treatments-xml/data/2E/3D/7C/2E3D7C439A2EA22C60FAC98CFD7FFCD9.xml
2024-06-21 12:32:37 +02:00

595 lines
73 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="4A156ADF5D755CB59AC73785143E77C1" ID-DOI="10.11646/zootaxa.5005.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5141384" ID-ZooBank="96006414-D7CC-488F-B927-696DFA2AE0CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627463447026" checkinUser="plazi" docAuthor="González, Javier Arcos" docDate="2021" docId="2E3D7C439A2EA22C60FAC98CFD7FFCD9" docLanguage="en" docName="zootaxa.5005.2.2.pdf" docOrigin="Zootaxa 5005 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Temnothorax estel González 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="154" masterDocId="D204043B9A2BA225606DCD66FFB6FFE0" masterDocTitle="Description of Temnothorax estel sp. nov. (Hymenoptera: Formicidae), with a review of the Iberian species of the sordidulus species-complex" masterLastPageNumber="160" masterPageNumber="145" pageNumber="150" updateTime="1699025309733" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="7B9C5F7FD8445DE9CF36D6992C0AE844" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AF0BE51B59913AE8A8F3AA4861E027C5">
<mods:title id="53834E3478DC53C039BB1E746667BE46">Description of Temnothorax estel sp. nov. (Hymenoptera: Formicidae), with a review of the Iberian species of the sordidulus species-complex</mods:title>
</mods:titleInfo>
<mods:name id="522CACFCD794CE879E490D3A02ABA042" type="personal">
<mods:role id="42F5DED3075A63DCB6FA820987DD7756">
<mods:roleTerm id="461B85DC086C9F82934EAD8A75FEF6D5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="614848A2A50CC1CE5188A07ED1E7243E">González, Javier Arcos</mods:namePart>
<mods:affiliation id="8E53EE1DEFF87710374045E27868C7CE">Independent researcher, Barcelona. javier. arcos @ e-campus. uab. cat</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A0ECB28715586A46F2ABFF10F986CD8F">text</mods:typeOfResource>
<mods:relatedItem id="C7BA7FDE7EAA509947E236B02BF93BA0" type="host">
<mods:titleInfo id="B4BD8AF1039F6B8CE87E39A7B68186B0">
<mods:title id="67F2345E82EBA7E9D73EA75741B079C8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2E8DF2E9C7736FB08494D4E1E23A9AED">
<mods:date id="A8547AF90F64FDAFC45A914290F6F240">2021</mods:date>
<mods:detail id="8BD94D44E4208188D7FED27033125D8C" type="pubDate">
<mods:number id="71AA54D2330AAB350BE3F5DACA32267A">2021-07-26</mods:number>
</mods:detail>
<mods:detail id="25BC4300C090456DD1D668E8E16C65E0" type="volume">
<mods:number id="DD95E61B20115E61D5C7C063316EDAD1">5005</mods:number>
</mods:detail>
<mods:detail id="390ECDBB54781EBF45307FE6C334406A" type="issue">
<mods:number id="22A72E031EA0D6F964CA14B78E12D98D">2</mods:number>
</mods:detail>
<mods:extent id="7BBE1957C7E3FB2C9B8BD1A8AB0ADDF6" unit="page">
<mods:start id="0036D649E867BE304D080BEEBCAEB063">145</mods:start>
<mods:end id="F4564DB94209BCB738ACD2FE17F649BD">160</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FEE14B724A00B16D96701FE38FE01178">journal article</mods:classification>
<mods:identifier id="08C291B864837B8BD4F5C2A382B56F91" type="DOI">10.11646/zootaxa.5005.2.2</mods:identifier>
<mods:identifier id="1AC68EA2272BD95DB033C69EED49C0EF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F93AD8232FC9188C532880FDD599FB04" type="Zenodo-Dep">5141384</mods:identifier>
<mods:identifier id="AC6E721D3EDCA64A5A22FEA4539437F2" type="ZooBank">96006414-D7CC-488F-B927-696DFA2AE0CD</mods:identifier>
</mods:mods>
<treatment id="2E3D7C439A2EA22C60FAC98CFD7FFCD9" ID-DOI="http://doi.org/10.5281/zenodo.5150493" ID-Zenodo-Dep="5150493" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9" httpUri="http://treatment.plazi.org/id/2E3D7C439A2EA22C60FAC98CFD7FFCD9" lastPageId="9" lastPageNumber="154" pageId="5" pageNumber="150">
<subSubSection id="EE8E9EDE9A2EA22060FAC98CFE6DFAE5" box="[151,475,1258,1285]" pageId="5" pageNumber="150" type="nomenclature">
<paragraph id="A62BCD559A2EA22060FAC98CFE6DFAE5" blockId="5.[151,596,1258,1321]" box="[151,475,1258,1285]" pageId="5" pageNumber="150">
<heading id="FD637A399A2EA22060FAC98CFE6DFAE5" bold="true" box="[151,475,1258,1285]" fontSize="11" level="1" pageId="5" pageNumber="150" reason="1">
<emphasis id="94E011479A2EA22060FAC98CFE6DFAE5" bold="true" box="[151,475,1258,1285]" pageId="5" pageNumber="150">
<taxonomicName id="6194B6D69A2EA22060FAC98CFEC0FAE5" authority="González, 2021" authorityName="González" authorityYear="2021" box="[151,374,1258,1285]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A2EA22060FAC98CFEC0FAE5" bold="true" box="[151,374,1258,1285]" italics="true" pageId="5" pageNumber="150">Temnothorax estel</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8FD3AC3C9A2EA2206113C98DFE6DFAE5" box="[382,475,1259,1285]" pageId="5" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A2EA22060FAC869FDE2FAC9" box="[151,596,1294,1321]" pageId="5" pageNumber="150" type="description">
<paragraph id="A62BCD559A2EA22060FAC869FDE2FAC9" blockId="5.[151,596,1258,1321]" box="[151,596,1294,1321]" pageId="5" pageNumber="150">
(
<figureCitation id="3EAFD1D09A2EA22060CDC869FF42FAC9" box="[160,244,1295,1321]" captionStart="FIGURE 2" captionStartId="7.[151,250,1956,1980]" captionTargetBox="[184,1409,186,1928]" captionTargetId="figure-16@7.[178,1409,181,1932]" captionTargetPageId="7" captionText="FIGURE 2. Head (left) and lateral view (right) of the workers of (A) T. conatensis (Mont-Rebei, Lleida), (B) T. estel sp. nov. (Sierra de Cazorla, Jaén), (C) T. ibericus (Sierra de Javalambre, Teruel), and (D) T. platycephalus (Sierra de Cazorla, Jaén)." figureDoi="http://doi.org/10.5281/zenodo.5141388" httpUri="https://zenodo.org/record/5141388/files/figure.png" pageId="5" pageNumber="150">Fig. 2B</figureCitation>
and
<figureCitation id="3EAFD1D09A2EA2206146C868FEF8FAC8" box="[299,334,1294,1320]" captionStart="FIGURE 6" captionStartId="11.[151,250,1929,1953]" captionTargetBox="[158,1421,419,1897]" captionTargetId="figure-364@11.[151,1436,403,1906]" captionTargetPageId="11" captionText="FIGURE 6. Typical coloration and morphological appearance of the species in the Iberian sordidulus species-complex. Sculpture of frons (left column) and lateral view of workers (right column); T. conatensis (A), T. estel sp. nov. (B), T. ibericus (C) and T. platycephalus (D)." figureDoi="http://doi.org/10.5281/zenodo.5141396" httpUri="https://zenodo.org/record/5141396/files/figure.png" pageId="5" pageNumber="150">6B</figureCitation>
, worker;
<figureCitation id="3EAFD1D09A2EA22061D5C869FE4CFAC9" box="[440,506,1295,1321]" captionStart="FIGURE 3" captionStartId="8.[151,250,808,832]" captionTargetBox="[151,1436,181,784]" captionTargetId="figure-30@8.[151,1436,181,784]" captionTargetPageId="8" captionText="FIGURE 3. Head (left) and lateral view (right) of the queen of T. estel sp. nov. Individual from Sierra de Cazorla, Jaén (see “additional studied material” under the T. estel sp. nov. description above)." figureDoi="http://doi.org/10.5281/zenodo.5141390" httpUri="https://zenodo.org/record/5141390/files/figure.png" pageId="5" pageNumber="150">Fig. 3</figureCitation>
, queen)
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A2EA22060FAC831FB89F971" pageId="5" pageNumber="150" type="materials_examined">
<paragraph id="A62BCD559A2EA22060FAC831FB0FFA3D" blockId="5.[151,1437,1366,2041]" pageId="5" pageNumber="150">
<emphasis id="94E011479A2EA22060FAC831FEF0FA90" bold="true" box="[151,326,1366,1393]" pageId="5" pageNumber="150">Type material.</emphasis>
<materialsCitation id="16FCC7089A2EA2206120C831FDAEFA75" collectingDate="2020-06-20" country="Spain" elevation="1282" latitude="38.651558" location="Serra d'Aitana" longLatPrecision="1" longitude="-0.2371389" pageId="5" pageNumber="150" specimenCount="1" typeStatus="holotype">
<typeStatus id="792F73F79A2EA2206120C831FE03FA91" box="[333,437,1367,1393]" pageId="5" pageNumber="150" type="holotype">Holotype</typeStatus>
worker:
<collectingCountry id="DE838DC59A2EA2206276C831FDE8FA91" box="[539,606,1367,1393]" name="Spain" pageId="5" pageNumber="150">Spain</collectingCountry>
,
<location id="A34B9B8E9A2EA2206207C831FCB8FA91" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9:A34B9B8E9A2EA2206207C831FCB8FA91" box="[618,782,1367,1393]" country="Spain" latitude="38.651558" longLatPrecision="1" longitude="-0.2371389" name="Serra d'Aitana" pageId="5" pageNumber="150">Serra dAitana</location>
(Alacant),
<date id="D22AEB959A2EA22063E3C831FBA7FA91" box="[910,1041,1367,1393]" pageId="5" pageNumber="150" value="2020-06-20">
<collectingDate id="C26E127D9A2EA22063E3C831FBA7FA91" box="[910,1041,1367,1393]" pageId="5" pageNumber="150" value="2020-06-20">20.VI.2020</collectingDate>
</date>
,
<geoCoordinate id="C3A0AB929A2EA2206471C830FB0CFA91" box="[1052,1210,1366,1393]" degrees="38" direction="north" minutes="39" orientation="latitude" pageId="5" pageNumber="150" precision="1" seconds="05.6" value="38.651558">38°3905.6”N</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22064AFC831FAEEFA91" box="[1218,1368,1367,1393]" degrees="0" direction="west" minutes="14" orientation="longitude" pageId="5" pageNumber="150" precision="1" seconds="13.7" value="-0.2371389">0°1413.7”W</geoCoordinate>
,
<quantity id="616C60B09A2EA2206509C831FF1BFA75" metricMagnitude="3" metricUnit="m" metricValue="1.2819999999999998" pageId="5" pageNumber="150" unit="m" value="1282.0">
<elevation id="2DB92A669A2EA2206509C831FF1BFA75" metricMagnitude="3" metricUnit="m" metricValue="1.2819999999999998" pageId="5" pageNumber="150" unit="m" value="1282.0">1282 m</elevation>
</quantity>
(MCNB code MZB 2021-0727)
</materialsCitation>
.
<typeStatus id="792F73F79A2EA220624CC81DFD24FA75" box="[545,658,1403,1429]" pageId="5" pageNumber="150" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="B09206DC9A2EA22062F1C81DFCBBFA75" box="[668,781,1403,1429]" pageId="5" pageNumber="150" type="worker">4 workers</specimenCount>
, same nest and date as
<typeStatus id="792F73F79A2EA2206464C81DFBDDFA75" box="[1033,1131,1403,1429]" pageId="5" pageNumber="150" type="holotype">holotype</typeStatus>
(MCNB codes MZB 2021- 0728, MZB 2021-0729, MZB 2021-0730 and MZB 2021-0731);
<specimenCount id="B09206DC9A2EA2206301C8F8FC68FA59" box="[876,990,1438,1465]" pageId="5" pageNumber="150" type="worker">5 workers</specimenCount>
, same nest and date as
<typeStatus id="792F73F79A2EA22064B3C8F9FAF6FA59" box="[1246,1344,1439,1465]" pageId="5" pageNumber="150" type="holotype">holotype</typeStatus>
(will be deposited in MNHN);
<specimenCount id="B09206DC9A2EA22061F9C8A4FDB0FA3D" box="[404,518,1474,1501]" pageId="5" pageNumber="150" type="worker">5 workers</specimenCount>
, same nest and date as
<typeStatus id="792F73F79A2EA2206364C8A5FCDDFA3D" box="[777,875,1475,1501]" pageId="5" pageNumber="150" type="holotype">holotype</typeStatus>
(will be deposited in CASC).
</paragraph>
<paragraph id="A62BCD559A2EA22060AAC880FB89F971" blockId="5.[151,1437,1366,2041]" pageId="5" pageNumber="150">
<emphasis id="94E011479A2EA22060AAC880FDA3F9E0" bold="true" box="[199,533,1510,1536]" pageId="5" pageNumber="150">Additional studied material</emphasis>
.
<materialsCitation id="16FCC7089A2EA2206248C881FF46F9C5" collectingDate="2012-04-09" elevation="1247" latitude="38.650112" location="Serra d'Aitana" longLatPrecision="1" longitude="-0.2246389" pageId="5" pageNumber="150" specimenCount="2" specimenCount-worker="2">
<specimenCount id="B09206DC9A2EA2206248C881FD2FF9E1" box="[549,665,1511,1537]" pageId="5" pageNumber="150" type="worker">2 workers</specimenCount>
,
<location id="A34B9B8E9A2EA22062CBC881FCFAF9E1" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9:A34B9B8E9A2EA22062CBC881FCFAF9E1" box="[678,844,1511,1537]" latitude="38.650112" longLatPrecision="1" longitude="-0.2246389" name="Serra d'Aitana" pageId="5" pageNumber="150">Serra dAitana</location>
(Alacant),
<date id="D22AEB959A2EA22063A3C881FBFBF9E1" box="[974,1101,1511,1537]" pageId="5" pageNumber="150" value="2012-04-09">
<collectingDate id="C26E127D9A2EA22063A3C881FBFBF9E1" box="[974,1101,1511,1537]" pageId="5" pageNumber="150" value="2012-04-09">09.IV.2012</collectingDate>
</date>
,
<geoCoordinate id="C3A0AB929A2EA2206437C881FB4EF9E1" box="[1114,1272,1511,1537]" degrees="38" direction="north" minutes="39" orientation="latitude" pageId="5" pageNumber="150" precision="1" seconds="00.4" value="38.650112">38°3900.4”N</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220656CC881FA21F9E1" box="[1281,1431,1511,1537]" degrees="0" direction="west" minutes="13" orientation="longitude" pageId="5" pageNumber="150" precision="1" seconds="28.7" value="-0.2246389">0°1328.7”W</geoCoordinate>
,
<quantity id="616C60B09A2EA22060FACB6DFF46F9C5" box="[151,240,1547,1573]" metricMagnitude="3" metricUnit="m" metricValue="1.247" pageId="5" pageNumber="150" unit="m" value="1247.0">
<elevation id="2DB92A669A2EA22060FACB6DFF46F9C5" box="[151,240,1547,1573]" metricMagnitude="3" metricUnit="m" metricValue="1.247" pageId="5" pageNumber="150" unit="m" value="1247.0">1247 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="16FCC7089A2EA2206090CB6DFF44F9A9" collectingDate="2016-06-30" elevation="1425" latitude="38.377415" location="Siles" longLatPrecision="1" longitude="-2.5176945" municipality="Sierra de Cazorla" pageId="5" pageNumber="150" specimenCount="28" specimenCount-male="13" specimenCount-queen="1" specimenCount-worker="14">
<specimenCount id="B09206DC9A2EA2206090CB6DFEC8F9C5" box="[253,382,1547,1573]" pageId="5" pageNumber="150" type="worker">14 workers</specimenCount>
,
<specimenCount id="B09206DC9A2EA22061E7CB6DFE50F9C5" box="[394,486,1547,1573]" pageId="5" pageNumber="150" type="queen">1 queen</specimenCount>
,
<specimenCount id="B09206DC9A2EA220619FCB6DFDECF9C5" box="[498,602,1547,1573]" pageId="5" pageNumber="150" type="male">13 males</specimenCount>
,
<location id="A34B9B8E9A2EA220620ACB6DFD16F9C5" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9:A34B9B8E9A2EA220620ACB6DFD16F9C5" box="[615,672,1547,1573]" latitude="38.377415" longLatPrecision="1" longitude="-2.5176945" municipality="Sierra de Cazorla" name="Siles" pageId="5" pageNumber="150">Siles</location>
,
<collectingMunicipality id="464F572F9A2EA22062C1CB6DFCC7F9C5" box="[684,881,1547,1573]" pageId="5" pageNumber="150">Sierra de Cazorla</collectingMunicipality>
(Jaén),
<date id="D22AEB959A2EA22063A0CB6CFBF9F9C4" box="[973,1103,1546,1572]" pageId="5" pageNumber="150" value="2016-06-30">
<collectingDate id="C26E127D9A2EA22063A0CB6CFBF9F9C4" box="[973,1103,1546,1572]" pageId="5" pageNumber="150" value="2016-06-30">30.VI.2016</collectingDate>
</date>
,
<geoCoordinate id="C3A0AB929A2EA2206436CB6DFB4FF9C5" box="[1115,1273,1547,1573]" degrees="38" direction="north" minutes="22" orientation="latitude" pageId="5" pageNumber="150" precision="1" seconds="38.7" value="38.377415">38°2238.7”N</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220656CCB6DFA2EF9C5" box="[1281,1432,1547,1573]" degrees="2" direction="west" minutes="31" orientation="longitude" pageId="5" pageNumber="150" precision="1" seconds="03.7" value="-2.5176945">2°3103.7”W</geoCoordinate>
,
<quantity id="616C60B09A2EA22060FACB48FF44F9A9" box="[151,242,1582,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="5" pageNumber="150" unit="m" value="1425.0">
<elevation id="2DB92A669A2EA22060FACB48FF44F9A9" box="[151,242,1582,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.425" pageId="5" pageNumber="150" unit="m" value="1425.0">1425 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="16FCC7089A2EA220616CCB49FAC7F9A9" box="[257,1393,1582,1610]" collectingDate="2020-06-20" elevation="1282" latitude="38.651558" location="Serra d'Aitana" longLatPrecision="1" longitude="-0.2371389" pageId="5" pageNumber="150" specimenCount="20" specimenCount-male="10" specimenCount-worker="10">
<specimenCount id="B09206DC9A2EA220616CCB49FE32F9A9" box="[257,388,1583,1609]" pageId="5" pageNumber="150" type="worker">10 workers</specimenCount>
,
<specimenCount id="B09206DC9A2EA22061FECB49FE4BF9A9" box="[403,509,1583,1609]" pageId="5" pageNumber="150" type="male">10 males</specimenCount>
,
<location id="A34B9B8E9A2EA2206261CB49FD05F9A9" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9:A34B9B8E9A2EA2206261CB49FD05F9A9" box="[524,691,1583,1609]" latitude="38.651558" longLatPrecision="1" longitude="-0.2371389" name="Serra d'Aitana" pageId="5" pageNumber="150">Serra dAitana</location>
(Alacant),
<date id="D22AEB959A2EA2206355CB49FC0DF9A9" box="[824,955,1583,1609]" pageId="5" pageNumber="150" value="2020-06-20">
<collectingDate id="C26E127D9A2EA2206355CB49FC0DF9A9" box="[824,955,1583,1609]" pageId="5" pageNumber="150" value="2020-06-20">20.VI.2020</collectingDate>
</date>
,
<geoCoordinate id="C3A0AB929A2EA22063A4CB48FBD1F9A9" box="[969,1127,1582,1609]" degrees="38" direction="north" minutes="39" orientation="latitude" pageId="5" pageNumber="150" precision="1" seconds="05.6" value="38.651558">38°3905.6”N</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220641CCB49FAB1F9A9" box="[1137,1287,1583,1609]" degrees="0" direction="west" minutes="14" orientation="longitude" pageId="5" pageNumber="150" precision="1" seconds="13.7" value="-0.2371389">0°1413.7”W</geoCoordinate>
,
<quantity id="616C60B09A2EA220657BCB49FAC7F9A9" box="[1302,1393,1583,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.2819999999999998" pageId="5" pageNumber="150" unit="m" value="1282.0">
<elevation id="2DB92A669A2EA220657BCB49FAC7F9A9" box="[1302,1393,1583,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.2819999999999998" pageId="5" pageNumber="150" unit="m" value="1282.0">1282 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="16FCC7089A2EA22065EDCB49FB61F98D" collectingDate="2020-09-26" elevation="1796" latitude="37.813362" location="Sierra de Cazorla" longLatPrecision="1" longitude="-2.9601667" pageId="5" pageNumber="150" specimenCount="20" specimenCount-worker="20">
<specimenCount id="B09206DC9A2EA22065EDCB49FF47F98D" pageId="5" pageNumber="150" type="worker">20 workers</specimenCount>
from 2 nests,
<location id="A34B9B8E9A2EA22061FDCB35FDE4F98D" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9:A34B9B8E9A2EA22061FDCB35FDE4F98D" box="[400,594,1619,1645]" latitude="37.813362" longLatPrecision="1" longitude="-2.9601667" name="Sierra de Cazorla" pageId="5" pageNumber="150">Sierra de Cazorla</location>
(Jaén),
<date id="D22AEB959A2EA22062C6CB34FC9BF98C" box="[683,813,1618,1644]" pageId="5" pageNumber="150" value="2020-09-26">
<collectingDate id="C26E127D9A2EA22062C6CB34FC9BF98C" box="[683,813,1618,1644]" pageId="5" pageNumber="150" value="2020-09-26">26.IX.2020</collectingDate>
</date>
,
<geoCoordinate id="C3A0AB929A2EA2206355CB35FC60F98D" box="[824,982,1619,1645]" degrees="37" direction="north" minutes="48" orientation="latitude" pageId="5" pageNumber="150" precision="1" seconds="48.1" value="37.813362">37°4848.1”N</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22063B0CB34FBC2F98D" box="[989,1140,1618,1645]" degrees="2" direction="west" minutes="57" orientation="longitude" pageId="5" pageNumber="150" precision="1" seconds="36.6" value="-2.9601667">2°5736.6”W</geoCoordinate>
,
<quantity id="616C60B09A2EA2206412CB34FB61F98D" box="[1151,1239,1618,1645]" metricMagnitude="3" metricUnit="m" metricValue="1.796" pageId="5" pageNumber="150" unit="m" value="1796.0">
<elevation id="2DB92A669A2EA2206412CB34FB61F98D" box="[1151,1239,1618,1645]" metricMagnitude="3" metricUnit="m" metricValue="1.796" pageId="5" pageNumber="150" unit="m" value="1796.0">1796 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="16FCC7089A2EA220648FCB35FB8CF971" collectingDate="2020-09-26" elevation="1649" latitude="37.802887" location="Sierra de Cazorla" longLatPrecision="1" longitude="-2.9706666" pageId="5" pageNumber="150" specimenCount="30" specimenCount-worker="30">
<specimenCount id="B09206DC9A2EA220648FCB35FAE9F98D" box="[1250,1375,1619,1645]" pageId="5" pageNumber="150" type="worker">30 workers</specimenCount>
from 2 nests,
<location id="A34B9B8E9A2EA220609FCB11FE02F971" LSID="urn:lsid:plazi:treatment:2E3D7C439A2EA22C60FAC98CFD7FFCD9:A34B9B8E9A2EA220609FCB11FE02F971" box="[242,436,1655,1681]" latitude="37.802887" longLatPrecision="1" longitude="-2.9706666" name="Sierra de Cazorla" pageId="5" pageNumber="150">Sierra de Cazorla</location>
(Jaén),
<date id="D22AEB959A2EA2206263CB10FD26F970" box="[526,656,1654,1680]" pageId="5" pageNumber="150" value="2020-09-26">
<collectingDate id="C26E127D9A2EA2206263CB10FD26F970" box="[526,656,1654,1680]" pageId="5" pageNumber="150" value="2020-09-26">26.IX.2020</collectingDate>
</date>
,
<geoCoordinate id="C3A0AB929A2EA22062F1CB11FC8FF971" box="[668,825,1655,1681]" degrees="37" direction="north" minutes="48" orientation="latitude" pageId="5" pageNumber="150" precision="1" seconds="10.4" value="37.802887">37°4810.4”N</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220632CCB10FC61F971" box="[833,983,1654,1681]" degrees="2" direction="west" minutes="58" orientation="longitude" pageId="5" pageNumber="150" precision="1" seconds="14.4" value="-2.9706666">2°5814.4”W</geoCoordinate>
,
<quantity id="616C60B09A2EA220638FCB10FB8CF971" box="[994,1082,1654,1681]" metricMagnitude="3" metricUnit="m" metricValue="1.649" pageId="5" pageNumber="150" unit="m" value="1649.0">
<elevation id="2DB92A669A2EA220638FCB10FB8CF971" box="[994,1082,1654,1681]" metricMagnitude="3" metricUnit="m" metricValue="1.649" pageId="5" pageNumber="150" unit="m" value="1649.0">1649 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A2EA22060AACBFCFEEEF8C1" pageId="5" pageNumber="150" type="diagnosis">
<paragraph id="A62BCD559A2EA22060AACBFCFEEEF8C1" blockId="5.[151,1437,1366,2041]" pageId="5" pageNumber="150">
<emphasis id="94E011479A2EA22060AACBFCFE8CF954" bold="true" box="[199,314,1690,1716]" pageId="5" pageNumber="150">Diagnosis</emphasis>
. Characterized within the Iberian
<taxonomicName id="6194B6D69A2EA22062AACBFDFC8AF955" baseAuthorityName="Muller" baseAuthorityYear="1923" box="[711,828,1691,1717]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="sordidulus">
<emphasis id="94E011479A2EA22062AACBFDFC8AF955" box="[711,828,1691,1717]" italics="true" pageId="5" pageNumber="150">sordidulus</emphasis>
</taxonomicName>
species-complex by being bicolored, with yellowish mesosoma and darker head dorsum; sculpture of frons entirely reticulated; head elongate (mean HL/HWb 1.275); metanotal groove inconspicuous to very shallow; and with the longest propodeal spines of the complex (mean SPST/CS 0.313).
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A2EA22360AACA4CFC58F8E5" lastPageId="6" lastPageNumber="151" pageId="5" pageNumber="150" type="description">
<paragraph id="A62BCD559A2EA22360AACA4CFAD7FEFD" blockId="5.[151,1437,1366,2041]" lastBlockId="6.[151,1437,150,2013]" lastPageId="6" lastPageNumber="151" pageId="5" pageNumber="150">
<emphasis id="94E011479A2EA22060AACA4CFE7AF8A5" bold="true" box="[199,460,1834,1861]" pageId="5" pageNumber="150">Description of worker</emphasis>
(n=50). HL: 0.635 ± 0.031 (
<geoCoordinate id="C3A0AB929A2EA2206368CA4CFCF0F8A5" box="[773,838,1834,1861]" degrees="0.570" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.57">0.570</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220632BCA4CFC23F8A5" box="[838,917,1834,1861]" degrees="0.693" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.693">0.693</geoCoordinate>
) mm; HWb: 0.499 ± 0.032 (
<geoCoordinate id="C3A0AB929A2EA22064A1CA4CFABAF8A5" box="[1228,1292,1834,1861]" degrees="0.453" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.453">0.453</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA2206561CA4CFAEAF8A5" box="[1292,1372,1834,1861]" degrees="0.564" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.564">0.564</geoCoordinate>
) mm; PoOC: 0.246 ± 0.015 (
<geoCoordinate id="C3A0AB929A2EA22061FECA28FE62F889" box="[403,468,1870,1897]" degrees="0.222" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.222">0.222</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22061B9CA28FD95F889" box="[468,547,1870,1897]" degrees="0.268" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.268">0.268</geoCoordinate>
) mm; EL: 0.126 ± 0.007 (
<geoCoordinate id="C3A0AB929A2EA2206328CA29FC35F889" box="[837,899,1871,1897]" degrees="0.111" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.111">0.111</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22063E9CA29FC65F889" box="[900,979,1871,1897]" degrees="0.139" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.139">0.139</geoCoordinate>
) mm; FRS: 0.192 ± 0.013 (
<geoCoordinate id="C3A0AB929A2EA2206568CA28FAF0F889" box="[1285,1350,1870,1897]" degrees="0.176" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.176">0.176</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220652BCA28FA23F889" box="[1350,1429,1870,1897]" degrees="0.213" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.213">0.213</geoCoordinate>
) mm; SL: 0.472 ± 0.029 (
<geoCoordinate id="C3A0AB929A2EA22061C7CA14FE5CF86D" box="[426,490,1906,1933]" degrees="0.425" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.425">0.425</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA2206187CA14FD8FF86D" box="[490,569,1906,1933]" degrees="0.527" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.527">0.527</geoCoordinate>
) mm; LDFS: 0.222 ± 0.008 (
<geoCoordinate id="C3A0AB929A2EA2206312CA14FC09F86D" box="[895,959,1906,1933]" degrees="0.208" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.208">0.208</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22063D2CA14FBB9F86D" box="[959,1039,1906,1933]" degrees="0.236" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.236">0.236</geoCoordinate>
) mm; LMH: 0.062 ± 0.009 (
<geoCoordinate id="C3A0AB929A2EA2206520CA15FA3BF86D" box="[1357,1421,1907,1933]" degrees="0.048" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.048">0.048</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22065E0CA15FF6EF851" degrees="0.083" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.083"> 0.083</geoCoordinate>
) mm; ML: 0.714 ± 0.039 (
<geoCoordinate id="C3A0AB929A2EA2206262CAF0FDF9F851" box="[527,591,1942,1969]" degrees="0.656" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.656">0.656</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220623DCAF0FD28F851" box="[592,670,1942,1969]" degrees="0.786" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.786">0.786</geoCoordinate>
) mm; MH: 0.292 ± 0.024 (
<geoCoordinate id="C3A0AB929A2EA22063B4CAF0FBACF851" box="[985,1050,1942,1969]" degrees="0.254" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.254">0.254</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA2206477CAF0FBDFF851" box="[1050,1129,1942,1969]" degrees="0.333" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.333">0.333</geoCoordinate>
) mm; MGr: 0.001 ± 0.002 (00.007) mm; SPST: 0.174 ± 0.014 (
<geoCoordinate id="C3A0AB929A2EA220625BCADCFDC0F835" box="[566,630,1978,2005]" degrees="0.157" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.157">0.157</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA220621BCADCFD73F835" box="[630,709,1978,2005]" degrees="0.194" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.194">0.194</geoCoordinate>
) mm; SPL: 0.112 ± 0.015 (
<geoCoordinate id="C3A0AB929A2EA2206383CADDFB98F835" box="[1006,1070,1979,2005]" degrees="0.083" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.083">0.083</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA2206443CADDFBC8F835" box="[1070,1150,1979,2005]" degrees="0.139" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.139">0.139</geoCoordinate>
) mm; PEL: 0.236 ± 0.025 (
<geoCoordinate id="C3A0AB929A2EA22060F2CAB9FF69F819" box="[159,223,2015,2041]" degrees="0.208" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.208">0.208</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22060B2CAB9FE98F819" box="[223,302,2015,2041]" degrees="0.287" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.287">0.287</geoCoordinate>
) mm; PEH: 0.204 ± 0.015 (
<geoCoordinate id="C3A0AB929A2EA2206232CAB9FD16F819" box="[607,672,2015,2041]" degrees="0.179" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.179">0.179</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA22062CCCAB9FD59F819" box="[673,751,2015,2041]" degrees="0.231" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.231">0.231</geoCoordinate>
) mm; NOL: 0.161 ± 0.014 (
<geoCoordinate id="C3A0AB929A2EA2206448CAB8FBD0F819" box="[1061,1126,2014,2041]" degrees="0.139" direction="north" orientation="latitude" pageId="5" pageNumber="150" precision="55" value="0.139">0.139</geoCoordinate>
<geoCoordinate id="C3A0AB929A2EA2206408CAB8FB02F819" box="[1125,1204,2014,2041]" degrees="0.185" direction="west" orientation="longitude" pageId="5" pageNumber="150" precision="55" value="-0.185">0.185</geoCoordinate>
) mm; SPEPH: 0.038 ± 0.006 (
<geoCoordinate id="C3A0AB929A2DA2236094CDF1FE8FFF52" box="[249,313,151,178]" degrees="0.028" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.028">0.028</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA2236154CDF1FE3FFF52" box="[313,393,151,178]" degrees="0.046" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.046">0.046</geoCoordinate>
) mm; PECW: 0.130 ± 0.008 (
<geoCoordinate id="C3A0AB929A2DA22362BDCDF1FCA7FF51" box="[720,785,151,177]" degrees="0.120" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA223637CCDF1FCE9FF51" box="[785,863,151,177]" degrees="0.139" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.139">0.139</geoCoordinate>
) mm; PPL: 0.141 ± 0.010 (
<geoCoordinate id="C3A0AB929A2DA22364E6CDF0FB7AFF51" box="[1163,1228,150,177]" degrees="0.129" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.129">0.129</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA22364A6CDF0FAACFF51" box="[1227,1306,150,177]" degrees="0.157" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.157">0.157</geoCoordinate>
) mm; PPH: 0.168 ± 0.012 (
<geoCoordinate id="C3A0AB929A2DA2236151CDDCFECBFF35" box="[316,381,186,213]" degrees="0.152" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.152">0.152</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA2236111CDDCFE7DFF35" box="[380,459,186,213]" degrees="0.194" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.194">0.194</geoCoordinate>
) mm; USH: 9.385 ± 1.293 (610.5) mm; SPTI: 0.180 ± 0.019 (
<geoCoordinate id="C3A0AB929A2DA2236411CDDCFB0AFF35" box="[1148,1212,186,213]" degrees="0.153" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.153">0.153</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA22364D0CDDCFABAFF35" box="[1213,1292,186,213]" degrees="0.213" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.213">0.213</geoCoordinate>
) mm; SPWI: 0.191 ± 0.018 (
<geoCoordinate id="C3A0AB929A2DA223612DCDB8FE36FF19" box="[320,384,222,249]" degrees="0.159" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.159">0.159</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA22361EDCDB8FE66FF19" box="[384,464,222,249]" degrees="0.224" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.224">0.224</geoCoordinate>
) mm; SPBA: 0.132 ± 0.019 (
<geoCoordinate id="C3A0AB929A2DA2236379CDB9FCE2FF1A" box="[788,852,223,250]" degrees="0.110" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.11">0.110</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA2236338CDB9FC15FF1A" box="[853,931,223,250]" degrees="0.166" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.166">0.166</geoCoordinate>
) mm; PW: 0.341 ± 0.022 (
<geoCoordinate id="C3A0AB929A2DA22364A6CDB9FABDFF19" box="[1227,1291,223,249]" degrees="0.313" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.313">0.313</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA2236566CDB9FAECFF19" box="[1291,1370,223,249]" degrees="0.383" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.383">0.383</geoCoordinate>
) mm; PEW: 0.139 ± 0.013 (
<geoCoordinate id="C3A0AB929A2DA22361E7CC65FE7CFEFE" box="[394,458,259,286]" degrees="0.116" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.116">0.116</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA22361A7CC65FDAFFEFE" box="[458,537,259,286]" degrees="0.166" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.166">0.166</geoCoordinate>
) mm; PPW: 0.188 ± 0.014 (
<geoCoordinate id="C3A0AB929A2DA223633ECC65FC22FEFE" box="[851,916,259,286]" degrees="0.162" direction="north" orientation="latitude" pageId="6" pageNumber="151" precision="55" value="0.162">0.162</geoCoordinate>
<geoCoordinate id="C3A0AB929A2DA22363F9CC65FC54FEFE" box="[916,994,259,286]" degrees="0.213" direction="west" orientation="longitude" pageId="6" pageNumber="151" precision="55" value="-0.213">0.213</geoCoordinate>
) mm. See
<tableCitation id="EB16F8EE9A2DA2236434CC65FB1AFEFD" box="[1113,1196,259,285]" captionStart="TABLE 2" captionStartId="12.[151,239,491,517]" captionText="TABLE 2. Measurements for Iberian species of the Temnothorax sordidulus complex. Mean ± standard error is shown, followed by the range in brackets. If a species has the largest or smallest mean among the species, the mean is in bold. If two species are tied for largest or smallest, they both appear in bold. Values for T. sordidulus and T. tergestinus are extracted from Csősz et al. (2015) using measurements of individuals from the supplementary material (not the reported nest means)." pageId="6" pageNumber="151">Table 2</tableCitation>
for the indexes.
</paragraph>
<paragraph id="A62BCD559A2DA22360AACC40FB94FEA1" blockId="6.[151,1437,150,2013]" box="[199,1058,294,321]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACC40FF41FEA0" bold="true" box="[199,247,294,320]" pageId="6" pageNumber="151">Size</emphasis>
. Small species in the
<taxonomicName id="6194B6D69A2DA2236181CC41FDD7FEA1" baseAuthorityName="Muller" baseAuthorityYear="1923" box="[492,609,295,321]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="species" species="sordidulus">
<emphasis id="94E011479A2DA2236181CC41FDD7FEA1" box="[492,609,295,321]" italics="true" pageId="6" pageNumber="151">sordidulus</emphasis>
</taxonomicName>
species-complex (mean CS
<quantity id="616C60B09A2DA22363CDCC40FBA0FEA1" box="[928,1046,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="5.67" pageId="6" pageNumber="151" unit="mm" value="0.567">0.567 mm</quantity>
).
</paragraph>
<paragraph id="A62BCD559A2DA22360AACC2CFB5EFE4D" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACC2CFE1DFE85" bold="true" box="[199,427,330,357]" pageId="6" pageNumber="151">Coloration pattern</emphasis>
. Bicolored species. Dorsum of head dark brown, contrasting with yellowish mesosoma. Antennal club not darkened. Ventral side of head, mandibles, antennae and legs yellowish. Femora occasionally darkened. Tergites of gaster with a dark and diffuse band on their distal third; rest of gaster yellow.
</paragraph>
<paragraph id="A62BCD559A2DA22360AACCD1FAD1FD49" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACCD1FEB1FE31" bold="true" box="[199,263,439,465]" pageId="6" pageNumber="151">Head</emphasis>
. Antenna of 12 segments. Head strongly elongated (mean HL/HWb 1.275). In frontal view, sides of head parallel, converging posteriorly. Posterior margin of head straight. Anterior margin of clypeus rounded. Eyes small (mean EL/CS
<quantity id="616C60B09A2DA2236155CC99FE18FDF9" box="[312,430,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="2.23" pageId="6" pageNumber="151" unit="mm" value="0.223">0.223 mm</quantity>
). Postocular distance similar to the rest of species (PoOC/HL 0.387). Frontal carinae very narrow (mean FRS/CS 0.339). Scape long, but not reaching the posterior margin of head (mean SL/CS 0.831). Mandible of five teeth. Frons with reticulated ground sculpture and distinct but not strong superimposed longitudinal striae, with very narrow and smoother median region. Clypeus surface smooth and shiny, with 911 longitudinal carinae; a median and two paramedian carinae are often more pronounced. Concentric rugae visible on genae.
</paragraph>
<paragraph id="A62BCD559A2DA22360AACFD5FC22FC61" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACFD5FEF3FD2D" bold="true" box="[199,325,691,717]" pageId="6" pageNumber="151">Mesosoma</emphasis>
. Indistinct to shallowly impressed metanotal groove (mean MGr/CS 0.220%). Pronotum narrow (mean PW/CS
<quantity id="616C60B09A2DA223612FCFB1FE0FFD11" box="[322,441,727,753]" metricMagnitude="-4" metricUnit="m" metricValue="6.01" pageId="6" pageNumber="151" unit="mm" value="0.601">0.601 mm</quantity>
). Propodeal spines longer than any other species in the complex (mean SPST/CS
<quantity id="616C60B09A2DA2236530CFB1FF71FCF5" metricMagnitude="-4" metricUnit="m" metricValue="3.08" pageId="6" pageNumber="151" unit="mm" value="0.308">0.308 mm</quantity>
), curved and deviating from longitudinal axis of mesosoma by 45°; in smaller individuals, spines may be much thinner and longer. Declivity before propodeal spines usually present in lateral view. Whole surface of mesosoma with reticulated ground sculpture and superimposed longitudinal striae, which are more developed on lateral margins of pronotum. Setae on mesosoma short (mean LMH/CS 0.110).
</paragraph>
<paragraph id="A62BCD559A2DA22360AACEECFBFFFBD5" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACEECFE62FC44" bold="true" box="[199,468,906,933]" pageId="6" pageNumber="151">Petiole and postpetiole</emphasis>
. Petiole high, with concave anterior face and truncated apex. Usually, a transverse and pronounced carina crosses the petiole in lateral view. Petiolar cylinder narrow than any other species in the complex (mean PECW/PEH 0.229). Subpetiolar process well developed (mean SPEPH/CS 0.067), with rounded apex and occasional concave anterior face. Postpetiole subquadrate in dorsal view, getting narrower towards the base. Petiole and postpetiole narrow in dorsal view (mean PEW/CS 0.245; mean PPW/CS 0.332).
</paragraph>
<paragraph id="A62BCD559A2DA22360AAC958FAAFFAB5" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AAC958FE76FBB9" bold="true" box="[199,448,1086,1113]" pageId="6" pageNumber="151">Description of queen</emphasis>
(n=1). HL:
<quantity id="616C60B09A2DA2236225C959FD0BFBB9" box="[584,701,1087,1113]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="6" pageNumber="151" unit="mm" value="0.73">0.730 mm</quantity>
; HWb:
<quantity id="616C60B09A2DA2236379C958FC3CFBB9" box="[788,906,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" pageId="6" pageNumber="151" unit="mm" value="0.675">0.675 mm</quantity>
; PoOC:
<quantity id="616C60B09A2DA2236384C959FBE8FBB9" box="[1001,1118,1087,1113]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="151" unit="mm" value="0.29">0.290 mm</quantity>
; EL:
<quantity id="616C60B09A2DA22364F6C958FAA7FBB9" box="[1179,1297,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="1.85" pageId="6" pageNumber="151" unit="mm" value="0.185">0.185 mm</quantity>
; FRS:
<quantity id="616C60B09A2DA2236530C959FF70FB9D" metricMagnitude="-4" metricUnit="m" metricValue="2.68" pageId="6" pageNumber="151" unit="mm" value="0.268">0.268 mm</quantity>
; SL:
<quantity id="616C60B09A2DA223616CC904FEC0FB9D" box="[257,374,1122,1149]" metricMagnitude="-4" metricUnit="m" metricValue="5.36" pageId="6" pageNumber="151" unit="mm" value="0.536">0.536 mm</quantity>
; LDFS:
<quantity id="616C60B09A2DA22361B9C905FDFFFB9D" box="[468,585,1123,1149]" metricMagnitude="-4" metricUnit="m" metricValue="2.31" pageId="6" pageNumber="151" unit="mm" value="0.231">0.231 mm</quantity>
; LMH:
<quantity id="616C60B09A2DA22362CCC905FCA0FB9D" box="[673,790,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="151" unit="mm" value="66.0">0,066 mm</quantity>
; ML:
<quantity id="616C60B09A2DA2236337C905FC79FB9D" box="[858,975,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="1.282" pageId="6" pageNumber="151" unit="mm" value="1.282">1.282 mm</quantity>
; MH:
<quantity id="616C60B09A2DA223647AC904FB3DFB9D" box="[1047,1163,1122,1149]" metricMagnitude="-4" metricUnit="m" metricValue="6.56" pageId="6" pageNumber="151" unit="mm" value="0.656">0.656 mm</quantity>
; SPST:
<quantity id="616C60B09A2DA2236489C904FAEFFB9D" box="[1252,1369,1122,1149]" metricMagnitude="-4" metricUnit="m" metricValue="2.59" pageId="6" pageNumber="151" unit="mm" value="0.259">0.259 mm</quantity>
; SPL:
<quantity id="616C60B09A2DA22360FAC9E1FEBAFB41" box="[151,268,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="1.39" pageId="6" pageNumber="151" unit="mm" value="0.139">0.139 mm</quantity>
; PEL:
<quantity id="616C60B09A2DA223613AC9E1FE7AFB41" box="[343,460,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="3.24" pageId="6" pageNumber="151" unit="mm" value="0.324">0.324 mm</quantity>
; PEH:
<quantity id="616C60B09A2DA2236277C9E1FD39FB41" box="[538,655,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="2.8699999999999997" pageId="6" pageNumber="151" unit="mm" value="0.287">0.287 mm</quantity>
; NOL:
<quantity id="616C60B09A2DA223628FC9E1FCE0FB41" box="[738,854,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="2.13" pageId="6" pageNumber="151" unit="mm" value="0.213">0.213 mm</quantity>
; SPEPH:
<quantity id="616C60B09A2DA22363A9C9E1FB8FFB41" box="[964,1081,1159,1185]" metricMagnitude="-5" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="151" unit="mm" value="0.046">0.046 mm</quantity>
; PECW:
<quantity id="616C60B09A2DA22364F2C9E1FAA5FB41" box="[1183,1299,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="1.94" pageId="6" pageNumber="151" unit="mm" value="0.194">0.194 mm</quantity>
; PPL:
<quantity id="616C60B09A2DA2236530C9E1FF70FB25" metricMagnitude="-4" metricUnit="m" metricValue="1.94" pageId="6" pageNumber="151" unit="mm" value="0.194">0.194 mm</quantity>
; PPH:
<quantity id="616C60B09A2DA223617AC9CCFE38FB25" box="[279,398,1194,1221]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="151" unit="mm" value="0.25">0.250 mm</quantity>
; SPTI:
<quantity id="616C60B09A2DA2236188C9CDFDEDFB25" box="[485,603,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="2.8699999999999997" pageId="6" pageNumber="151" unit="mm" value="0.287">0.287 mm</quantity>
; SPWI:
<quantity id="616C60B09A2DA22362D1C9CCFC84FB25" box="[700,818,1194,1221]" metricMagnitude="-4" metricUnit="m" metricValue="3.05" pageId="6" pageNumber="151" unit="mm" value="0.305">0.305 mm</quantity>
; SPBA:
<quantity id="616C60B09A2DA22363FBC9CDFBBBFB25" box="[918,1037,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="2.77" pageId="6" pageNumber="151" unit="mm" value="0.277">0.277 mm</quantity>
; PW:
<quantity id="616C60B09A2DA223643EC9CDFB7CFB25" box="[1107,1226,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="8.04" pageId="6" pageNumber="151" unit="mm" value="0.804">0.804 mm</quantity>
; PEW:
<quantity id="616C60B09A2DA223654CC9CDFA2EFB25" box="[1313,1432,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="2.03" pageId="6" pageNumber="151" unit="mm" value="0.203">0.203 mm</quantity>
; PPW:
<quantity id="616C60B09A2DA223608DC9A9FEE0FB09" box="[224,342,1231,1257]" metricMagnitude="-4" metricUnit="m" metricValue="2.77" pageId="6" pageNumber="151" unit="mm" value="0.277">0.277 mm</quantity>
. Indexes: CS: 0.703; HL/HWb: 1.080; PoOC/HL: 0.397; FRS/CS: 0.382; SL/CS: 0.763; EL/CS: 0.263; LDFS/CS: 0.329; LMH/CS: 0.092; ML/CS: 1.825; MH/CS: 0.934; SPST/CS: 0.368; SPL/CS: 0.197; PEL/ CS: 0.461; NOL/CS: 0.303; PEH/CS: 0.408; PPL/CS: 0.276; PPH/CS: 0.355; SPEPH/CS: 0.066; PECW/CS: 0.276; PEW/CS: 0.289; PPW/CS: 0.395; SPWI/CS: 0.434; SPTI/CS: 0.408; SPBA/CS: 0.395; PW/CS: 1.145.
</paragraph>
<paragraph id="A62BCD559A2DA22360AAC839FAC7F9E9" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">Head and mesosoma brown, with yellowish appendages; abdomen brown with a yellowish macule on the base of the first gastral tergite. Head with reticulated ground sculpture and coarse superimposed longitudinal striae. Scutum longitudinally striated, without median shiny region. Scutellum with a median smooth and shiny region, sides longitudinally striated. Propodeal spines well developed. Petiole high and long, with concave anterior face and triangular apex. Subpetiolar process medium-sized. Sides of postpetiole on dorsal view converging posteriorly.</paragraph>
<paragraph id="A62BCD559A2DA22360AACB74FC58F8E5" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACB74FEF1F9CC" bold="true" box="[199,327,1554,1580]" pageId="6" pageNumber="151">Variability</emphasis>
. Some workers (7% of all examined individuals) may exhibit a darker coloration pattern, with black to dark brown head dorsum contrasting with brown mesosoma and dark brown gaster with a basal yellowish macule on the first tergite. The mesosoma declivity preceding propodeal spines may be pronounced (18% of individuals), present but not pronounced (54%) or absent (27%). The absolute length of the propodeal spines may be bigger in the smaller specimens of the colony. The petiolar apex appears rounded instead of truncated in 25% of individuals. A poorly-developed mesopropodeal depression is present in 23% of individuals, while it appears as inconspicuous in the rest. Subpetiolar process may be reduced in smaller specimens (12%).
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A2DA22360AACA69FDB9F8AD" pageId="6" pageNumber="151" type="etymology">
<paragraph id="A62BCD559A2DA22360AACA69FDB9F8AD" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACA69FEFEF8C9" bold="true" box="[199,328,1807,1833]" pageId="6" pageNumber="151">Etymology</emphasis>
.
<collectingRegion id="645003B79A2DA2236138CA69FE1DF8C9" box="[341,427,1807,1833]" country="Spain" name="Catalunya" pageId="6" pageNumber="151">Catalan</collectingRegion>
for star, in reference to the color of the mesosoma in contrast with the dark head and gaster, which resembles a star in the sky.
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A2DA22C60AACA30FC4CFD81" lastPageId="9" lastPageNumber="154" pageId="6" pageNumber="151" type="discussion">
<paragraph id="A62BCD559A2DA22360AACA30FF51F83D" blockId="6.[151,1437,150,2013]" pageId="6" pageNumber="151">
<emphasis id="94E011479A2DA22360AACA30FEF9F890" bold="true" box="[199,335,1878,1904]" pageId="6" pageNumber="151">Comments.</emphasis>
<taxonomicName id="6194B6D69A2DA223613BCA31FE13F891" authority="González, 2021" authorityName="González" authorityYear="2021" box="[342,421,1879,1905]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A2DA223613BCA31FE13F891" box="[342,421,1879,1905]" italics="true" pageId="6" pageNumber="151">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A2DA22361C1CA31FDB0F891" bold="true" box="[428,518,1879,1905]" pageId="6" pageNumber="151">
<taxonomicNameLabel id="8FD3AC3C9A2DA22361C1CA31FDB0F891" box="[428,518,1879,1905]" pageId="6" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily distinguished from any other taxa of the Iberian
<taxonomicName id="6194B6D69A2DA22364E1CA31FAB7F891" baseAuthorityName="Muller" baseAuthorityYear="1923" box="[1164,1281,1879,1905]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="species" species="sordidulus">
<emphasis id="94E011479A2DA22364E1CA31FAB7F891" box="[1164,1281,1879,1905]" italics="true" pageId="6" pageNumber="151">sordidulus</emphasis>
</taxonomicName>
species-complex by the unique combination of bicolored body, long head (mean HL/HWb 1.275), inconspicuous to poorly-developed metanotal groove (mean MGr/CS 0.297%) and longest propodeal spines of the complex (mean SPST/CS 0.313).
</paragraph>
<caption id="F2EB9DDD9A2CA22260FACAC2FACEF800" ID-DOI="http://doi.org/10.5281/zenodo.5141388" ID-Zenodo-Dep="5141388" httpUri="https://zenodo.org/record/5141388/files/figure.png" pageId="7" pageNumber="152" startId="7.[151,250,1956,1980]" targetBox="[184,1409,186,1928]" targetPageId="7">
<paragraph id="A62BCD559A2CA22260FACAC2FACEF800" blockId="7.[151,1436,1956,2016]" pageId="7" pageNumber="152">
<emphasis id="94E011479A2CA22260FACAC2FEA5F85C" bold="true" box="[151,275,1956,1980]" pageId="7" pageNumber="152">FIGURE 2.</emphasis>
Head (left) and lateral view (right) of the workers of (A)
<taxonomicName id="6194B6D69A2CA222630FCAC3FB7FF85C" authority="(Mont-Rebei, Lleida)" baseAuthorityName="Mont-Rebei, Lleida" box="[866,1225,1956,1980]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="152" phylum="Arthropoda" rank="species" species="conatensis">
<emphasis id="94E011479A2CA222630FCAC3FC50F85C" box="[866,998,1957,1980]" italics="true" pageId="7" pageNumber="152">T. conatensis</emphasis>
(Mont-Rebei, Lleida)
</taxonomicName>
, (B)
<taxonomicName id="6194B6D69A2CA2226491CAC3FAF5F85C" authority="González, 2021" authorityName="González" authorityYear="2021" box="[1276,1347,1956,1980]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="152" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A2CA2226491CAC3FAF5F85C" box="[1276,1347,1956,1980]" italics="true" pageId="7" pageNumber="152">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A2CA2226526CAC2FA2AF85C" bold="true" box="[1355,1436,1956,1980]" pageId="7" pageNumber="152">
<taxonomicNameLabel id="8FD3AC3C9A2CA2226526CAC2FA2AF85C" box="[1355,1436,1956,1980]" pageId="7" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Sierra de Cazorla, Jaén), (C)
<taxonomicName id="6194B6D69A2CA22261ABCAAFFCDAF800" authority="(Sierra de Javalambre, Teruel)" baseAuthorityName="Sierra de Javalambre, Teruel" box="[454,876,1992,2016]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="152" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="94E011479A2CA22261ABCAAFFD87F800" box="[454,561,1992,2016]" italics="true" pageId="7" pageNumber="152">T. ibericus</emphasis>
(Sierra de Javalambre, Teruel)
</taxonomicName>
, and (D)
<taxonomicName id="6194B6D69A2CA22263A7CAAFFAC5F800" authority="(Sierra de Cazorla, Jaen)" baseAuthorityName="Sierra de Cazorla, Jaen" box="[970,1395,1992,2016]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="152" phylum="Arthropoda" rank="species" species="platycephalus">
<emphasis id="94E011479A2CA22263A7CAAFFBC6F800" box="[970,1136,1992,2016]" italics="true" pageId="7" pageNumber="152">T. platycephalus</emphasis>
(Sierra de Cazorla, Jaén)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="F2EB9DDD9A23A22D60FACE4EFC39FC84" ID-DOI="http://doi.org/10.5281/zenodo.5141390" ID-Zenodo-Dep="5141390" httpUri="https://zenodo.org/record/5141390/files/figure.png" pageId="8" pageNumber="153" startId="8.[151,250,808,832]" targetBox="[151,1436,181,784]" targetPageId="8">
<paragraph id="A62BCD559A23A22D60FACE4EFC39FC84" blockId="8.[151,1437,808,868]" pageId="8" pageNumber="153">
<emphasis id="94E011479A23A22D60FACE4EFEA2FCA0" bold="true" box="[151,276,808,832]" pageId="8" pageNumber="153">FIGURE 3.</emphasis>
Head (left) and lateral view (right) of the queen of
<taxonomicName id="6194B6D69A23A22D6341CE4FFCC2FCA0" authority="González, 2021" authorityName="González" authorityYear="2021" box="[812,884,808,832]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A23A22D6341CE4FFCC2FCA0" box="[812,884,808,832]" italics="true" pageId="8" pageNumber="153">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A23A22D6311CE4EFC78FCA0" bold="true" box="[892,974,808,832]" pageId="8" pageNumber="153">
<taxonomicNameLabel id="8FD3AC3C9A23A22D6311CE4EFC78FCA0" box="[892,974,808,832]" pageId="8" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Individual from Sierra de Cazorla, Jaén (see “additional studied material” under the
<taxonomicName id="6194B6D69A23A22D624ACE2BFDD8FC84" authority="González, 2021" authorityName="González" authorityYear="2021" box="[551,622,844,868]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A23A22D624ACE2BFDD8FC84" box="[551,622,844,868]" italics="true" pageId="8" pageNumber="153">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A23A22D6218CE2AFD70FC84" bold="true" box="[629,710,844,868]" pageId="8" pageNumber="153">
<taxonomicNameLabel id="8FD3AC3C9A23A22D6218CE2AFD70FC84" box="[629,710,844,868]" pageId="8" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
description above).
</paragraph>
</caption>
<caption id="F2EB9DDD9A23A22D60FACA3AFC2AF85C" ID-DOI="http://doi.org/10.5281/zenodo.5141392" ID-Zenodo-Dep="5141392" httpUri="https://zenodo.org/record/5141392/files/figure.png" pageId="8" pageNumber="153" startId="8.[151,250,1884,1908]" targetBox="[151,1436,892,1860]" targetPageId="8">
<paragraph id="A62BCD559A23A22D60FACA3AFC2AF85C" blockId="8.[151,1437,1884,1980]" pageId="8" pageNumber="153">
<emphasis id="94E011479A23A22D60FACA3AFEA5F894" bold="true" box="[151,275,1884,1908]" pageId="8" pageNumber="153">FIGURE 4.</emphasis>
A: Nest of
<taxonomicName id="6194B6D69A23A22D61E4CA3BFE66F894" authority="González, 2021" authorityName="González" authorityYear="2021" box="[393,464,1884,1908]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A23A22D61E4CA3BFE66F894" box="[393,464,1884,1908]" italics="true" pageId="8" pageNumber="153">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A23A22D61BACA3AFD91F894" bold="true" box="[471,551,1884,1908]" pageId="8" pageNumber="153">
<taxonomicNameLabel id="8FD3AC3C9A23A22D61BACA3AFD91F894" box="[471,551,1884,1908]" pageId="8" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in a rock crevice in Serra dAitana, Alacant. B: Worker of
<taxonomicName id="6194B6D69A23A22D6411CA3BFA94F894" box="[1148,1314,1884,1908]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="platycephalus">
<emphasis id="94E011479A23A22D6411CA3BFA94F894" box="[1148,1314,1884,1908]" italics="true" pageId="8" pageNumber="153">T. platycephalus</emphasis>
</taxonomicName>
from Sierra de Cazorla, Jaén. C: big rocks where both
<taxonomicName id="6194B6D69A23A22D6227CAE7FD27F878" authority="González, 2021" authorityName="González" authorityYear="2021" box="[586,657,1920,1944]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A23A22D6227CAE7FD27F878" box="[586,657,1920,1944]" italics="true" pageId="8" pageNumber="153">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A23A22D62F4CAE6FD5CF878" bold="true" box="[665,746,1920,1944]" pageId="8" pageNumber="153">
<taxonomicNameLabel id="8FD3AC3C9A23A22D62F4CAE6FD5CF878" box="[665,746,1920,1944]" pageId="8" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="6194B6D69A23A22D6370CAE7FC72F878" box="[797,964,1920,1944]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="platycephalus">
<emphasis id="94E011479A23A22D6370CAE7FC72F878" box="[797,964,1920,1944]" italics="true" pageId="8" pageNumber="153">T. platycephalus</emphasis>
</taxonomicName>
establish their nest in Sierra de Cazorla, Jaén. D: general view of the habitat of
<taxonomicName id="6194B6D69A23A22D618ACAC3FD98F85C" authority="González, 2021" authorityName="González" authorityYear="2021" box="[487,558,1956,1980]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A23A22D618ACAC3FD98F85C" box="[487,558,1956,1980]" italics="true" pageId="8" pageNumber="153">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A23A22D6259CAC2FD33F85C" bold="true" box="[564,645,1956,1980]" pageId="8" pageNumber="153">
<taxonomicNameLabel id="8FD3AC3C9A23A22D6259CAC2FD33F85C" box="[564,645,1956,1980]" pageId="8" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Serra dAitana, Alacant.
</paragraph>
</caption>
<paragraph id="A62BCD559A22A22C60AACDF1FA9FFE4D" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="154">
The most closely related species to
<taxonomicName id="6194B6D69A22A22C6234CDF1FD1FFF51" authority="González, 2021" authorityName="González" authorityYear="2021" box="[601,681,151,177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A22A22C6234CDF1FD1FFF51" box="[601,681,151,177]" italics="true" pageId="9" pageNumber="154">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A22A22C62DCCDF1FCBDFF51" bold="true" box="[689,779,151,177]" pageId="9" pageNumber="154">
<taxonomicNameLabel id="8FD3AC3C9A22A22C62DCCDF1FCBDFF51" box="[689,779,151,177]" pageId="9" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is
<taxonomicName id="6194B6D69A22A22C6340CDF1FC75FF50" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[813,963,151,176]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C6340CDF1FC75FF50" box="[813,963,151,176]" italics="true" pageId="9" pageNumber="154">T. tergestinus</emphasis>
</taxonomicName>
, from Central Europe. The first has much longer head (mean HL/HWb 1.252 vs. 1.206), narrower frontal carina (mean FRS/CS 0.343 vs. 0.361), longer scape (mean SL/CS 0.826 vs. 0.799), inconspicuous to poorly-developed metanotal groove (vs. poorly to well-developed metanotal groove in
<taxonomicName id="6194B6D69A22A22C61E5CC65FD97FEFC" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[392,545,259,284]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C61E5CC65FD97FEFC" box="[392,545,259,284]" italics="true" pageId="9" pageNumber="154">T. tergestinus</emphasis>
</taxonomicName>
) and longer and more diverging propodeal spines (mean SPST/CS 0.313 vs. 0.276; SPWI/CS 0.342 vs. 0.333). Both species exhibit the same coloration pattern, but
<taxonomicName id="6194B6D69A22A22C640BCC41FB4CFEA0" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[1126,1274,295,320]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C640BCC41FB4CFEA0" box="[1126,1274,295,320]" italics="true" pageId="9" pageNumber="154">T. tergestinus</emphasis>
</taxonomicName>
can also show extreme dark forms (
<bibRefCitation id="C205B0A49A22A22C61EBCC2DFDE6FE85" author="Csosz, S. &amp; Heinze, J. &amp; Miko, I." box="[390,592,330,357]" pageId="9" pageNumber="154" pagination="0140000" refId="ref10287" refString="Csosz, S., Heinze, J. &amp; Miko, I. (2015) Taxonomic Synopsis of the Ponto-Mediterranean Ants of Temnothorax nylanderi Species-Group. PLoS ONE, 10 (11), e 0140000. https: // doi. org / 10.1371 / journal. pone. 0140000" type="journal article" year="2015">
Csősz
<emphasis id="94E011479A22A22C61BECC2DFDB3FE85" box="[467,517,331,357]" italics="true" pageId="9" pageNumber="154">et al</emphasis>
. 2015
</bibRefCitation>
;
<bibRefCitation id="C205B0A49A22A22C6230CC2DFCCDFE85" author="Borowiec, L. &amp; Salata, S." box="[605,891,331,357]" pageId="9" pageNumber="154" pagination="217 - 248" refId="ref10120" refString="Borowiec, L. &amp; Salata, S. (2018) Ants from Thessaly, Greece (Hymenoptera: Formicidae). Polish Journal of Entomology, 87, 217 - 248. https: // doi. org / 10.2478 / pjen- 2018 - 0016" type="journal article" year="2018">Borowiec &amp; Salata 2018</bibRefCitation>
).
<taxonomicName id="6194B6D69A22A22C63E3CC2DFB12FE84" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[910,1188,331,357]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C63E3CC2DFB12FE84" box="[910,1188,331,357]" italics="true" pageId="9" pageNumber="154">Temnothorax tergestinus</emphasis>
</taxonomicName>
has a wide Palearctic distribution from
<collectingCountry id="DE838DC59A22A22C610DCC09FE76FE69" box="[352,448,367,393]" name="Bulgaria" pageId="9" pageNumber="154">Bulgaria</collectingCountry>
to
<collectingCountry id="DE838DC59A22A22C6185CC09FD81FE69" box="[488,567,367,393]" name="France" pageId="9" pageNumber="154">France</collectingCountry>
, but has not been yet recorded from
<collectingCountry id="DE838DC59A22A22C63B1CC09FB97FE69" box="[988,1057,367,393]" name="Georgia" pageId="9" pageNumber="154">Iberia</collectingCountry>
. It is unknown whether a contact zone between
<taxonomicName id="6194B6D69A22A22C615ACCF5FE30FE4D" authority="González, 2021" authorityName="González" authorityYear="2021" box="[311,390,403,429]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A22A22C615ACCF5FE30FE4D" box="[311,390,403,429]" italics="true" pageId="9" pageNumber="154">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A22A22C61E0CCF5FE51FE4D" bold="true" box="[397,487,403,429]" pageId="9" pageNumber="154">
<taxonomicNameLabel id="8FD3AC3C9A22A22C61E0CCF5FE51FE4D" box="[397,487,403,429]" pageId="9" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="6194B6D69A22A22C6270CCF5FD05FE4C" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[541,691,403,428]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C6270CCF5FD05FE4C" box="[541,691,403,428]" italics="true" pageId="9" pageNumber="154">T. tergestinus</emphasis>
</taxonomicName>
could exist somewhere in North
<collectingCountry id="DE838DC59A22A22C644BCCF5FBD1FE4D" box="[1062,1127,403,429]" name="Georgia" pageId="9" pageNumber="154">Iberia</collectingCountry>
or South
<collectingCountry id="DE838DC59A22A22C64B8CCF5FA93FE4D" box="[1237,1317,403,429]" name="France" pageId="9" pageNumber="154">France</collectingCountry>
.
</paragraph>
<paragraph id="A62BCD559A22A22C60AACCD1FC4CFD81" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="154">
MGr is probably one of the best discriminant characters between
<taxonomicName id="6194B6D69A22A22C63C5CCD1FC41FE31" authority="González, 2021" authorityName="González" authorityYear="2021" box="[936,1015,439,465]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A22A22C63C5CCD1FC41FE31" box="[936,1015,439,465]" italics="true" pageId="9" pageNumber="154">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A22A22C6392CCD1FBEFFE31" bold="true" box="[1023,1113,439,465]" pageId="9" pageNumber="154">
<taxonomicNameLabel id="8FD3AC3C9A22A22C6392CCD1FBEFFE31" box="[1023,1113,439,465]" pageId="9" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="6194B6D69A22A22C64FDCCD1FA90FE30" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[1168,1318,439,464]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C64FDCCD1FA90FE30" box="[1168,1318,439,464]" italics="true" pageId="9" pageNumber="154">T. tergestinus</emphasis>
</taxonomicName>
, but it has not been measured for the latter species in this paper. Queens of both taxa appear very similar, and the only available queen of
<taxonomicName id="6194B6D69A22A22C6092CC99FEFBFDF9" authority="González, 2021" authorityName="González" authorityYear="2021" box="[255,333,511,537]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="estel" status="sp. nov.">
<emphasis id="94E011479A22A22C6092CC99FEFBFDF9" box="[255,333,511,537]" italics="true" pageId="9" pageNumber="154">T. estel</emphasis>
</taxonomicName>
<emphasis id="94E011479A22A22C6139CC99FE1BFDF9" bold="true" box="[340,429,511,537]" pageId="9" pageNumber="154">
<taxonomicNameLabel id="8FD3AC3C9A22A22C6139CC99FE1BFDF9" box="[340,429,511,537]" pageId="9" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does not allow a safe separation with the queens of
<taxonomicName id="6194B6D69A22A22C639FCC99FB31FDF8" baseAuthorityName="Finzi" baseAuthorityYear="1928" box="[1010,1159,511,536]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="94E011479A22A22C639FCC99FB31FDF8" box="[1010,1159,511,536]" italics="true" pageId="9" pageNumber="154">T. tergestinus</emphasis>
</taxonomicName>
. Males of the
<taxonomicName id="6194B6D69A22A22C654ACC99FA2AFDF9" baseAuthorityName="Muller" baseAuthorityYear="1923" box="[1319,1436,511,537]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="154" phylum="Arthropoda" rank="species" species="sordidulus">
<emphasis id="94E011479A22A22C654ACC99FA2AFDF9" box="[1319,1436,511,537]" italics="true" pageId="9" pageNumber="154">sordidulus</emphasis>
</taxonomicName>
species-complex are largely unknown and rarely described, since the external morphology is very variable within the same nest and the study of the genitalia is not routinely used in the genus.
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A22A22C60AACF0CFD97FCF5" pageId="9" pageNumber="154" type="biology_ecology">
<paragraph id="A62BCD559A22A22C60AACF0CFD97FCF5" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="154">
<emphasis id="94E011479A22A22C60AACF0CFE97FD64" bold="true" box="[199,289,618,644]" pageId="9" pageNumber="154">Biology</emphasis>
. Mostly unknown. Nests were found at high altitude in rock crevices and once under a plain stone, and populations were located from stony open habitats to semi-open pine forests (
<figureCitation id="3EAFD1D09A22A22C642CCFE9FB3FFD49" box="[1089,1161,655,681]" captionStart="FIGURE 4" captionStartId="8.[151,250,1884,1908]" captionTargetBox="[151,1436,892,1860]" captionTargetId="figure-67@8.[151,1436,892,1860]" captionTargetPageId="8" captionText="FIGURE 4. A: Nest of T. estel sp. nov. in a rock crevice in Serra dAitana, Alacant. B: Worker of T. platycephalus from Sierra de Cazorla, Jaén. C: big rocks where both T. estel sp. nov. and T. platycephalus establish their nest in Sierra de Cazorla, Jaén. D: general view of the habitat of T. estel sp. nov. in Serra dAitana, Alacant." figureDoi="http://doi.org/10.5281/zenodo.5141392" httpUri="https://zenodo.org/record/5141392/files/figure.png" pageId="9" pageNumber="154">Fig. 4</figureCitation>
). All colonies were monogynous. Nest densities were always low. Two colonies kept in an artificial nest were fed with insects and nectar and produced reproductive individuals in early July. The elongated head may be an adaptation to move into rock crevices and nest in narrow spaces.
</paragraph>
</subSubSection>
<subSubSection id="EE8E9EDE9A22A22C60AACE78FD7FFCD9" box="[199,713,798,825]" pageId="9" pageNumber="154" type="distribution">
<paragraph id="A62BCD559A22A22C60AACE78FD7FFCD9" blockId="9.[151,1437,151,825]" box="[199,713,798,825]" pageId="9" pageNumber="154">
<emphasis id="94E011479A22A22C60AACE78FEEFFCD8" bold="true" box="[199,345,798,824]" pageId="9" pageNumber="154">Distribution</emphasis>
. Known from SE
<collectingCountry id="DE838DC59A22A22C624BCE79FDD1FCD9" box="[550,615,799,825]" name="Georgia" pageId="9" pageNumber="154">Iberia</collectingCountry>
(
<figureCitation id="3EAFD1D09A22A22C621ACE79FD0DFCD9" box="[631,699,798,825]" captionStart="FIGURE 5" captionStartId="9.[151,250,1887,1911]" captionTargetBox="[151,1437,849,1864]" captionTargetId="figure-343@9.[151,1437,849,1864]" captionTargetPageId="9" captionText="FIGURE 5. Distribution of the southern species in the sordidulus species-complex, including the French records of T. tergestinus and T. sordidulus. Colors: T. conatensis (dark blue circles), T. estel sp. nov. (gold circles), T. ibericus (purple circles), T. platycephalus (red circles), T. sordidulus (green circle) and T. tergestinus (yellow circle)." figureDoi="http://doi.org/10.5281/zenodo.5141394" httpUri="https://zenodo.org/record/5141394/files/figure.png" pageId="9" pageNumber="154">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>