treatments-xml/data/28/86/3F/28863FA1D53E5C49A6C2A1A77875FF02.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="28863FA1D53E5C49A6C2A1A77875FF02" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Effossana circumnota Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="F67F1709-2F57-497D-BFD2-D1C5730F75EF" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
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<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="https://zoobank.org/F67F1709-2F57-497D-BFD2-D1C5730F75EF" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Effossana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Effossana circumnota" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="circumnota" status="sp. nov.">Effossana circumnota</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
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<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">Figs 37</figureCitation>
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">, 38</figureCitation>
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Species of Effossana gen. nov., dorsal (left) and lateral (right) views. A, B E. assimulata sp. nov., holotype male, DZUP. C, D E. circumnota sp. nov., holotype male, DZUP. E, F E. fructa comb. nov., holotype male, USNM. G, H E. gatuna comb. nov., holotype male, OSU." figureDoi="10.3897/asp.81.e81961.figure42" httpUri="https://binary.pensoft.net/fig/798195" pageId="0" pageNumber="79">, 42C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37C</figureCitation>
), crown-face transition thick. Pygofer (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37G</figureCitation>
) without processes. Style, in lateral view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37K</figureCitation>
), ventral margin with subapical process truncated apically and serrated on anterior margin. Aedeagus (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37L</figureCitation>
) with shaft very long, basal portion strongly curved forming wide ring.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 9.2 mm; paratypes, males (n = 28) 8.7-10.3 mm, females (n = 12) 9.6-10.4 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37A-B</figureCitation>
, 
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Species of Effossana gen. nov., dorsal (left) and lateral (right) views. A, B E. assimulata sp. nov., holotype male, DZUP. C, D E. circumnota sp. nov., holotype male, DZUP. E, F E. fructa comb. nov., holotype male, USNM. G, H E. gatuna comb. nov., holotype male, OSU." figureDoi="10.3897/asp.81.e81961.figure42" httpUri="https://binary.pensoft.net/fig/798195" pageId="0" pageNumber="79">42C, D</figureCitation>
) yellowish brown. Crown (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37A</figureCitation>
) with coronal suture black. Face (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37B</figureCitation>
) gena yellow, brown maculae near subgenal suture, ventral and inner margins of eye. Pronotum (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37A</figureCitation>
) with brown punctures, pair of elongated and oblique black maculae behind ocelli, near anterior margin; posterior margin with black transverse band; proepimeron (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37C</figureCitation>
), with dorsal half black. Mesonotum (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37A</figureCitation>
) with yellow markings, pair of black ring-shaped maculae near lateral angles and pair of rounded black spots, near scutoscutelar suture. Scutellum (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37A</figureCitation>
) black with large yellow macula. Forewing (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37D</figureCitation>
) strongly marked by dark-brown mottling; membrane translucent light yellow; veins brown outlined by dark brown. Metatibia (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in dorsal view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37A</figureCitation>
) transocular width 8.2 tenths of humeral width of pronotum. Head, in frontal view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37B</figureCitation>
), frons 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; clypeus 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; lateral margins slightly divergent toward apex. Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37C</figureCitation>
), crown-face transition thick, smooth medially and with few striae near of eye. Forewing (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37D</figureCitation>
) with appendix reduced, narrower than the width of the first apical cell. Protibia AD row without differentiated setae. Metatibia PD, AD and AV rows with 23-26, 12-13 and 16-17 macrosetae respectively. Metatarsomere I moderately elongated, about 3 
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of ventral surface with 5-6 setae; pecten with 6-7 platellae. Metatarsomere II pecten with 3 platellae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37E</figureCitation>
) 1.7 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, lateral margins weakly rounded; posterior margin with median third deeply excavated and truncate. Valve (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37F</figureCitation>
) 3.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin narrowly excavated; in lateral view, strongly convex. Pygofer, in lateral view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37G</figureCitation>
), 1.8 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; anteroventral margin rounded; posterodorsal and posteroventral margins slightly rounded; apex broadly rounded. Subgenital plate, in ventral view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37H</figureCitation>
), elongated, 4.4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near half length; inner margin straight; outer margin slightly rounded; apex tapered and rounded. Connective (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37I</figureCitation>
) with thin arms. Style, in lateral view (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37K</figureCitation>
), with blade narrow tapered toward apex; ventral margin with subapical process truncated apically and serrated on anterior margin; apex slightly curved dorsally, abruptly tapered, and acute. Aedeagus (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37L, M</figureCitation>
) dorsal apodeme reduced, dorsal margin excavated, lateral margins not projected laterally; apodemal process elongated and curved dorsally, dorsal margin excavated medially, apex rounded; shaft cylindrical and very long, basal portion strongly curved forming wide ring, portion after the ring straight; apex twisted laterally, with two pairs of processes directed dorsally: one apical very short and one subapical 3 
<normalizedToken originalValue="×">x</normalizedToken>
longer than the apical. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female terminalia">
<paragraph pageId="0" pageNumber="79">Female terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VII (Fig. 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">38A</figureCitation>
) 1.7 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterolateral angles produced as far as posterior margin; posterior margin deeply V-shaped excavated each side of triangular median lobe. Internal sternite VIII membranous. Pygofer (Fig. 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">38A, B</figureCitation>
) 1.9 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; apex rounded. First valvifer (Fig. 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">38C</figureCitation>
) sub-rectangular 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; anterior and dorsal margins straight; posterior margin rounded. First valvula (Fig. 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">38C</figureCitation>
) 4.1 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Second valvula (Fig. 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">38E</figureCitation>
) 4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; dorsal protuberance rounded. Gonoplac (Fig. 
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Effossana circumnota sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure38" httpUri="https://binary.pensoft.net/fig/798191" pageId="0" pageNumber="79">38G</figureCitation>
) 3.6 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Other characteristics as in the diagnosis of the genus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the ringed basal portion of the aedeagal shaft.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2011-03-05" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="1" stateProvince="Goias" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male: 
<collectingCountry name="Brazil">Brazil</collectingCountry>
: 
<normalizedToken originalValue="Goiás">
<collectingRegion country="Brazil" name="Goias">Goias</collectingRegion>
</normalizedToken>
</emphasis>
: &quot;Brasil, GO, Novo Mundo,\ Armadilha Malaise\ 
<geoCoordinate degrees="13.9292" direction="south" orientation="latitude" precision="5" value="-13.9292">13.9292°S</geoCoordinate>
<geoCoordinate degrees="49.9716" direction="west" orientation="longitude" precision="5" value="-49.9716">49.9716°W</geoCoordinate>
,\ 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.82" unit="m" value="282.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.82" unit="m" value="282.0">282m</elevation>
</quantity>
, 
<collectingDate value="2011-03-05">05.III.2011</collectingDate>
&quot; (DZUP)
</materialsCitation>
. 
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" elevation="282" latitude="-13.9292" location="Paratypes" longLatPrecision="7" longitude="-49.9716" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Paratypes">
<typeStatus>Paratypes</typeStatus>
</collectingRegion>
</emphasis>
: 
<specimenCount type="male">1♂</specimenCount>
, same data as holotype (DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="2010-11-26" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">2♀</specimenCount>
, same data as holotype except 
<collectingDate value="2010-11-26">26.XI.2010</collectingDate>
(DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="2010-12-03" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
same data as holotype except 
<collectingDate value="2010-12-03">03.XII.2010</collectingDate>
(DZRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2010-12-10" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, same data as holotype except 
<collectingDate value="2010-12-10">10.XII.2010</collectingDate>
(DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="2011-01-03" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as holotype except 
<collectingDate value="2011-01-03">03.I.2011</collectingDate>
(DZRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2012-03-10" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="2" specimenCount-female="2" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="female">2♀</specimenCount>
, same data as holotype except 
<collectingDate value="2012-03-10">10.III.2012</collectingDate>
(DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="2012-10-06" country="Brazil" elevation="282" latitude="-13.9292" location="Goias" longLatPrecision="7" longitude="-49.9716" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="male">4♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
, same data as holotype except 
<collectingDate value="2012-10-06">06.X.2012</collectingDate>
(
<specimenCount type="male">3♂</specimenCount>
DZUP, 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
MNRJ)
</materialsCitation>
; 
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" county="Brasil" elevation="282" isTailEnumeration="true" latitude="-13.9292" location="Sao Domingos" longLatPrecision="7" longitude="-49.9716" municipality="Goias" specimenCount="18" specimenCount-male="18" stateProvince="Distrito Federal" typeStatus="paratype">
<specimenCount type="male">18♂</specimenCount>
, &quot; 
<collectingCounty>Brasil</collectingCounty>
, 
<normalizedToken originalValue="Goiás">
<collectingMunicipality>Goias</collectingMunicipality>
</normalizedToken>
\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:6EB93A8B7A90D083AA751644129455E1" country="Brazil" county="Brasil" latitude="-13.9292" longLatPrecision="7" longitude="-49.9716" municipality="Goias" name="Sao Domingos" stateProvince="Distrito Federal">
<normalizedToken originalValue="São">Sao</normalizedToken>
Domingos
</location>
\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:D7C2515E5D7FF377934F1ACB472BB29A" country="Brazil" county="Brasil" latitude="-13.9292" longLatPrecision="7" longitude="-49.9716" municipality="Goias" name="Fazenda Cipasa" stateProvince="Distrito Federal">Fazenda Cipasa</location>
\ 18-IX a 2-X-93&quot;, 
<normalizedToken originalValue="“Dionísio">
&quot; 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:B756BB8609A63DE3AD8AFF2B3D60C18C" country="Brazil" county="Brasil" latitude="-13.9292" longLatPrecision="7" longitude="-49.9716" municipality="Goias" name="Dionisio Pimentel" stateProvince="Distrito Federal">Dionisio Pimentel</location>
</normalizedToken>
&quot;, 
<normalizedToken originalValue="“Malaise”">
&quot; 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:2A98A05215936CD2EEB14F55B10E6DB7" country="Brazil" county="Brasil" latitude="-13.9292" longLatPrecision="7" longitude="-49.9716" municipality="Goias" name="Malaise" stateProvince="Distrito Federal">Malaise</location>
&quot;
</normalizedToken>
, &quot; MPEG 05029105, 05029106, 05029109, 05029113-05029117, 05029119-05029121, 05029123-05029128 and 05029132&quot; (MPEG). - 
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Distrito Federal">Distrito Federal</collectingRegion>
</emphasis>
</materialsCitation>
: 
<materialsCitation collectingDate="2012-05-31" country="Brazil" county="Brasil" latitude="-15.9594" location="Faz." longLatPrecision="7" longitude="-47.9328" municipality="Brasilia" specimenCount="1" specimenCount-male="1" stateProvince="Distrito Federal" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<collectingCounty>Brasil</collectingCounty>
, 
<normalizedToken originalValue="Brasília">
<collectingMunicipality>Brasilia</collectingMunicipality>
</normalizedToken>
- DF, 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:C3C088F36566E4F3DEE55178F921B96C" country="Brazil" county="Brasil" latitude="-15.9594" longLatPrecision="7" longitude="-47.9328" municipality="Brasilia" name="Faz." stateProvince="Distrito Federal">Faz.</location>
\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:3B3E6BBCECCD2169C51542E37D98E372" country="Brazil" county="Brasil" latitude="-15.9594" longLatPrecision="7" longitude="-47.9328" municipality="Brasilia" name="Agua Limpa" stateProvince="Distrito Federal">
<normalizedToken originalValue="Água">Agua</normalizedToken>
Limpa
</location>
, 
<geoCoordinate degrees="15.9594" direction="south" orientation="latitude" precision="5" value="-15.9594">15.9594°S</geoCoordinate>
<geoCoordinate degrees="47.9328" direction="west" orientation="longitude" precision="5" value="-47.9328">47.9328°W</geoCoordinate>
, 
<collectingDate value="2012-05-31">31.V.2012</collectingDate>
\ Malaise, A.J.C. Aguiar&quot; (DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="2017-11-23" collectorName="J. R. P. Luz" country="Brazil" county="Brasil" location="Fazenda Agua Limpa" municipality="Brasilia" specimenCount="1" specimenCount-female="1" stateProvince="Distrito Federal" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, &quot; 
<collectingCounty>Brasil</collectingCounty>
, 
<normalizedToken originalValue="Brasília">
<collectingMunicipality>Brasilia</collectingMunicipality>
</normalizedToken>
- DF,\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:026FD59139C8456EFC36C06C7FB1E764" country="Brazil" county="Brasil" municipality="Brasilia" name="Fazenda Agua Limpa" stateProvince="Distrito Federal">
Fazenda 
<normalizedToken originalValue="Água">Agua</normalizedToken>
Limpa
</location>
,\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:30CE0FAE4619616418234C75F0EDE452" country="Brazil" county="Brasil" municipality="Brasilia" name="Cerrado" stateProvince="Distrito Federal">Cerrado</location>
, malaise, 09-\ 
<collectingDate value="2017-11-23">23.XI.2017</collectingDate>
, 
<collectorName>J.R.P. Luz</collectorName>
&quot; (DZUP)
</materialsCitation>
; 
<specimenCount type="female">1♀</specimenCount>
, same as preceding except 
<date value="2017-12-22">22.XII.2017</date>
(DZUP). - 
<materialsCitation collectingDate="2021-08-18" collectingDateMax="2021-09-24" collectingDateMin="2021-08-18" collectorName="R. N. Guedes, H. C. Onody" country="Brazil" county="Brasil" latitude="-10.43412" location="Corrente" longLatPrecision="1" longitude="-45.16392" municipality="Piaui" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Piaui" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="79">
<normalizedToken originalValue="Piauí">
<collectingRegion country="Brazil" name="Piaui">Piaui</collectingRegion>
</normalizedToken>
</emphasis>
: 
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">2♀</specimenCount>
, &quot; 
<collectingCounty>Brasil</collectingCounty>
: 
<normalizedToken originalValue="Piauí">
<collectingMunicipality>Piaui</collectingMunicipality>
</normalizedToken>
, 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:2894C0B7FAE3A02D85A9DFF9E0D42777" country="Brazil" county="Brasil" latitude="-10.43412" longLatPrecision="1" longitude="-45.16392" municipality="Piaui" name="Corrente" stateProvince="Piaui">Corrente</location>
, PARNA \ das 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:4D76CA30F57BE39F41D822E7AC4F8C0D" country="Brazil" county="Brasil" latitude="-10.43412" longLatPrecision="1" longitude="-45.16392" municipality="Piaui" name="Nascentes do Rio Parnaiba" stateProvince="Piaui">
Nascentes do Rio 
<normalizedToken originalValue="Parnaíba">Parnaiba</normalizedToken>
</location>
\ - 
<geoCoordinate degrees="10.43412" direction="south" orientation="latitude" precision="1" value="-10.43412">10.43412 S</geoCoordinate>
, - 
<geoCoordinate degrees="45.16392" direction="west" orientation="longitude" precision="1" value="-45.16392">45.16392 W</geoCoordinate>
Tri- \ lha 1, 
<collectingDate value="2021-08-18" valueMax="2021-09-24" valueMin="2021-08-18">18.viii-24.ix.2021</collectingDate>
Malaise \ 
<collectorName>R.N.Guedes</collectorName>
&amp; 
<collectorName>H.C.Onody</collectorName>
cols&quot; (
<specimenCount type="male">1♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
DZUP, 
<specimenCount type="female">1♀</specimenCount>
UESPI). - 
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Tocantins">Tocantins</collectingRegion>
</emphasis>
</materialsCitation>
: 
<materialsCitation collectingDate="2012-08-03" collectingDateMax="2012-08-10" collectingDateMin="2012-08-03" collectorName="Lima, H. I. L." country="Brazil" county="Brasil" latitude="-10.250639" location="Palmas" longLatPrecision="1" longitude="-48.126" municipality="Tocantins" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<collectingCounty>Brasil</collectingCounty>
, 
<collectingMunicipality>Tocantins</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:A2C4A2D2D7B55261ADDF1DBCC03CD922" country="Brazil" county="Brasil" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Tocantins" name="Palmas" stateProvince="Minas Gerais">Palmas</location>
\ 
<normalizedToken originalValue="Taquaruçu">
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:F99CF6C65F47A1C8CECBF0AE9CC38B12" country="Brazil" county="Brasil" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Tocantins" name="Taquarucu" stateProvince="Minas Gerais">Taquarucu</location>
</normalizedToken>
, 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:9DF71AEFB22F83AE0E5298D10577A150" country="Brazil" county="Brasil" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Tocantins" name="Fazenda" stateProvince="Minas Gerais">Fazenda</location>
\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:0DEFDBFA6DFF87E16A5270A158A837B5" country="Brazil" county="Brasil" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Tocantins" name="Encantada" stateProvince="Minas Gerais">Encantada</location>
<geoCoordinate degrees="10" direction="south" minutes="15" orientation="latitude" precision="1" seconds="2.3" value="-10.250639">10°15′2.3″S</geoCoordinate>
<geoCoordinate degrees="48" direction="west" minutes="07" orientation="longitude" precision="1" seconds="33.6" value="-48.126">48°07′33.6″W</geoCoordinate>
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:6BF421FCD6BE3405CA5833FB73D5C3A8" country="Brazil" county="Brasil" latitude="-10.250639" longLatPrecision="1" longitude="-48.126" municipality="Tocantins" name="Malaise" stateProvince="Minas Gerais">Malaise</location>
\ 
<collectingDate value="2012-08-03" valueMax="2012-08-10" valueMin="2012-08-03">03-10.viii.2012</collectingDate>
Krolow,\ T.K. &amp; 
<collectorName>Lima, H.I.L.</collectorName>
leg.&quot; (DZUP). - 
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Minas Gerais">Minas Gerais</collectingRegion>
</emphasis>
</materialsCitation>
: 
<materialsCitation collectingDate="1983-12" collectorName="M. Alvarenga" country="Brazil" county="Brasil" location="Gerais" municipality="Minas" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, &quot; 
<collectingCounty>Brasil</collectingCounty>
, 
<collectingMunicipality>Minas</collectingMunicipality>
\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:502FBD2029F335BBDFCE17890AB18126" country="Brazil" county="Brasil" municipality="Minas" name="Gerais" stateProvince="Minas Gerais">Gerais</location>
, 
<normalizedToken originalValue="Águas">
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:7D3FA9A62C026970B3BEC28634C656D6" country="Brazil" county="Brasil" municipality="Minas" name="Aguas" stateProvince="Minas Gerais">Aguas</location>
</normalizedToken>
\ 
<location LSID="urn:lsid:plazi:treatment:28863FA1D53E5C49A6C2A1A77875FF02:FB336708C237CA761E5A5BEA6A164996" country="Brazil" county="Brasil" municipality="Minas" name="Vermelhas" stateProvince="Minas Gerais">Vermelhas</location>
<collectingDate value="1983-12">xii.1983</collectingDate>
\ 
<collectorName>M. Alvarenga</collectorName>
leg.&quot; (DZUP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Effossana" kingdom="Animalia" lsidName="Effossana circumnota" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="circumnota">
<emphasis italics="true" pageId="0" pageNumber="79">Effossana circumnota</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
was recovered as sister to 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. intrinseca" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="intrinseca">
<emphasis italics="true" pageId="0" pageNumber="79">E. intrinseca</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
with high branch support (SR = 99), sustained by one synapomorphy, 134:2, aedeagal shaft with basal portion directed anterad, and four homoplasies, as for example: 58:0, forewing with appendix reduced; and 135:0, aedeagal shaft with apical portion straight in lateral view. 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Effossana" kingdom="Animalia" lsidName="Effossana circumnota" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="circumnota">
<emphasis italics="true" pageId="0" pageNumber="79">Effossana circumnota</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can easily be separated from 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="E. intrinseca" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="intrinseca">
<emphasis italics="true" pageId="0" pageNumber="79">E. intrinseca</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
by the pygofer (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37G</figureCitation>
) without processes and the style (Fig. 
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Effossana circumnota sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure37" httpUri="https://binary.pensoft.net/fig/798190" pageId="0" pageNumber="79">37K</figureCitation>
) with ventral margin of blade forming a truncated subapical process.
</paragraph>
</subSubSection>
</treatment>
</document>