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<document id="A9108A6081EE26E57B1318FF99691B4B" ID-DOI="10.5281/zenodo.212695" ID-GBIF-Dataset="2ec4a0e5-2653-49a3-b8d9-a130b353c02b" ID-ISSN="1175-5326" ID-Zenodo-Dep="212695" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460536255904" checkinUser="plazi" docAuthor="Larsen, Kim, Nagaoka, Ryoko &amp; Froufe, Elsa" docDate="2012" docId="36728B7AFF86FFE1FF1344DFFD756EB3" docLanguage="en" docName="zt03498p044.pdf" docOrigin="Zootaxa 3498" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptochelia parasavignyi Larsen, Nagaoka &amp; Froufe, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="CA4BF302FF87FFEBFF844065FFF16D74" masterDocTitle="Tanaidacea (Crustacea) from Macaronesia III. The shallow-water Tanaidomorpha from the Cape Verde archipelago" masterLastPageNumber="44" masterPageNumber="24" pageNumber="25" updateTime="1698316476309" updateUser="plazi">
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<mods:title id="866BCD3C93651750D5FF6E5973B61942">Tanaidacea (Crustacea) from Macaronesia III. The shallow-water Tanaidomorpha from the Cape Verde archipelago</mods:title>
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<mods:namePart id="793064FFA7DE7166CEA0575E346909E5">Larsen, Kim</mods:namePart>
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<mods:namePart id="AF3BB29DA37B3D3BBB5B6B3E93128309">Nagaoka, Ryoko</mods:namePart>
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<mods:namePart id="6D4EEEF73CDEB3213281975BB4888BCB">Froufe, Elsa</mods:namePart>
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<mods:date id="4E894A4714E67BE6E4EB1677C365A84C">2012</mods:date>
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<mods:number id="75361266380F798B64D4E41BE859D473">3498</mods:number>
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<treatment id="36728B7AFF86FFE1FF1344DFFD756EB3" ID-DOI="http://doi.org/10.5281/zenodo.6178273" ID-GBIF-Taxon="119527630" ID-Zenodo-Dep="6178273" LSID="urn:lsid:plazi:treatment:36728B7AFF86FFE1FF1344DFFD756EB3" httpUri="http://treatment.plazi.org/id/36728B7AFF86FFE1FF1344DFFD756EB3" lastPageId="10" lastPageNumber="34" pageId="1" pageNumber="25">
<subSubSection id="F6C169E7FF86FFEAFF1344DFFEE36981" pageId="1" pageNumber="25" type="nomenclature">
<paragraph id="BE643A6CFF86FFEAFF1344DFFD9F69A0" blockId="1.[151,622,1210,1269]" box="[151,622,1210,1236]" pageId="1" pageNumber="25">
<heading id="E52C8D00FF86FFEAFF1344DFFD9F69A0" bold="true" box="[151,622,1210,1236]" fontSize="11" level="1" pageId="1" pageNumber="25" reason="1">
<emphasis id="8CAFE67EFF86FFEAFF1344DFFD9F69A0" bold="true" box="[151,622,1210,1236]" pageId="1" pageNumber="25">
<taxonomicName id="79DB41EFFF86FFEAFF1344DFFE3669A0" ID-CoL="6PFHZ" box="[151,455,1210,1236]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="25" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF86FFEAFF1344DFFE3669A0" bold="true" box="[151,455,1210,1236]" italics="true" pageId="1" pageNumber="25">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="979C5B05FF86FFEAFE4A44DFFDFE69A0" box="[462,527,1210,1236]" pageId="1" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
Larsen
</emphasis>
</heading>
</paragraph>
<paragraph id="BE643A6CFF86FFEAFF1344B8FEE36981" blockId="1.[151,622,1210,1269]" box="[151,274,1245,1269]" pageId="1" pageNumber="25">
(
<figureCitation id="26E026E9FF86FFEAFF1B44B8FEF86981" box="[159,265,1245,1269]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,249,1953,1976]" captionStartId-1="4.[151,249,1953,1976]" captionStartId-2="5.[151,249,1860,1883]" captionStartId-3="6.[151,249,1953,1976]" captionStartId-4="7.[151,249,1955,1978]" captionStartId-5="8.[151,249,1723,1746]" captionTargetBox-0="[262,1284,195,1931]" captionTargetBox-1="[209,1368,202,1905]" captionTargetBox-2="[159,1428,193,1829]" captionTargetBox-3="[203,1376,203,1929]" captionTargetBox-4="[154,1425,196,1923]" captionTargetBox-5="[159,1431,201,1687]" captionTargetId-0="figure@3.[250,1336,193,1932]" captionTargetId-1="figure@4.[206,1380,193,1932]" captionTargetId-2="figure@5.[151,1436,193,1839]" captionTargetId-3="figure@6.[202,1384,193,1932]" captionTargetId-4="figure@7.[151,1436,193,1934]" captionTargetId-5="figure@8.[151,1436,193,1703]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Leptochelia parasavignyi n. sp., female. A holotype, lateral view, scale bar 1.0 mm. B E paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod." captionText-1="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." captionText-2="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." captionText-3="FIGURE 4. Leptochelia parasavignyi n. sp., primary male, paratype. A, lateral view, scale bar 1.0 mm. B, antennule. C, antenna, scale bar 0.5 mm. D, same, article 1 3, scale bar 0.1 mm." captionText-4="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." captionText-5="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri-0="https://zenodo.org/record/212696/files/figure.png" httpUri-1="https://zenodo.org/record/212697/files/figure.png" httpUri-2="https://zenodo.org/record/212698/files/figure.png" httpUri-3="https://zenodo.org/record/212699/files/figure.png" httpUri-4="https://zenodo.org/record/212700/files/figure.png" httpUri-5="https://zenodo.org/record/212701/files/figure.png" pageId="1" pageNumber="25">Figs. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F6C169E7FF86FFEAFF134540FA6D68F1" pageId="1" pageNumber="25" type="materials_examined">
<paragraph id="BE643A6CFF86FFEAFF134540FA6D68F1" blockId="1.[151,1437,1317,2025]" pageId="1" pageNumber="25">
<emphasis id="8CAFE67EFF86FFEAFF134540FE72684A" bold="true" box="[151,387,1317,1342]" pageId="1" pageNumber="25">Material examined.</emphasis>
<typeStatus id="616084CEFF86FFEAFE084540FE056849" box="[396,500,1317,1341]" pageId="1" pageNumber="25" type="holotype">Holotype</typeStatus>
non-ovigerous female. Body length
<quantity id="79239789FF86FFEAFC194540FC086849" box="[925,1017,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="25" unit="mm" value="2.3">2.3 mm</quantity>
. (Genbank # T327
<accessionNumber id="A188A78FFF86FFEAFB5F4540FABF6849" box="[1243,1358,1317,1341]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JX402116" pageId="1" pageNumber="25">JX402116</accessionNumber>
Reg #
<collectionCode id="D8CAA2A9FF86FFEAFF13452FFF296815" box="[151,216,1354,1377]" httpUri="http://grbio.org/cool/mme0-gx6r" name="Museu Municipal do Funchal" pageId="1" pageNumber="25">MMF</collectionCode>
42308), Mindelo beach, St. Vicente,
<quantity id="79239789FF86FFEAFD00452CFD376815" box="[644,710,1353,1377]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="25" unit="m" value="0.5">0.5 m</quantity>
depth, hard bottom algae.
<typeStatus id="616084CEFF86FFEAFC7E452FFB966816" box="[1018,1127,1354,1378]" pageId="1" pageNumber="25" type="paratype">Paratypes</typeStatus>
(Reg #
<collectionCode id="D8CAA2A9FF86FFEAFB42452FFAF66815" box="[1222,1287,1354,1377]" httpUri="http://grbio.org/cool/mme0-gx6r" name="Museu Municipal do Funchal" pageId="1" pageNumber="25">MMF</collectionCode>
42309): 50+ specimens of both genders and most developmental stages, same locality and Mindelo Marina, sandy bottom algae.
</paragraph>
</subSubSection>
<subSubSection id="F6C169E7FF86FFEAFF4345F4FAD86B99" pageId="1" pageNumber="25" type="diagnosis">
<paragraph id="BE643A6CFF86FFEAFF4345F4FF146BF5" blockId="1.[151,1437,1317,2025]" pageId="1" pageNumber="25">
<emphasis id="8CAFE67EFF86FFEAFF4345F4FEB068DE" bold="true" box="[199,321,1425,1450]" pageId="1" pageNumber="25">Diagnosis.</emphasis>
Female. Pereonite 6 longer than pereonite 2. Antennule proximal article just over three times as long as wide, article 2 longer than article 3. Maxilliped basis with four to five long distal setae, palp article 2 with one outer and four inner setae. Cheliped basis only 1.5 times as long as wide, carpus 1.9 times as long as wide. Pereopod 1 basis 4.5 times as long as wide, carpus longest seta 0.45 times length of propodus, dactylus 1.5 times as long as unguis, combined longer than propodus; pereopods 2 and 3 carpus as long as merus and 0.6 to 0.7 times as long as propodus; Uropod endopod with five articles, exopod uniarticulate, 0.8 times as long as first endopod article.
</paragraph>
<paragraph id="BE643A6CFF86FFEAFF4346E8FAD86B99" blockId="1.[151,1437,1317,2025]" pageId="1" pageNumber="25">Primary male. Body smaller than adult female. Antennule with eight articles (flagellum with only five articles). Antenna spiniform setae slimmer. Cheliped carpus longer and slimmer. Pereopod 1 dactylus and unguis much shorter than in female. Pereopod 6 basis without dorsal marginal flange. Uropodal exopod uniarticulate..</paragraph>
</subSubSection>
<subSubSection id="F6C169E7FF86FFEAFF43469FFB976A65" box="[199,1126,1785,1810]" pageId="1" pageNumber="25" type="etymology">
<paragraph id="BE643A6CFF86FFEAFF43469FFB976A65" blockId="1.[151,1437,1317,2025]" box="[199,1126,1785,1810]" pageId="1" pageNumber="25">
<emphasis id="8CAFE67EFF86FFEAFF43469FFEBF6A66" bold="true" box="[199,334,1786,1810]" pageId="1" pageNumber="25">Etymology.</emphasis>
Named after the morphological similarities with
<taxonomicName id="79DB41EFFF86FFEAFCF2469CFBAF6A65" box="[886,1118,1785,1809]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="1" pageNumber="25" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="8CAFE67EFF86FFEAFCF2469CFBAF6A65" box="[886,1118,1785,1809]" italics="true" pageId="1" pageNumber="25">Leptochelia savignyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F6C169E7FF86FFE2FF434778FF276B94" lastPageId="9" lastPageNumber="33" pageId="1" pageNumber="25" type="description">
<paragraph id="BE643A6CFF86FFEAFF434778FA9A6A42" blockId="1.[151,1437,1317,2025]" box="[199,1387,1821,1846]" pageId="1" pageNumber="25">
<emphasis id="8CAFE67EFF86FFEAFF434778FEA86A42" bold="true" box="[199,345,1821,1846]" pageId="1" pageNumber="25">Description.</emphasis>
Female. Body from non-ovigerous
<typeStatus id="616084CEFF86FFEAFD6D4778FCBF6A41" box="[745,846,1821,1845]" pageId="1" pageNumber="25" type="holotype">holotype</typeStatus>
, appendages from dissected ovigerous
<typeStatus id="616084CEFF86FFEAFA80477BFA996A42" box="[1284,1384,1822,1846]" pageId="1" pageNumber="25" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="BE643A6CFF86FFE9FF434724FE7D6DA0" blockId="1.[151,1437,1317,2025]" lastBlockId="2.[151,1437,151,1976]" lastPageId="2" lastPageNumber="26" pageId="1" pageNumber="25">
<emphasis id="8CAFE67EFF86FFEAFF434724FEF06A2D" box="[199,257,1857,1881]" italics="true" pageId="1" pageNumber="25">Body</emphasis>
(
<figureCitation id="26E026E9FF86FFEAFE8B4724FEBE6A2D" box="[271,335,1857,1881]" captionStart="FIGURE 1" captionStartId="3.[151,249,1953,1976]" captionTargetBox="[262,1284,195,1931]" captionTargetId="figure@3.[250,1336,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. Leptochelia parasavignyi n. sp., female. A holotype, lateral view, scale bar 1.0 mm. B E paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod." httpUri="https://zenodo.org/record/212696/files/figure.png" pageId="1" pageNumber="25">Fig. 1</figureCitation>
A). Elongate, slender, 8.5 times as long as wide.
<emphasis id="8CAFE67EFF86FFEAFCF64727FC116A2D" box="[882,992,1858,1881]" italics="true" pageId="1" pageNumber="25">Carapace</emphasis>
subrectangular, almost twice as long as wide, longer than pereonites 1 and 2 combined, with slight rounded rostrum, naked.
<emphasis id="8CAFE67EFF86FFEAFBD24703FB3C6A09" box="[1110,1229,1894,1917]" italics="true" pageId="1" pageNumber="25">Pereonites</emphasis>
, each bearing one short, simple anterolateral setae. Pereonite 1 shortest, pereonites 2, 3, 4, and 5 progressively longer; pereonite 6 longer than pereonite 2.
<emphasis id="8CAFE67EFF86FFEAFE2947C8FE1F6AB1" box="[429,494,1965,1989]" italics="true" pageId="1" pageNumber="25">Pleon</emphasis>
slightly wider than pereon, about one-quarter of total length.
<emphasis id="8CAFE67EFF86FFEAFB2947C8FAE46AB1" box="[1197,1301,1965,1989]" italics="true" pageId="1" pageNumber="25">Pleonites</emphasis>
about three times as wide as long; anterior three pleonites with one- posterior two pleonites with three short, simple mediolateral setae.
<emphasis id="8CAFE67EFF85FFE9FEF340FDFE1C6DC4" box="[375,493,152,176]" italics="true" pageId="2" pageNumber="26">Pleotelson</emphasis>
only marginally longer than pleonite 5, with one simple lateral and two pairs of terminal simple setae.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF434085FB726C10" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF434085FEC86D8C" box="[199,313,224,248]" italics="true" pageId="2" pageNumber="26">Antennule</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FECC40BAFE766D83" box="[328,391,223,247]" captionStart="FIGURE 1" captionStartId="3.[151,249,1953,1976]" captionTargetBox="[262,1284,195,1931]" captionTargetId="figure@3.[250,1336,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. Leptochelia parasavignyi n. sp., female. A holotype, lateral view, scale bar 1.0 mm. B E paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod." httpUri="https://zenodo.org/record/212696/files/figure.png" pageId="2" pageNumber="26">Fig. 1</figureCitation>
B). Article 1 just over three times as long as wide, with medial and distal setulose and simple setae of which one is longer than article 2. Article 2 longer than article 3, with two distal setae of which one is longer than article 3. Article 3 marginally longer than one quarter of article 1, with small ventrodistal process and two simple distal setae. Article 4 minute, with four simple distal setae and one aesthetasc.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF434115FBE96F63" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF434115FED46CFC" box="[199,293,368,392]" italics="true" pageId="2" pageNumber="26">Antenna</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FEB0410AFE856CF3" box="[308,372,367,391]" captionStart="FIGURE 1" captionStartId="3.[151,249,1953,1976]" captionTargetBox="[262,1284,195,1931]" captionTargetId="figure@3.[250,1336,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. Leptochelia parasavignyi n. sp., female. A holotype, lateral view, scale bar 1.0 mm. B E paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod." httpUri="https://zenodo.org/record/212696/files/figure.png" pageId="2" pageNumber="26">Fig. 1</figureCitation>
C). Article 1 with small ventrodistal projection and one ventrodistal seta. Article 2 longer than article 1, with conspicuous single ventrodistal and dorsodistal spiniform setae arising from small projections. Article 3 shorter than article 2, with dorsodistal spiniform seta. Article 4 longest, less than twice as long as article 2, with two long simple and three setulated distal setae. Article 5 slender, less than half as long as article 4, with two simple distal setae. Article 6 minute with three long and one minute distal setae.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF434241FDB86EB3" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF434241FE436F48" box="[199,434,548,572]" italics="true" pageId="2" pageNumber="26">Mouthparts. Labrum</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FE464241FDF46F48" box="[450,517,548,572]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
A) with conspicuous lateral projections, setulose.
<emphasis id="8CAFE67EFF85FFE9FBBF4241FB266F48" box="[1083,1239,548,572]" italics="true" pageId="2" pageNumber="26">Left mandible</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FB6C4241FADD6F48" box="[1256,1324,548,572]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
B) incisor with one distal and two apical denticles,
<emphasis id="8CAFE67EFF85FFE9FD05422DFCDC6F14" box="[641,813,584,608]" italics="true" pageId="2" pageNumber="26">lacinia mobilis</emphasis>
with wide, crenulate distal edge and fine setulation/ serration on proximal edge, molar with strongly chitinized crushing surface with small pointy teeth.
<emphasis id="8CAFE67EFF85FFE9FB744209FA6A6FF0" box="[1264,1435,620,644]" italics="true" pageId="2" pageNumber="26">Right mandible</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FF1B42EAFF156FD3" box="[159,228,655,679]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
C) incisor bifurcate, with small single dorsal crenulation, and fine setulation/serration on proximal edge, molar as in left mandible.
<emphasis id="8CAFE67EFF85FFE9FE3E42D1FDE16FB8" box="[442,528,692,716]" italics="true" pageId="2" pageNumber="26">Labium</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FD9A42D1FD916FB8" box="[542,608,692,716]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
D) with two lobes, distal margin with multiple setae and setules.
<emphasis id="8CAFE67EFF85FFE9FAB542D1FA6D6FB8" box="[1329,1436,692,716]" italics="true" pageId="2" pageNumber="26">Maxillule</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FF1B42B2FF156F9B" box="[159,228,727,751]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
E) endite with ten spiniform distal setae and distally setulose outer- and sparsely medial inner setulation; palp with two long distal setae.
<emphasis id="8CAFE67EFF85FFE9FE7A4299FDA46E60" box="[510,597,764,788]" italics="true" pageId="2" pageNumber="26">Maxilla</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FDE14299FD596E60" box="[613,680,764,788]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
F) simple, ovoid, naked.
<emphasis id="8CAFE67EFF85FFE9FC464299FBCA6E60" box="[962,1083,764,788]" italics="true" pageId="2" pageNumber="26">Maxilliped</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FBCF4299FB7F6E60" box="[1099,1166,764,788]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
G) Basis with four right and five left long (exceeding distal margin of palp article 2) setae; endites distally with single outer seta and three flat spiniform setae, inner one shorter than outer two, lateral corners finely setulose; palp article 1 naked; article 2 with outer-distal seta and four inner distal setae, inner proximal margin finely setulose; article 3 with eight inner setae; article 4 with ten inner and one outer subdistal setae.
<emphasis id="8CAFE67EFF85FFE9FCCD43E9FC416ED0" box="[841,944,908,932]" italics="true" pageId="2" pageNumber="26">Epignath</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FC4543E9FBF46ED0" box="[961,1029,908,932]" captionStart="FIGURE 2" captionStartId="4.[151,249,1953,1976]" captionTargetBox="[209,1368,202,1905]" captionTargetId="figure@4.[206,1380,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Leptochelia parasavignyi n. sp., female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="2" pageNumber="26">Fig. 2</figureCitation>
H) apparently naked but the tip has broken off so this might be an artefact.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF4343B1FDA069B0" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF4343B1FEDD6E98" box="[199,300,980,1004]" italics="true" pageId="2" pageNumber="26">Cheliped</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FEB943B1FE716E98" box="[317,384,980,1004]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="2" pageNumber="26">Fig. 3</figureCitation>
A) attached to carapace via a large triangular sclerite. Basis only 1.5 times as long as wide, with sub-dorsodistal seta. Merus triangular, with four ventral setae. Carpus almost twice as long as wide, with two dorsoproximal and one dorsodistal setae and three ventrodistal setae. Propodus, inclusive fixed finger, almost twice as long as wide, with inner row of six pinnate setae of which the most ventral is longer than the others, with one setae at dactylus insertion. Fixed finger shorter than body of propodus, with chitinised ridge overlapping unguis, with three setae on cutting edge and four ventral setae. Dactylus with dorsoproximal seta, with process on cutting edge but without small spiniform setae.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF4344B5FBCF68E8" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF4344B5FEB6699C" box="[199,327,1232,1256]" italics="true" pageId="2" pageNumber="26">Pereopod 1</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FED244B5FE66699C" box="[342,407,1232,1256]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="2" pageNumber="26">Fig. 3</figureCitation>
B) longer than other pereopods. Coxa with seta. Basis slender, shorter than combined length of propodus, dactylus, and unguis, with one simple, and one setulose dorsoproximal setae. Ischium with one ventral seta. Merus longer than carpus, with three small distal setae. Carpus about one-third as long as basis, with three short and two longer distal setae. Propodus shorter than carpus and merus combined, with one ventral subdistal seta and three dorsal subdistal setae, with small dorsal spine just proximal to dorsal setae and distal spine. Dactylus with dorsoproximal seta, combined with unguis as long as propodus, longer than unguis.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF4345CDFE146BEC" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF4345CDFEB868B4" box="[199,329,1448,1472]" italics="true" pageId="2" pageNumber="26">Pereopod 2</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FEDD45CDFE6D68B4" box="[345,412,1448,1472]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="2" pageNumber="26">Fig. 3</figureCitation>
C) shorter than pereopod 1. Coxa with seta. Basis wider than that of pereopod 1, with two setulose and one simple dorsoproximal setae. Ischium with two ventral setae. Merus as long as carpus, combined longer than propodus, with one simple and one spiniform ventrodistal setae. Carpus with one simple and one spiniform ventrodistal setae and scales, with additional two dorsodistal setae. Propodus longer than half of basis, with one small spiniform ventral subdistal seta and three dorsal/medial subdistal setae, with small dorsal spine just proximal to dorsal seta and small distal spine. Dactylus with proximal spines combined with unguis shorter than propodus, longer than unguis.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF4346C1FCB06B94" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF4346C1FEBC6BC8" box="[199,333,1700,1724]" italics="true" pageId="2" pageNumber="26">Pereopod 3</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FEE546C1FE596BC8" box="[353,424,1700,1724]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="2" pageNumber="26">Fig. 3</figureCitation>
D) similar to pereopod 2 except: basis with just one dorsoproximal setulose seta and propodus with only two dorsal subdistal seta and large spine.
</paragraph>
<paragraph id="BE643A6CFF85FFE9FF434689FAAC6ACC" blockId="2.[151,1437,151,1976]" pageId="2" pageNumber="26">
<emphasis id="8CAFE67EFF85FFE9FF434689FEB96A70" box="[199,328,1772,1796]" italics="true" pageId="2" pageNumber="26">Pereopod 4</emphasis>
(
<figureCitation id="26E026E9FF85FFE9FEDC4689FE6B6A70" box="[344,410,1772,1796]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="2" pageNumber="26">Fig. 3</figureCitation>
E) coxa fused with pereonite. Basis stout, more than twice as long as wide and longer than rest of pereopod, with one ventroproximal setulose seta. Ischium with two ventral setae. Merus as long as carpus, with paired ventrodistal spiniform setae. Carpus with two dorsodistal setae, one inner and two outer curved spiniform setae. Propodus about as long as merus, with three stout and one spiniform pinnate dorsodistal setae and small spine proximal from dorsal setae, with ventrodistal pair of spiniform setae. Dactylus curved and incompletely fused with unguis, distally with small spines. Unguis combined with dactylus about half as long as propodus.
</paragraph>
<caption id="EAA46AE4FF84FFE8FF1347C4FD486AA2" httpUri="https://zenodo.org/record/212696/files/figure.png" pageId="3" pageNumber="27" targetBox="[262,1284,195,1931]" targetPageId="3">
<paragraph id="BE643A6CFF84FFE8FF1347C4FD486AA2" blockId="3.[151,1436,1953,2006]" pageId="3" pageNumber="27">
<emphasis id="8CAFE67EFF84FFE8FF1347C4FEE26AC2" bold="true" box="[151,275,1953,1976]" pageId="3" pageNumber="27">FIGURE 1.</emphasis>
<taxonomicName id="79DB41EFFF84FFE8FE9F47C4FDEF6AC3" box="[283,542,1953,1975]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="3" pageNumber="27" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF84FFE8FE9F47C4FDEF6AC3" box="[283,542,1953,1975]" italics="true" pageId="3" pageNumber="27">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<emphasis id="8CAFE67EFF84FFE8FDA147C7FDAE6AC3" bold="true" box="[549,607,1954,1975]" pageId="3" pageNumber="27">
<taxonomicNameLabel id="979C5B05FF84FFE8FDA147C7FDAE6AC3" box="[549,607,1954,1975]" pageId="3" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A holotype, lateral view, scale bar 1.0 mm. BE paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod.
</paragraph>
</caption>
<caption id="EAA46AE4FF83FFEFFF1347C4FCBF6AA2" httpUri="https://zenodo.org/record/212697/files/figure.png" pageId="4" pageNumber="28" targetBox="[209,1368,202,1905]" targetPageId="4">
<paragraph id="BE643A6CFF83FFEFFF1347C4FCBF6AA2" blockId="4.[151,1436,1953,2006]" pageId="4" pageNumber="28">
<emphasis id="8CAFE67EFF83FFEFFF1347C4FEE26AC2" bold="true" box="[151,275,1953,1976]" pageId="4" pageNumber="28">FIGURE 2.</emphasis>
<taxonomicName id="79DB41EFFF83FFEFFE9F47C4FDEF6AC3" box="[283,542,1953,1975]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF83FFEFFE9F47C4FDEF6AC3" box="[283,542,1953,1975]" italics="true" pageId="4" pageNumber="28">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<emphasis id="8CAFE67EFF83FFEFFDA247C7FD916AC3" bold="true" box="[550,608,1954,1975]" pageId="4" pageNumber="28">
<taxonomicNameLabel id="979C5B05FF83FFEFFDA247C7FD916AC3" box="[550,608,1954,1975]" pageId="4" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, paratype. A, labrum. B, left mandible. C, right mandible. D, labium. E, maxillule. F, maxilla. G, maxilliped. H, epignath. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="EAA46AE4FF82FFEEFF134721FD0E6A0D" httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="5" pageNumber="29" targetBox="[159,1428,193,1829]" targetPageId="5">
<paragraph id="BE643A6CFF82FFEEFF134721FD0E6A0D" blockId="5.[151,1436,1860,1913]" pageId="5" pageNumber="29">
<emphasis id="8CAFE67EFF82FFEEFF134721FEE26A2E" bold="true" box="[151,275,1860,1883]" pageId="5" pageNumber="29">FIGURE 3.</emphasis>
<taxonomicName id="79DB41EFFF82FFEEFE9F4720FDEF6A2F" box="[283,542,1861,1883]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="5" pageNumber="29" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF82FFEEFE9F4720FDEF6A2F" box="[283,542,1861,1883]" italics="true" pageId="5" pageNumber="29">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<emphasis id="8CAFE67EFF82FFEEFDA14720FDAE6A2E" bold="true" box="[549,607,1861,1882]" pageId="5" pageNumber="29">
<taxonomicNameLabel id="979C5B05FF82FFEEFDA14720FDAE6A2E" box="[549,607,1861,1882]" pageId="5" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="EAA46AE4FF81FFEDFF1347C4FCC96AA2" httpUri="https://zenodo.org/record/212699/files/figure.png" pageId="6" pageNumber="30" targetBox="[203,1376,203,1929]" targetPageId="6">
<paragraph id="BE643A6CFF81FFEDFF1347C4FCC96AA2" blockId="6.[151,1435,1953,2006]" pageId="6" pageNumber="30">
<emphasis id="8CAFE67EFF81FFEDFF1347C4FEE76AC2" bold="true" box="[151,278,1953,1976]" pageId="6" pageNumber="30">FIGURE 4.</emphasis>
<taxonomicName id="79DB41EFFF81FFEDFE9B47C4FDD46AC3" box="[287,549,1953,1975]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="6" pageNumber="30" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF81FFEDFE9B47C4FDD46AC3" box="[287,549,1953,1975]" italics="true" pageId="6" pageNumber="30">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<emphasis id="8CAFE67EFF81FFEDFDAB47C7FD9A6AC3" bold="true" box="[559,619,1954,1975]" pageId="6" pageNumber="30">
<taxonomicNameLabel id="979C5B05FF81FFEDFDAB47C7FD9A6AC3" box="[559,619,1954,1975]" pageId="6" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, primary male, paratype. A, lateral view, scale bar 1.0 mm. B, antennule. C, antenna, scale bar 0.5 mm. D, same, article 13, scale bar 0.1 mm.
</paragraph>
</caption>
<caption id="EAA46AE4FF80FFECFF1347C6FB1A6AAC" httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="7" pageNumber="31" targetBox="[154,1425,196,1923]" targetPageId="7">
<paragraph id="BE643A6CFF80FFECFF1347C6FB1A6AAC" blockId="7.[151,1436,1955,2008]" pageId="7" pageNumber="31">
<emphasis id="8CAFE67EFF80FFECFF1347C6FEE36ACD" bold="true" box="[151,274,1955,1978]" pageId="7" pageNumber="31">FIGURE 5.</emphasis>
<taxonomicName id="79DB41EFFF80FFECFE9D47C6FDEA6ACD" box="[281,539,1955,1977]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="7" pageNumber="31" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF80FFECFE9D47C6FDEA6ACD" box="[281,539,1955,1977]" italics="true" pageId="7" pageNumber="31">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<emphasis id="8CAFE67EFF80FFECFDA647C1FDAA6ACD" bold="true" box="[546,603,1956,1977]" pageId="7" pageNumber="31">
<taxonomicNameLabel id="979C5B05FF80FFECFDA647C1FDAA6ACD" box="[546,603,1956,1977]" pageId="7" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, primary male, paratype. AD &amp; FH scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod.
</paragraph>
</caption>
<caption id="EAA46AE4FF8FFFE3FF1346DEFD0E6B85" httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="8" pageNumber="32" targetBox="[159,1431,201,1687]" targetPageId="8">
<paragraph id="BE643A6CFF8FFFE3FF1346DEFD0E6B85" blockId="8.[151,1436,1723,1777]" pageId="8" pageNumber="32">
<emphasis id="8CAFE67EFF8FFFE3FF1346DEFEE66BA5" bold="true" box="[151,279,1723,1746]" pageId="8" pageNumber="32">FIGURE 6.</emphasis>
<taxonomicName id="79DB41EFFF8FFFE3FEA546D9FDD96BA6" box="[289,552,1724,1746]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="parasavignyi" status="sp. nov.">
<emphasis id="8CAFE67EFF8FFFE3FEA546D9FDD96BA6" box="[289,552,1724,1746]" italics="true" pageId="8" pageNumber="32">Leptochelia parasavignyi</emphasis>
</taxonomicName>
<emphasis id="8CAFE67EFF8FFFE3FDB746D9FD816BA6" bold="true" box="[563,624,1724,1746]" pageId="8" pageNumber="32">
<taxonomicNameLabel id="979C5B05FF8FFFE3FDB746D9FD816BA6" box="[563,624,1724,1746]" pageId="8" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="BE643A6CFF8FFFE3FF434778FD6C6A2D" blockId="8.[151,1436,1821,2026]" pageId="8" pageNumber="32">
<emphasis id="8CAFE67EFF8FFFE3FF434778FEBB6A41" box="[199,330,1821,1845]" italics="true" pageId="8" pageNumber="32">Pereopod 5</emphasis>
(
<figureCitation id="26E026E9FF8FFFE3FED94778FE506A41" box="[349,417,1821,1845]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="8" pageNumber="32">Fig. 3</figureCitation>
F) similar to pereopod 4 except: basis with one simple and two setulose ventroproximal setae and propodus with several dorsal spines.
</paragraph>
<paragraph id="BE643A6CFF8FFFE3FF434703FBF56AD5" blockId="8.[151,1436,1821,2026]" pageId="8" pageNumber="32">
<emphasis id="8CAFE67EFF8FFFE3FF434703FEBB6A0A" box="[199,330,1894,1918]" italics="true" pageId="8" pageNumber="32">Pereopod 6</emphasis>
(
<figureCitation id="26E026E9FF8FFFE3FEDF4700FE6E6A09" box="[347,415,1893,1917]" captionStart="FIGURE 3" captionStartId="5.[151,249,1860,1883]" captionTargetBox="[159,1428,193,1829]" captionTargetId="figure@5.[151,1436,193,1839]" captionTargetPageId="5" captionText="FIGURE 3. Leptochelia parasavignyi n. sp., female. paratype. A, cheliped. B, pereopod 1. C, pereopod 2. D, pereopod 3. E, Pereopod 4. F, pereopod 5. G, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212698/files/figure.png" pageId="8" pageNumber="32">Fig. 3</figureCitation>
G) as pereopod 4 except: basis with one setulose dorsoproximal seta. Propodus with five stiff pinnate and two longer simple dorsodistal setae and several dorsal spines.
</paragraph>
<paragraph id="BE643A6CFF8FFFE2FF4347CBFB476DA0" blockId="8.[151,1436,1821,2026]" lastBlockId="9.[151,1437,151,2012]" lastPageId="9" lastPageNumber="33" pageId="8" pageNumber="32">
<emphasis id="8CAFE67EFF8FFFE3FF4347CBFEC16AB2" box="[199,304,1966,1990]" italics="true" pageId="8" pageNumber="32">Pleopods</emphasis>
(
<figureCitation id="26E026E9FF8FFFE3FEBA47C8FE8D6AB1" box="[318,380,1965,1989]" captionStart="FIGURE 1" captionStartId="3.[151,249,1953,1976]" captionTargetBox="[262,1284,195,1931]" captionTargetId="figure@3.[250,1336,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. Leptochelia parasavignyi n. sp., female. A holotype, lateral view, scale bar 1.0 mm. B E paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod." httpUri="https://zenodo.org/record/212696/files/figure.png" pageId="8" pageNumber="32">Fig. 1</figureCitation>
D) all five pairs similar. Basal article with one circumplumose inner seta. Endopod with single inner plumose seta and 18 outer plumose setae, distal seta shorter and thicker and with curved serrated apex, gap between most proximal seta and other outer setae. Exopod with 25 outer plumose setae, most proximal seta circumplumose, seems to arise from a process and is separated by a gap from remaining setae.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF434085FAE76C68" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434085FEEC6D8C" box="[199,285,224,248]" italics="true" pageId="9" pageNumber="33">Uropod</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEA840BAFE986D83" box="[300,361,223,247]" captionStart="FIGURE 1" captionStartId="3.[151,249,1953,1976]" captionTargetBox="[262,1284,195,1931]" captionTargetId="figure@3.[250,1336,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. Leptochelia parasavignyi n. sp., female. A holotype, lateral view, scale bar 1.0 mm. B E paratype, scale bar 0.5 mm. B, antennule. C, antenna. D, pleopod. E, uropod." httpUri="https://zenodo.org/record/212696/files/figure.png" pageId="9" pageNumber="33">Fig. 1</figureCitation>
E) basal article naked. Exopod uniarticulate, more than half as long as first endopod article, with one medial and one distal setae. Endopod with five articles, distal three articles each with a single seta.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF434142FE0F6C4B" blockId="9.[151,1437,151,2012]" box="[199,510,295,320]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434142FEA86C34" bold="true" box="[199,345,295,320]" pageId="9" pageNumber="33">Description.</emphasis>
Primary male.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF434129FA876F48" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434129FEF06C10" box="[199,257,332,356]" italics="true" pageId="9" pageNumber="33">Body</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FE9F4129FE986C10" box="[283,361,332,356]" captionStart="FIGURE 4" captionStartId="6.[151,249,1953,1976]" captionTargetBox="[203,1376,203,1929]" captionTargetId="figure@6.[202,1384,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Leptochelia parasavignyi n. sp., primary male, paratype. A, lateral view, scale bar 1.0 mm. B, antennule. C, antenna, scale bar 0.5 mm. D, same, article 1 3, scale bar 0.1 mm." httpUri="https://zenodo.org/record/212699/files/figure.png" pageId="9" pageNumber="33">Fig. 4</figureCitation>
A) shorter than female with relative longer pleon and shorter pereon.
<emphasis id="8CAFE67EFF8EFFE2FB704129FA6D6C10" box="[1268,1436,332,356]" italics="true" pageId="9" pageNumber="33">Cephalothorax</emphasis>
subrectangular, longer than wide, almost as long as pereonites 1 to 3 combined, eyes larger than those of female.
<emphasis id="8CAFE67EFF8EFFE2FF1341F0FEFF6CD8" box="[151,270,405,428]" italics="true" pageId="9" pageNumber="33">Pereonites</emphasis>
, bearing one short, simple lateral seta. Pereonite 1 shortest. Pereonites 2 and 3 each 1.3 times as long as pereonite 1. Pereonites 4 and 5 longest, almost twice as long as pereonite 1. Pereonite 6 only marginally longer than pereonite 1.
<emphasis id="8CAFE67EFF8EFFE2FEAD41B9FE9A6C80" box="[297,363,476,500]" italics="true" pageId="9" pageNumber="33">Pleon</emphasis>
wider than pereon. Pleonites about four times as wide as long, each with one short, simple mediolateral seta. Pleotelson (
<figureCitation id="26E026E9FF8EFFE2FE7F419AFDB26F63" box="[507,579,511,535]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
H) short, 1.2 times as long as pleonite 5, twice as wide as long, with mediolateral simple seta and two pairs of simple apical setae of unequal length and one pair of sub apical setae.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF43422DFBEC6FB8" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF43422DFEC86F14" box="[199,313,584,608]" italics="true" pageId="9" pageNumber="33">Antennule</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEC84222FE626F2B" box="[332,403,583,607]" captionStart="FIGURE 4" captionStartId="6.[151,249,1953,1976]" captionTargetBox="[203,1376,203,1929]" captionTargetId="figure@6.[202,1384,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Leptochelia parasavignyi n. sp., primary male, paratype. A, lateral view, scale bar 1.0 mm. B, antennule. C, antenna, scale bar 0.5 mm. D, same, article 1 3, scale bar 0.1 mm." httpUri="https://zenodo.org/record/212699/files/figure.png" pageId="9" pageNumber="33">Fig. 4</figureCitation>
B) consists of eight articles. Article 1 only about three times as long as wide, with one dorsodistal simple seta. Article 2 longer than half of article 1, with one ventrodistal simple seta. Article 3 about half as long as article 2, with three simple distal setae. Article 47 serially repeating, all with aesthestascs (many more than illustrated). Article minute, with three long and one tiny simple distal setae.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF4342BDFE9D6EF4" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF4342BDFED46F84" box="[199,293,728,752]" italics="true" pageId="9" pageNumber="33">Antenna</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEB242B2FE8C6F9B" box="[310,381,727,751]" captionStart="FIGURE 4" captionStartId="6.[151,249,1953,1976]" captionTargetBox="[203,1376,203,1929]" captionTargetId="figure@6.[202,1384,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Leptochelia parasavignyi n. sp., primary male, paratype. A, lateral view, scale bar 1.0 mm. B, antennule. C, antenna, scale bar 0.5 mm. D, same, article 1 3, scale bar 0.1 mm." httpUri="https://zenodo.org/record/212699/files/figure.png" pageId="9" pageNumber="33">Fig. 4</figureCitation>
C,D) Article 1 square and with one tiny simple seta. Article 2 with ventro- and dorsodistal spiniform setae (thinner than in female). Article 3 about as long as article 2, with dorsodistal spiniform seta (thinner than in female). Article 4 twice as long as article 3, with three long simple and one setulose distal setae. Article 5 slender, shorter than article 4, with one long simple distal seta. Article 6 minute with four long and two shorter simple distal setae.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF4343E9FBE66E98" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF4343E9FEBB6ED0" box="[199,330,908,932]" italics="true" pageId="9" pageNumber="33">Mouthparts</emphasis>
reduced.
<emphasis id="8CAFE67EFF8EFFE2FE3843E9FDE56ED0" box="[444,532,908,932]" italics="true" pageId="9" pageNumber="33">Labrum</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FDA243E9FD986ED0" box="[550,617,908,932]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
A) prominent but naked, fused to clypeus.
<emphasis id="8CAFE67EFF8EFFE2FBDD43E9FB3E6ED0" box="[1113,1231,908,932]" italics="true" pageId="9" pageNumber="33">Mandibles</emphasis>
absent.
<emphasis id="8CAFE67EFF8EFFE2FAB543E9FA6D6ED0" box="[1329,1436,908,932]" italics="true" pageId="9" pageNumber="33">Maxillule</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FF1B43CAFF156EB3" box="[159,228,943,967]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
B) endite absent, basis naked, palp pseudo-uniarticulate.
<emphasis id="8CAFE67EFF8EFFE2FCF043D5FC1C6EBC" box="[884,1005,944,968]" italics="true" pageId="9" pageNumber="33">Maxilliped</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FC7B43CAFBB56EB3" box="[1023,1092,943,967]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
C) basis naked, endite absent, palp pseudoarticulate.
<emphasis id="8CAFE67EFF8EFFE2FE1043B1FE0A6E98" box="[404,507,980,1004]" italics="true" pageId="9" pageNumber="33">Epignath</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FD8E43B1FDBA6E98" box="[522,587,980,1004]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
D) fully developed with distal setulation.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF43439DFB9B699C" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF43439DFEDD6964" box="[199,300,1016,1040]" italics="true" pageId="9" pageNumber="33">Cheliped</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEBF439DFE8C6964" box="[315,381,1016,1040]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
E,F) slender, about half as long as body, attached to body via triangular sclerite bearing a seta. Basis less than twice as long as wide, with single, outer sub-dorsodistal seta. Merus triangular, with three ventral setae. Carpus longer than propodus inclusive fixed finger, with two dorsoproximal and one dorsodistal setae, with two longer and one short ventro-subdistal setae. Propodus including fixed finger as long as basis, with one setae at dactyls insertion and inner row of six pinnate setae of which the most ventral is longer than others. Fixed finger with two small projections on cutting edge, not overlapping unguis, with three setae on cutting edge and four ventral setae. Dactylus with dorsoproximal seta and inner row of small spiniform setae.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF434491FB2D6878" blockId="9.[151,1437,151,2012]" box="[199,1244,1268,1292]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434491FEB96878" box="[199,328,1268,1292]" italics="true" pageId="9" pageNumber="33">Pereopod 1</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FED34491FE686878" box="[343,409,1268,1292]" captionStart="FIGURE 6" captionStartId="8.[151,249,1723,1746]" captionTargetBox="[159,1431,201,1687]" captionTargetId="figure@8.[151,1436,193,1703]" captionTargetPageId="8" captionText="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="9" pageNumber="33">Fig. 6</figureCitation>
A) as in female except: dactylus/unguis much shorter relative to propodus.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF43457DFDB56820" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
Pereopod 2 (
<figureCitation id="26E026E9FF8EFFE2FED0457DFE676844" box="[340,406,1304,1328]" captionStart="FIGURE 6" captionStartId="8.[151,249,1723,1746]" captionTargetBox="[159,1431,201,1687]" captionTargetId="figure@8.[151,1436,193,1703]" captionTargetPageId="8" captionText="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="9" pageNumber="33">Fig. 6</figureCitation>
B) as in female except: carpus with two small spiniform setae. Dactylus ventral spine absent small spines present on dorsal margin.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF434505FE4C68E8" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434505FEBB680C" box="[199,330,1376,1400]" italics="true" pageId="9" pageNumber="33">Pereopod 3</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEDF4505FE51680C" box="[347,416,1376,1400]" captionStart="FIGURE 6" captionStartId="8.[151,249,1723,1746]" captionTargetBox="[159,1431,201,1687]" captionTargetId="figure@8.[151,1436,193,1703]" captionTargetPageId="8" captionText="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="9" pageNumber="33">Fig. 6</figureCitation>
C) similar to pereopod 2 except: merus with spiniform ventral setae. Propodus with only two dorsal subdistal setae.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF4345CDFAAF68B4" blockId="9.[151,1437,151,2012]" box="[199,1374,1448,1472]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF4345CDFEB968B4" box="[199,328,1448,1472]" italics="true" pageId="9" pageNumber="33">Pereopod 4</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FED345CDFE6868B4" box="[343,409,1448,1472]" captionStart="FIGURE 6" captionStartId="8.[151,249,1723,1746]" captionTargetBox="[159,1431,201,1687]" captionTargetId="figure@8.[151,1436,193,1703]" captionTargetPageId="8" captionText="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="9" pageNumber="33">Fig. 6</figureCitation>
D) as in female except: basis slimmer. Merus to dactylus with scales or minute spines.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF4345A9FB4D6890" blockId="9.[151,1437,151,2012]" box="[199,1212,1484,1508]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF4345A9FEB96890" box="[199,328,1484,1508]" italics="true" pageId="9" pageNumber="33">Pereopod 5</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FED345A9FE686890" box="[343,409,1484,1508]" captionStart="FIGURE 6" captionStartId="8.[151,249,1723,1746]" captionTargetBox="[159,1431,201,1687]" captionTargetId="figure@8.[151,1436,193,1703]" captionTargetPageId="8" captionText="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="9" pageNumber="33">Fig. 6</figureCitation>
E) as pereopod 4 except: basis with additional ventromedial simple seta
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF434595FCD76B58" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434595FEBD6B7C" box="[199,332,1520,1544]" italics="true" pageId="9" pageNumber="33">Pereopod 6</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEDB4595FE576B7C" box="[351,422,1520,1544]" captionStart="FIGURE 6" captionStartId="8.[151,249,1723,1746]" captionTargetBox="[159,1431,201,1687]" captionTargetId="figure@8.[151,1436,193,1703]" captionTargetPageId="8" captionText="FIGURE 6. Leptochelia parasavignyi n. sp., primary male, paratype. A, pereopod 1. B, pereopod 2. C, pereopod 3. D, Pereopod 4. E, pereopod 5. F, pereopod 6. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/212701/files/figure.png" pageId="9" pageNumber="33">Fig. 6</figureCitation>
F) as pereopod 4 except: propodus with five pinnate and one simple dorsodistal setae. Dactylus with more demarcated spines at unguis insertion.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF43465DFED36B00" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF43465DFEC16B24" box="[199,304,1592,1616]" italics="true" pageId="9" pageNumber="33">Pleopods</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEBB465DFE706B24" box="[319,385,1592,1616]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
G) all similar. As in female except: with fewer but longer plumose setae not separated by gap on endopod.
</paragraph>
<paragraph id="BE643A6CFF8EFFE2FF4346E5FF276B94" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF4346E5FEEC6BEC" box="[199,285,1664,1688]" italics="true" pageId="9" pageNumber="33">Uropod</emphasis>
(
<figureCitation id="26E026E9FF8EFFE2FEA846E5FE9D6BEC" box="[300,364,1664,1688]" captionStart="FIGURE 5" captionStartId="7.[151,249,1955,1978]" captionTargetBox="[154,1425,196,1923]" captionTargetId="figure@7.[151,1436,193,1934]" captionTargetPageId="7" captionText="FIGURE 5. Leptochelia parasavignyi n. sp., primary male, paratype. A D &amp; F H scale bars 0.1 mm. E, scale bar 0.5 mm. A, labrum. B, maxillule palp. C, maxilliped. D, epignath. E, cheliped. F, same, propodus. G, pleopod. H, uropod." httpUri="https://zenodo.org/record/212700/files/figure.png" pageId="9" pageNumber="33">Fig. 5</figureCitation>
H) basal article with several distal and subdistal setae. Endopod with four articles, all with setae. Exopod uniarticulate, almost as long as first endopod article, with long medial seta and two unequally length distal setae.
</paragraph>
</subSubSection>
<subSubSection id="F6C169E7FF8EFFE1FF434689FDB36CD8" lastPageId="10" lastPageNumber="34" pageId="9" pageNumber="33" type="discussion">
<paragraph id="BE643A6CFF8EFFE1FF434689FDB36CD8" blockId="9.[151,1437,151,2012]" lastBlockId="10.[151,1436,151,967]" lastPageId="10" lastPageNumber="34" pageId="9" pageNumber="33">
<emphasis id="8CAFE67EFF8EFFE2FF434689FECA6A70" bold="true" box="[199,315,1772,1796]" pageId="9" pageNumber="33">Remarks.</emphasis>
The genus
<taxonomicName id="79DB41EFFF8EFFE2FE404689FDB96A70" box="[452,584,1772,1796]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="8CAFE67EFF8EFFE2FE404689FDB96A70" box="[452,584,1772,1796]" italics="true" pageId="9" pageNumber="33">Leptochelia</emphasis>
</taxonomicName>
is one of the genera in Tanaidomorpha with the most cryptic species and present serious obstacles for species identification (
<bibRefCitation id="DA4A479DFF8EFFE2FD7A476AFC556A53" author="Bamber" box="[766,932,1807,1831]" pageId="9" pageNumber="33" refString="Bamber, R. N. (2010) In the footsteps of Henrik Nikolaj Kroyer: the rediscovery and redescription of Leptochelia savignyi (Kroyer, 1842) sensu stricto (Crustacea: Tanaidacea: Leptocheliidae). Proceedings of the Biological Society of Washington, 123, 289 - 311." type="journal article" year="2010">Bamber 2010</bibRefCitation>
). At the same time it is one of the most commonly found genera in the world (
<bibRefCitation id="DA4A479DFF8EFFE2FDD94751FC706A38" author="Larsen" box="[605,897,1844,1869]" pageId="9" pageNumber="33" refString="Larsen, K. &amp; Rayment, H. (2002) New species of Leptochelia (Crustacea: Tanaidacea) from the Andaman Sea, North-eastern Indian Ocean. Phuket Marine Biological Center Special Publication, 23 (1), 17 - 31." type="journal article" year="2002">Larsen &amp; Rayment 2002</bibRefCitation>
) and thus it seems a proper target for genetic studies (
<bibRefCitation id="DA4A479DFF8EFFE2FF724732FE786A1B" author="Larsen" box="[246,393,1879,1903]" pageId="9" pageNumber="33" refString="Larsen, K. (2001) Morphological and molecular investigation of polymorphism and cryptic species in tanaid crustaceans: Implications for tanaid systematics and biodiversity estimates. Zoological Journal of the Linnean Society, 131 (3), 353 - 379." type="journal article" year="2001">Larsen 2001</bibRefCitation>
).
<taxonomicName id="79DB41EFFF8EFFE2FE1B473DFD4E6A04" box="[415,703,1880,1904]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="parasavignyi">
<emphasis id="8CAFE67EFF8EFFE2FE1B473DFD4E6A04" box="[415,703,1880,1904]" italics="true" pageId="9" pageNumber="33">Leptochelia parasavignyi</emphasis>
</taxonomicName>
is clearly closely related to
<taxonomicName id="79DB41EFFF8EFFE2FB8E473CFB766A04" box="[1034,1159,1881,1904]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="8CAFE67EFF8EFFE2FB8E473CFB766A04" box="[1034,1159,1881,1904]" italics="true" pageId="9" pageNumber="33">L. savignyi</emphasis>
</taxonomicName>
but genetic comparison with
<taxonomicName id="79DB41EFFF8EFFE2FF4B4718FEBB6AE0" box="[207,330,1917,1940]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="8CAFE67EFF8EFFE2FF4B4718FEBB6AE0" box="[207,330,1917,1940]" italics="true" pageId="9" pageNumber="33">L. savignyi</emphasis>
</taxonomicName>
from Gran Canarias and
<collectingCountry id="C6CC7AFCFF8EFFE2FDE14718FD406AE0" box="[613,689,1917,1940]" name="France" pageId="9" pageNumber="33">France</collectingCountry>
showed 25% overall difference in the COI gene. This is more than
<taxonomicName id="79DB41EFFF8EFFE2FF1347C4FEBB6ACD" authority="Kroyer" authorityName="Kroyer" box="[151,330,1952,1977]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="8CAFE67EFF8EFFE2FF1347C4FF056ACC" box="[151,244,1952,1976]" italics="true" pageId="9" pageNumber="33">L. dubia</emphasis>
Krøyer
</taxonomicName>
differs from
<taxonomicName id="79DB41EFFF8EFFE2FE5A47C4FC996ACC" authority="Stebbing (Drumm 2010)" authorityName="Stebbing (Drumm" authorityYear="2010" box="[478,872,1952,1976]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="forresti">
<emphasis id="8CAFE67EFF8EFFE2FE5A47C4FDBC6ACC" box="[478,589,1952,1976]" italics="true" pageId="9" pageNumber="33">L. forresti</emphasis>
Stebbing (Drumm 2010)
</taxonomicName>
and two to four times as much as between species of
<taxonomicName id="79DB41EFFF8EFFE2FF3147A0FF0B6AA8" box="[181,250,1989,2012]" class="Malacostraca" family="Tanaididae" genus="Zeuxo" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="8CAFE67EFF8EFFE2FF3147A0FF0B6AA8" box="[181,250,1989,2012]" italics="true" pageId="9" pageNumber="33">Zeuxo</emphasis>
</taxonomicName>
(Larsen
<emphasis id="8CAFE67EFF8EFFE2FED847A0FE656AA8" box="[348,404,1988,2012]" italics="true" pageId="9" pageNumber="33">et al.</emphasis>
research in progress). Morphologically
<taxonomicName id="79DB41EFFF8EFFE2FCD047A0FBF56AA8" box="[852,1028,1989,2012]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="parasavignyi">
<emphasis id="8CAFE67EFF8EFFE2FCD047A0FBF56AA8" box="[852,1028,1989,2012]" italics="true" pageId="9" pageNumber="33">L. parasavignyi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="79DB41EFFF8EFFE2FB1247A0FAE16AA8" box="[1174,1296,1989,2012]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="8CAFE67EFF8EFFE2FB1247A0FAE16AA8" box="[1174,1296,1989,2012]" italics="true" pageId="9" pageNumber="33">L. savignyi</emphasis>
</taxonomicName>
primarily in the longer antennule article 2 and the longer pereonite 6. Additional characters separating the female
<taxonomicName id="79DB41EFFF8DFFE1FA0140FCFED86DA0" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="species" species="parasavignyi">
<emphasis id="8CAFE67EFF8DFFE1FA0140FCFED86DA0" italics="true" pageId="10" pageNumber="34">L. parasavignyi</emphasis>
</taxonomicName>
from
<taxonomicName id="79DB41EFFF8DFFE1FEF440D8FE1D6DA0" box="[368,492,189,212]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis id="8CAFE67EFF8DFFE1FEF440D8FE1D6DA0" box="[368,492,189,212]" italics="true" pageId="10" pageNumber="34">L. savignyi</emphasis>
</taxonomicName>
are: Antennule article 2 longer than article 3. Antenna article 1 with seta; article 2 longer than article 1; article 3 with spiniform seta arising from ventrodistal process. Cheliped basis only 1.5 times as long as wide, dactylus and unguis, combined longer than propodus. Pereopod 2 ischium with two setae. Uropod endopod with five articles (see below); exopod 0.8 times as long as first endopod articles. For the primary male the separating characters are: Antennule with a total of eight articles (flagellum with only four serially repeating articles); article 3 shorter. Cheliped propodus shorter. Pereopod 6 basis without dorsal marginal flange. The uropod endopod has four articles (see below).
</paragraph>
</subSubSection>
<subSubSection id="F6C169E7FF8DFFE1FF4341DDFD756EB3" pageId="10" pageNumber="34" type="reference_group">
<paragraph id="BE643A6CFF8DFFE1FF4341DDFD006F63" blockId="10.[151,1436,151,967]" pageId="10" pageNumber="34">
<taxonomicName id="79DB41EFFF8DFFE1FF4341DDFE196CA4" ID-CoL="6PFHZ" box="[199,488,440,464]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="species" species="parasavignyi">
<emphasis id="8CAFE67EFF8DFFE1FF4341DDFE196CA4" box="[199,488,440,464]" italics="true" pageId="10" pageNumber="34">Leptochelia parasavignyi</emphasis>
</taxonomicName>
can be separated from another Macaronesian
<emphasis id="8CAFE67EFF8DFFE1FB8D41DDFAE66CA4" box="[1033,1303,440,464]" italics="true" pageId="10" pageNumber="34">
<taxonomicName id="79DB41EFFF8DFFE1FB8D41DDFB616CA4" ID-CoL="84PMK" box="[1033,1168,440,464]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="genus">Leptochelia</taxonomicName>
,
<taxonomicName id="79DB41EFFF8DFFE1FB2441DCFAE66CA4" ID-CoL="3TL8W" authority="Bamber" authorityName="Bamber" authorityYear="2009" box="[1184,1303,440,464]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="species" species="caldera">L. caldera</taxonomicName>
</emphasis>
<bibRefCitation id="DA4A479DFF8DFFE1FAA641D2FEEC6C80" author="Bamber" pageId="10" pageNumber="34" refString="Bamber, R. N. &amp; Costa. A. C. (2009) The Tanaidaceans (Arthropoda: Peracarida: Tanaidacea) of Sao Miguel, Azores, with description of two new species, and a new record from Tenerife. Acoreana, Suplemento, 6, 183 - 200." type="journal article" year="2009">Bamber &amp; Costa, 2009</bibRefCitation>
from (Azores) by: Female; antennule article 2 longer than article 3. Maxilliped basis with four to five setae. Primary male; cheliped propodus much shorter.
</paragraph>
<paragraph id="BE643A6CFF8DFFE1FF434241FE9C6FF0" blockId="10.[151,1436,151,967]" pageId="10" pageNumber="34">The new species can be separated from all other species, except those of the minuta group, by having four or five maxilliped basis setae. From the member of the minuta group, the new species differ by the short male cheliped propodus.</paragraph>
<paragraph id="BE643A6CFF8DFFE1FF4342F5FD896E43" blockId="10.[151,1436,151,967]" pageId="10" pageNumber="34">
It is interesting that the number of maxilliped basis setae differs between the left and right basis. The number of these setae is otherwise considered a stable diagnostic character on the species level, but
<bibRefCitation id="DA4A479DFF8DFFE1FB1442D1FA976FB8" author="Gutu" box="[1168,1382,692,716]" pageId="10" pageNumber="34" refString="Gutu, M. &amp; Iliffe, T. M. (2011) Leptochelia vatulelensis (Crustacea; Tanaidacea), a new species from anchialine caves of the south-western Pacific. Travaux du Museum National d´Histoire naturelle &quot; Grigore Antipa &quot;, 54 (2), 351 - 363." type="journal article" year="2011">Guţu &amp; Iliffe 2011</bibRefCitation>
also illustrate a variable number (three to four) of these setae in one specimen of
<taxonomicName id="79DB41EFFF8DFFE1FC6142BDFB986F84" ID-CoL="84PMK" box="[997,1129,728,752]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="8CAFE67EFF8DFFE1FC6142BDFB986F84" box="[997,1129,728,752]" italics="true" pageId="10" pageNumber="34">Leptochelia</emphasis>
</taxonomicName>
(
<taxonomicName id="79DB41EFFF8DFFE1FBFD42BCFAEB6F84" ID-CoL="6PFHL" box="[1145,1306,728,752]" class="Malacostraca" family="Leptocheliidae" genus="Leptochelia" kingdom="Animalia" order="Tanaidacea" pageId="10" pageNumber="34" phylum="Arthropoda" rank="species" species="vatulelensis">
<emphasis id="8CAFE67EFF8DFFE1FBFD42BCFAEB6F84" box="[1145,1306,728,752]" italics="true" pageId="10" pageNumber="34">L. vatulelensis</emphasis>
</taxonomicName>
). This cave dwelling species can be separated from the new species by the lack of a spiniform eye-lobe and by aforementioned difference in maxilliped basis seta number.
</paragraph>
<paragraph id="BE643A6CFF8DFFE1FF434321FD756EB3" blockId="10.[151,1436,151,967]" pageId="10" pageNumber="34">
The use of the uropodal article number as a diagnostic character, sparks some controversy. We are convinced that this character is not entirely stable, neither in this genus nor in many other taxa, while some authors are of the opposite view (
<bibRefCitation id="DA4A479DFF8DFFE1FECC43E9FDCF6ED0" author="Bird" box="[328,574,908,932]" pageId="10" pageNumber="34" refString="Bird, G. J. &amp; Bamber, R. N. (2000) Additions to the Tanaidomorph tanaidaceans (Crustacea: Peracarida) of Hong Kong. In: B. Morton (Ed.) The Marine Flora and Fauna of Hong Kong and Southern China IV. Proceedings of the Tenth International Marine Biological workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong, 2 - 26 April 1998. Hong Kong: Hong Kong University Press, pp. 66 - 104." type="journal article" year="2000">Bird &amp; Bamber 2000</bibRefCitation>
,
<bibRefCitation id="DA4A479DFF8DFFE1FDC843E9FD146ED0" author="Bamber" box="[588,741,908,932]" pageId="10" pageNumber="34" refString="Bamber, R. N. (2010) In the footsteps of Henrik Nikolaj Kroyer: the rediscovery and redescription of Leptochelia savignyi (Kroyer, 1842) sensu stricto (Crustacea: Tanaidacea: Leptocheliidae). Proceedings of the Biological Society of Washington, 123, 289 - 311." type="journal article" year="2010">Bamber 2010</bibRefCitation>
). We therefore recommend caution regarding the use of this character until further data can be collected.
</paragraph>
</subSubSection>
</treatment>
</document>