253 lines
26 KiB
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253 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.295.4905" ID-GBIF-Dataset="59cbf2a9-41ca-4f5a-a350-ef034991798b" ID-PMC="PMC3677376" ID-Pensoft-Pub="1313-2970-295-1" ID-PubMed="23794874" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-295-1" ModsDocOrigin="ZooKeys 295" ModsDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" checkinTime="1451247354359" checkinUser="pensoft" docAuthor="Branstetter, Michael G." docDate="2013" docId="2E0CEC734A0FC5640B5616BDBA24FC2A" docLanguage="en" docName="ZooKeys 295: 1-277" docOrigin="ZooKeys 295" docSource="http://dx.doi.org/10.3897/zookeys.295.4905" docTitle="Stenamma nonotch Branstetter, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="168" masterDocId="FFB3D3328201C177FF99FFC5FFA8A72C" masterDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="164" updateTime="1668155749461" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branstetter, Michael G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>295</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>277</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.295.4905</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.295.4905</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-295-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152043646" LSID="urn:lsid:zoobank.org:act:D8476855-E0CF-4042-AC45-E4AF73F9B95B" httpUri="http://treatment.plazi.org/id/2E0CEC734A0FC5640B5616BDBA24FC2A" lastPageId="167" lastPageNumber="168" pageId="163" pageNumber="164">
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<subSubSection pageId="163" pageNumber="164" type="nomenclature">
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<paragraph pageId="163" pageNumber="164">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:D8476855-E0CF-4042-AC45-E4AF73F9B95B" class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="163" pageNumber="164" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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<taxonomicNameLabel pageId="163" pageNumber="164">sp. n.</taxonomicNameLabel>
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Worker: Figures 131, 132; Queen: Figure 133; Map: Figure 130
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</paragraph>
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</subSubSection>
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<subSubSection pageId="163" pageNumber="164" type="reference_group">
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<paragraph pageId="163" pageNumber="164">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma" order="Hymenoptera" pageId="163" pageNumber="164" phylum="Arthropoda" rank="genus">Stenamma</taxonomicName>
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mgb11 [variant 2 below] Branstetter, 2012: phylogeny.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="163" pageNumber="164" type="type material">
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<paragraph pageId="163" pageNumber="164">Type material.</paragraph>
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<paragraph pageId="163" pageNumber="164">
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Holotype worker.
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<normalizedToken originalValue="MÉXICO">MEXICO</normalizedToken>
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, Oaxaca: 4.3km SW Valle Nacional,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.7432">17.74320°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.32729">96.32729°W</geoCoordinate>
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<normalizedToken originalValue="±">+/-</normalizedToken>
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40m, 370m, 13 Aug 2009, disturbed tropical rainforest, ex sifted leaf litter (L.
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<normalizedToken originalValue="Sáenz">Saenz</normalizedToken>
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, collection LSD329) [USNM, specimen CASENT0605789]. Paratypes: same data as holotype [1dq, 1w, CAS, CASENT0605782, CASENT0605788], [1w, EAPZ, CASENT0605790], [1w, ECOSCE, CASENT0623408], [1w, FMNH, CASENT0623409], [1w, ICN, CASENT0623410], [1w, INBio, CASENT0623411], [1w, JTLC, CASENT0623412], [1w, LACM, CASENT0623413], [1w, MGBPC, CASENT0623414], [1dq, 1w, MCZ, CASENT0605784, CASENT0623415], [1w, MZSP, CASENT0623416], [1w, UCD, CASENT0623417], [1dq, 1w, UNAM, CASENT0605783, CASENT0623418], [1dq, 1w, USNM, CASENT0623407, CASENT0623419], [1w, UVGC, CASENT0623420].
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="164" lastPageNumber="165" pageId="163" pageNumber="164" type="worker diagnosis">
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<paragraph pageId="163" pageNumber="164">Worker diagnosis.</paragraph>
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<paragraph lastPageId="164" lastPageNumber="165" pageId="163" pageNumber="164">
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Integument mostly dark red-brown (almost black) to brown; small- to medium-sized species (see HL, ML, PrW below); anterior clypeal
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<pageBreakToken pageId="164" pageNumber="165" start="start">margin</pageBreakToken>
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entire, or at most with a nearly imperceptible median notch (only visible from anterodorsal angle); basal margin of mandible straight; face densely sculptured, mostly rugoreticulate; dorsum of promesonotum rugoreticulate, or with many irregular rugulae; pilosity on gastral tergites indistinctly bilayered, forming a somewhat sparse layer of long suberect setae, and a very sparse layer of short decumbent setae; eye of small to moderate size (EL 0.09-0.11, REL 15-19), oval-shaped, with 4-7 ommatidia at greatest diameter; propodeal spines reduced to small triangular tubercles (PSL 0.08-0.14, PSI 1.1-1.4); frontal lobes of moderate width (FLD 0.14-0.19, FLI 25-29). Similar species:
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma ignotum" order="Hymenoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="ignotum">Stenamma ignotum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nanozoi" order="Hymenoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="nanozoi">Stenamma nanozoi</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma picopicucha" order="Hymenoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="picopicucha">Stenamma picopicucha</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma sandinista" order="Hymenoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="sandinista">Stenamma sandinista</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma schmidti" order="Hymenoptera" pageId="164" pageNumber="165" phylum="Arthropoda" rank="species" species="schmidti">Stenamma schmidti</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="164" pageNumber="165" type="geographic range">
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<paragraph pageId="164" pageNumber="165">Geographic range.</paragraph>
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<paragraph pageId="164" pageNumber="165">Southern Mexico to Guatemala.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="165" lastPageNumber="166" pageId="164" pageNumber="165" type="worker description">
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<paragraph pageId="164" pageNumber="165">Worker description.</paragraph>
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<paragraph pageId="164" pageNumber="165">(17 measured) HL 0.57-0.76 (0.61), HW 0.51-0.68 (0.57), FLD 0.14-0.19 (0.15), PCW 0.03-0.07 (0.03), SL 0.46-0.64 (0.50), EL 0.09-0.11 (0.09), ACL 0.45-0.59 (0.49), ML 0.68-1.00 (0.78), PrW 0.36-0.49 (0.40), PSL 0.08-0.14 (0.11), SDL 0.06-0.11 (0.09), PL 0.26-0.40 (0.31), PH 0.16-0.23 (0.19), PW 0.14-0.19 (0.16), PPL 0.12-0.22 (0.15), PPH 0.13-0.21 (0.16), PPW 0.16-0.22 (0.18), MFL 0.50-0.72 (0.55), MTL 0.41-0.57 (0.45), CI 86-92 (92), SI 88-97 (89), REL 15-19 (16), FLI 25-29 (26), PSI 1.1-1.4 (1.3), MFI 93-104 (103), ACI1 65-69 (67), ACI2 92-99 (97).</paragraph>
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<paragraph lastPageId="165" lastPageNumber="166" pageId="164" pageNumber="165">
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Small- to medium-sized species; general body color dark red-brown (almost black) to brown, with appendages lighter, brown to yellow-brown toward extremities; setae golden; mandible with 6-7 teeth (usually 7), consisting of 3 larger, more distinct apical teeth, a basal tooth, and 2-3 smaller denticulate teeth, which are usually more worn, and sometimes indistinct; basal margin of mandible straight, without a basal notch or depression; mandible mostly smooth, except for scattered piligerous punctae, and a few basal striae; anterior clypeal margin entire, or at most with a nearly imperceptible median notch, only visible from anterodorsal view; median lobe of clypeus with faint longitudinal carinulae and some punctations, apex of lobe with a few short transverse carinulae, remainder of clypeus mostly smooth; posterior extension of clypeus between antennal insertions of moderate width (PCW 0.03-0.07), with sides subparallel to slightly hour-glass-shaped; frontal lobes of moderate width (FLD 0.14-0.19, FLI 25-29), not obscuring torular lobes in full-face view; head roughly oval-shaped (CI 86-92), posterior margin slightly depressed medially; eye of small to moderate size (EL 0.09-0.11, REL 15-19), oval-shaped, with 4-7 ommatidia at greatest diameter; face densely sculptured, mostly rugoreticulate, with a few longitudinal rugae along midline near frontal lobes; scape of moderate length (SI 88-97), almost reaching posterior margin of head when laid back; scape surface mostly smooth, with scattered piligerous punctae; flagellum with a distinct 4-segmented antennal club; most of mesosoma densely sculptured, dorsum of promesonotum rugoreticulate, or with irregular longitudinal rugae that only sometimes merge, side of pronotum variably rugulose-punctate, sometimes with a large patch of smooth cuticle, mesopleuron mostly punctate, with a few rugulae,
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<pageBreakToken pageId="165" pageNumber="166" start="start">side</pageBreakToken>
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of propodeum rugulose-punctate, propodeal dorsum and upper half of declivity with a few transverse carinulae; promesonotum in profile low-domed, roughly symmetrical, sometimes dorsum slightly flattened; metanotal groove well-demarcated, moderate; propodeal spines tuberculate, often forming somewhat broad triangles (PSL 0.08-0.14, PSI 1.1-1.4); petiole appearing of moderate length (PL/HW 0.50-0.59), peduncle usually somewhat slender; petiolar node of moderate height to somewhat small (PH/PL 0.54-0.65), dorsum broadly rounded, and always pointing distinctly posteriad, area underneath node usually with a small concavity, or notch; postpetiole in profile subspherical to oblong, appearing slightly compressed anteroposteriorly; anterior faces of petiolar and postpetiolar nodes smooth, remaining surfaces mostly punctate, with a few rugulae on posterior half of postpetiole; gaster smooth, with scattered piligerous punctae; face with a mostly uniform layer of short decumbent setae; remainder of body dorsum with longer suberect setae; pilosity on gastral tergites indistinctly bilayered, with a relatively sparse layer of longer suberect to subdecumbent setae, and a sparser layer of decumbent to appressed setae; setae on scapes and legs mostly decumbent to appressed, with some suberect setae on femoral venters and coxae.
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</paragraph>
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<caption pageId="165" pageNumber="166">
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<paragraph pageId="165" pageNumber="166">
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Figure 131.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="165" pageNumber="166" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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holotype worker (CASENT060578) A Profile B Face C Dorsum D Anterior clypeal margin in anterodorsal view E Gaster.
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</paragraph>
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</caption>
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<caption pageId="165" pageNumber="166">
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<paragraph pageId="165" pageNumber="166">
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Figure 132.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="165" pageNumber="166" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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worker variants. Face, profile, dorsal views
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Variant 1 (CASENT0603920)
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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Variant 2 (CASENT0603896)
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<normalizedToken originalValue="G–I">G-I</normalizedToken>
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Variant 3 (CASENT0604711).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="166" lastPageNumber="167" pageId="165" pageNumber="166" type="queen description">
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<paragraph pageId="165" pageNumber="166">Queen description.</paragraph>
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<paragraph lastPageId="166" lastPageNumber="167" pageId="165" pageNumber="166">
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(5 measured) HL 0.61-0.76 (0.61), HW 0.56-0.70 (0.57), FLD 0.15-0.21 (0.16), PCW 0.04-0.08 (0.04), SL 0.50-0.64 (0.50), EL 0.15-0.20 (0.15),
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<pageBreakToken pageId="166" pageNumber="167" start="start">ACL</pageBreakToken>
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0.48-0.61 (0.49), ML 0.84-1.11 (0.86), PrW 0.49-0.63 (0.49), PSL 0.11-0.16 (0.14), SDL 0.09-0.12 (0.10), PL 0.33-0.44 (0.34), PH 0.19-0.25 (0.19), PW 0.17-0.22 (0.18), PPL 0.15-0.22 (0.18), PPH 0.19-0.25 (0.19), PPW 0.18-0.27 (0.21), MFL 0.55-0.73 (0.55), MTL 0.45-0.58 (0.45), CI 89-94 (94), SI 86-96 (86), REL 26-28 (26), FLI 27-31 (27), PSI 1.3-1.4 (1.4), MFI 93-104 (104), ACI1 64-67 (66), ACI2 94-98 (98).
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</paragraph>
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<paragraph pageId="166" pageNumber="167">Same as worker except for standard queen modifications and as follows: side of pronotum rugoreticulate to transversely rugose, center of pronotum transversely carinulate; mesoscutum mostly foveolate to longitudinally rugose, usually with some smooth cuticle along midline, foveolae when present piligerous; propodeum with transverse carinulae that wrap around surface; mesopleuron mostly smooth; pilosity on gastral tergites distinctly bilayered, with a sparse layer of longer suberect to subdecumbent setae and a dense layer of decumbent setae, lower layer usually almost pubescent, but sometimes less dense and longer; wing venation as in Figure 133D.</paragraph>
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<caption pageId="166" pageNumber="167">
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<paragraph pageId="166" pageNumber="167">
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Figure 133.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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A Queen (CASENT0605471), profile B Same, face C Same, dorsum D Queen (CASENT0605470), forewing.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="166" pageNumber="167" type="male">
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<paragraph pageId="166" pageNumber="167">Male.</paragraph>
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<paragraph pageId="166" pageNumber="167">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="166" pageNumber="167" type="biology">
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<paragraph pageId="166" pageNumber="167">Biology.</paragraph>
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<paragraph pageId="166" pageNumber="167">This species is known only from extracts of sifted leaf litter collected from the forest floor. It occupies a variety of wet forest habitat types (e.g. rainforest, mesophyll forest, 2° wet forest, Liquidambar forest, cloud forest, oak-pine forest) and occurs from near sea level to 2000 m. Dealate queens are commonly collected in samples, suggesting that the species may nest in the leaf litter microenvironment.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="167" lastPageNumber="168" pageId="166" pageNumber="167" type="comments">
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<paragraph pageId="166" pageNumber="167">Comments.</paragraph>
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<paragraph pageId="166" pageNumber="167">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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forms a clade with
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nanozoi" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="nanozoi">Stenamma nanozoi</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma sandinista" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="sandinista">Stenamma sandinista</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma schmidti" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="schmidti">Stenamma schmidti</taxonomicName>
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and is very similar to these species (Branstetter unpublished data). However, it is the only species within the clade to have the anterior clypeal margin entire, and the basal margin of the mandible straight. This feature makes it easy to distinguish from its close relatives.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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is also similar to
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma ignotum" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="ignotum">Stenamma ignotum</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma picopicucha" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="picopicucha">Stenamma picopicucha</taxonomicName>
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, which both have a straight basal margin of the mandible. However, neither of these species have the anterior clypeal margin entire.
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</paragraph>
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<paragraph lastPageId="167" lastPageNumber="168" pageId="166" pageNumber="167">
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There is some variation among populations of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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, with workers differing in size, sculpture, and shape of the propodeal spines and petiolar node. I split this diversity into three variant forms. The type form (Figure 131) includes all of the collections from Oaxaca, Mexico, which were collected below 700 m on the wetter side of the state. Variant 1 (Figure 132
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) occurs at the low-elevation locality Salto de Agua in Chiapas, Mexico. Compared to the type population, specimens are smaller, have a lighter red-brown coloration, have the propodeal spines slightly more robust and projecting dorsally, and have the petiolar node in profile somewhat anteroposteriorly compressed. Variant 2 (Figure 132
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) occurs at the mid-elevation locality
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<normalizedToken originalValue="Nahá">Naha</normalizedToken>
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in Chiapas, Mexico. It is smaller than the type form and has the propodeal spines noticeably reduced to sharp angles, rather than sharp, projecting tubercles. Lastly, variant 3 (Figure 132
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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) is a high elevation version of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma nonotch" order="Hymenoptera" pageId="166" pageNumber="167" phylum="Arthropoda" rank="species" species="nonotch">Stenamma nonotch</taxonomicName>
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, occurring mainly at the locality Coapilla in Chiapas, Mexico. Compared with the type form, it is larger, has darker coloration, and has a thicker petiolar node. Also, the ventral margin of the petiolar node is flat, rather than with a small concavity. Molecular data suggest that there is significant divergence among
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<pageBreakToken pageId="167" pageNumber="168" start="start">populations</pageBreakToken>
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, but I have not seen sympatry among forms, and some populations have intermediate phenotypes. Thus, I treat the above as intraspecific variants until more data are available.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="167" pageNumber="168" type="material examined">
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<paragraph pageId="167" pageNumber="168">Material examined.</paragraph>
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<paragraph pageId="167" pageNumber="168">
|
||
<normalizedToken originalValue="MÉXICO">MEXICO</normalizedToken>
|
||
: Chiapas: 15.1km NW Bochil, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="17.0912">17.0912°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-92.9914">92.9914°W</geoCoordinate>
|
||
], 1930m, 24 Sep 1992 (R. S. Anderson); 5km NE Coapilla,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.17558">17.17558°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-93.15012">93.15012°W</geoCoordinate>
|
||
, 1770m, 26 May 2008 (M. G. Branstetter); 5km NE Coapilla,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.17602">17.17602°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-93.13293">93.13293°W</geoCoordinate>
|
||
, 1990m, 25 May 2008 (LLAMA); Lagos de Montebello, Cinco Lagos,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="16.1012">16.1012°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-91.674">91.6740°W</geoCoordinate>
|
||
, 1600m, 21 Jul 2007 (R. S. Anderson); Lagos de Montebello, L. Pojoj,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.09913">16.09913°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-91.67125">91.67125°W</geoCoordinate>
|
||
, 1510m, 21 Jul 2007 (R. S. Anderson);
|
||
<normalizedToken originalValue="Nahá">Naha</normalizedToken>
|
||
,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="16.96426">16.96426°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-91.59234">91.59234°W</geoCoordinate>
|
||
, 985m, 8 Jun 2008 (LLAMA); 6km SW Ocosingo,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="16.8672">16.8672°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-92.0787">92.0787°W</geoCoordinate>
|
||
, 1400m, 22 Sep 1991 (R. S. Anderson); 2km S
|
||
<normalizedToken originalValue="Playón">Playon</normalizedToken>
|
||
de la Gloria,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="16.1385">16.1385°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-90.9015">90.9015°W</geoCoordinate>
|
||
, 170m, 22 Jul 2007 (R. S. Anderson); 13km N Pueblo Nuevo
|
||
<normalizedToken originalValue="Solistahuacán">Solistahuacan</normalizedToken>
|
||
, [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="17.211">17.211°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-92.964">92.964°W</geoCoordinate>
|
||
], 1860m, 26-27 Aug 1973 (A. F. Newton); 5km E
|
||
<normalizedToken originalValue="Rayón">Rayon</normalizedToken>
|
||
,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="17.217">17.217°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-92.967">92.967°W</geoCoordinate>
|
||
, 1700m, 23 Dec 1991 (P. S. Ward); 8.9km E Rayon,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="17.2">17.2000°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-92.9163">92.9163°W</geoCoordinate>
|
||
, 1500m, 19 Sep 1991 (R. S. Anderson); 21km SW Salto de Agua,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.38542">17.38542°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-92.42802">92.42802°W</geoCoordinate>
|
||
, 180m, 15 Jun 2008 (M. G. Branstetter);Oaxaca: 4.3km SW Valle Nacional,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.7432">17.74320°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.32729">96.32729°W</geoCoordinate>
|
||
, 370m, 13 Aug 2009 (L.
|
||
<normalizedToken originalValue="Sáenz">Saenz</normalizedToken>
|
||
); 7.5km S Valle Nacional,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="17.70752">17.70752°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-96.30516">96.30516°W</geoCoordinate>
|
||
, 680m, 12 Aug 2009 (M. G. Branstetter); Veracruz: Los Tuxtlas, 10km NNW Sontecomapan,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="55" value="18.583">18.583°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-95.083">95.083°W</geoCoordinate>
|
||
, 200m, 20 Mar 1985 (P. S. Ward).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |