFFFF0FFE3B950FA963D4CF7E7" pageId="20" pageNumber="411" type="biology_ecology">
|
||
<paragraph id="C4261544FFF0FFE3B950FA963D4CF7E7" blockId="20.[831,1437,1383,2053]" pageId="20" pageNumber="411">
|
||
<emphasis id="F6EDC956FFF0FFE3B950FA963B87FA9D" bold="true" box="[831,927,1383,1406]" pageId="20" pageNumber="411">Biology:</emphasis>
|
||
Spiders of this species were found under and between huge granite boulders (some of them touching each other and forming recesses) in a lush rain forest. These recesses probably correspond to the Gua Sinah or Gua Panah which is the exact
|
||
<typeStatus id="1B22ABE6FFF0FFE3BEF8FA1B3CDDF9E1" box="[1175,1221,1514,1538]" pageId="20" pageNumber="411">type</typeStatus>
|
||
locality and which were described as not being real caves (like their Malay names suggest) but just overhung caverns (
|
||
<bibRefCitation id="A00868B5FFF0FFE3BF4CF9D83C3EF981" author="Platnick N. & Sedgwick W. C." journalOrPublisher="American Museum Novitates" pageId="20" pageNumber="411" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick & Sedgwick, 1984: 29</bibRefCitation>
|
||
). Burrows were mostly dug into loamy soil at the base and between the bolders (a few small spiders had silken nests on the surface of bolders just above the ground, as described for juvenile spiders of
|
||
<taxonomicName id="03996EC7FFF0FFE3B90FF93D3BA2F900" authorityName="Platnick & Sedgwick" authorityYear="1984" box="[864,954,1740,1763]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="411" phylum="Arthropoda" rank="species" species="trang">
|
||
<emphasis id="F6EDC956FFF0FFE3B90FF93D3BA2F900" box="[864,954,1740,1763]" italics="true" pageId="20" pageNumber="411">L. trang</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="A00868B5FFF0FFE3B997F93B3CC3F900" author="Schwendinger P. J." box="[1016,1243,1738,1763]" journalOrPublisher="Natural History Bulletin of the Siam Society" pageId="20" pageNumber="411" pagination="19 - 25" part="35" refString="Schwendinger P. J. 1987. On the male of Liphistius trang (Araneae: Mesothelae), with notes on the natural history of the species. Natural History Bulletin of the Siam Society 35: 19 - 25." title="On the male of Liphistius trang (Araneae: Mesothelae), with notes on the natural history of the species" type="journal article" year="1987">Schwendinger, 1987</bibRefCitation>
|
||
: fig. 5), and they had up to eight exceptionally long (up to
|
||
<quantity id="0361B8A1FFF0FFE3BF68F91A3D54F8E0" box="[1287,1356,1771,1795]" metricMagnitude="-1" metricUnit="m" metricValue="3.4" pageId="20" pageNumber="411" unit="cm" value="34.0">34 cm</quantity>
|
||
) signal lines spread over soil or rock surface. The trapdoor of the adult male (before its final moult) was
|
||
<quantity id="0361B8A1FFF0FFE3BF76F8DD3D7CF8A0" box="[1305,1380,1836,1860]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="411" unit="cm" value="1.8">1.8 cm</quantity>
|
||
long and
|
||
<quantity id="0361B8A1FFF0FFE3B900F8BD3BA2F887" box="[879,954,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="20" pageNumber="411" unit="cm" value="3.2">3.2 cm</quantity>
|
||
wide, that of the largest female was
|
||
<quantity id="0361B8A1FFF0FFE3BF3DF8BD3D85F887" box="[1362,1437,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="411" unit="cm" value="1.9">1.9 cm</quantity>
|
||
and
|
||
<quantity id="0361B8A1FFF0FFE3B91FF89D3BA6F867" box="[880,958,1900,1924]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="20" pageNumber="411" unit="cm" value="3.2">3.2 cm</quantity>
|
||
, respectively. When collected at the end of June, four females had egg cases: one was not opened, another was empty, the remaining two were
|
||
<quantity id="0361B8A1FFF0FFE3BF48F85C3D85F827" box="[1319,1437,1965,1989]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="20" pageNumber="411" unit="cm" value="2.45" valueMax="2.7" valueMin="2.2">2.2-2.7 cm</quantity>
|
||
long,
|
||
<quantity id="0361B8A1FFF0FFE3B910F83C3BECF806" box="[895,1012,1997,2021]" metricMagnitude="-2" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="20" pageNumber="411" unit="cm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8-2.9 cm</quantity>
|
||
wide and
|
||
<quantity id="0361B8A1FFF0FFE3BE05F83C3CADF806" box="[1130,1205,1997,2021]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="20" pageNumber="411" unit="cm" value="1.9">1.9 cm</quantity>
|
||
high; they contained 3rd instar spiderlings (not collected, not counted).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |