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<document id="AF9F92C874F9A75D3F5D966C3E9AC4D4" ID-DOI="10.5281/zenodo.5326103" ID-ISSN="0253-116X" ID-Zenodo-Dep="5326103" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630152639660" checkinUser="carolina" docAuthor="Assing, V." docDate="2011" docId="281387FAFF86FFFFFF73F6901290308F" docLanguage="en" docName="LinzerbiolBeitr.2011.43.2.1179-1197.pdf" docOrigin="Linzer biologische Beiträge 43 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Tetartopeus bimaculatus" docType="treatment" docVersion="3" lastPageNumber="1191" masterDocId="D42AFF82FF8DFFF3FFC1F5051062347B" masterDocTitle="On some East Palaearctic Tetartopeus species (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1197" masterPageNumber="1179" pageNumber="1190" updateTime="1699165773312" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A237FF4568DF3C0F5AB8A37C333970C3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="36A34AEBA7F5794023810263FB374A03">
<mods:title id="95CEB1D8BAA053C28045A69A91AA3469">On some East Palaearctic Tetartopeus species (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="299EF2465F1FC6B3254B8DADB58AFFD2">Assing, V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2B7B8A92ECF8C5E7C2D5EA7522DC6FB9">text</mods:typeOfResource>
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<mods:title id="A70E9BE6BA9B0FAB0EA3C46210442EC7">Linzer biologische Beiträge</mods:title>
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<mods:part id="5BA9F31046035F8DD534A147B10CAC7D">
<mods:date id="18DCA8C2E94BA616F495D844443805AD">2011</mods:date>
<mods:detail id="F02A7CF7EAF401E003369A2848FA218E" type="pubDate">
<mods:number id="AAA68F7A5AE3E40C56D78E295A9C7266">2011-12-19</mods:number>
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<mods:number id="4599CC71EDA8D05E82D8FB2EE370B374">43</mods:number>
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<mods:classification id="B0DC431C1609A8FD90099391D7D5720F">journal article</mods:classification>
<mods:identifier id="31A19BC756913566C09FD42BC778E0A0" type="DOI">10.5281/zenodo.5326103</mods:identifier>
<mods:identifier id="95D8511F810299D501BF553C31AE9A98" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="79038D9581E948A92DB3B967AB009EFA" type="Zenodo-Dep">5326103</mods:identifier>
</mods:mods>
<treatment id="281387FAFF86FFFFFF73F6901290308F" LSID="urn:lsid:plazi:treatment:281387FAFF86FFFFFF73F6901290308F" httpUri="http://treatment.plazi.org/id/281387FAFF86FFFFFF73F6901290308F" lastPageId="12" lastPageNumber="1191" pageId="11" pageNumber="1190">
<subSubSection id="E8A06567FF86FFF8FF73F690133C37D7" box="[178,862,916,941]" pageId="11" pageNumber="1190" type="nomenclature">
<paragraph id="A00536ECFF86FFF8FF73F690133C37D7" blockId="11.[178,862,916,941]" box="[178,862,916,941]" pageId="11" pageNumber="1190">
<heading id="FB4D8180FF86FFF8FF73F690133C37D7" bold="true" box="[178,862,916,941]" fontSize="9" level="2" pageId="11" pageNumber="1190" reason="3">
<taxonomicName id="67BA4D6FFF86FFF8FF73F69012B037D7" authority="(LI, TANG &amp; ZHU 2007)" baseAuthorityName="LI, TANG &amp; ZHU" baseAuthorityYear="2007" box="[178,722,916,941]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1190" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="92CEEAFEFF86FFF8FF73F69012B037D7" bold="true" box="[178,722,916,941]" pageId="11" pageNumber="1190">
<emphasis id="92CEEAFEFF86FFF8FF73F69011A037D7" bold="true" box="[178,450,916,940]" italics="true" pageId="11" pageNumber="1190">Tetartopeus bimaculatus</emphasis>
(LI, TANG &amp; ZHU 2007)
</emphasis>
</taxonomicName>
(
<figureCitation id="38812A69FF86FFF8FD21F690133437D7" box="[736,854,917,940]" captionStart-0="Figs 39-47" captionStart-1="Figs 48-51" captionStartId-0="10.[178,218,1431,1453]" captionStartId-1="11.[178,218,810,832]" captionTargetBox-0="[184,1076,212,1413]" captionTargetBox-1="[182,1073,212,796]" captionTargetId-0="figure-79@10.[178,1085,212,1422]" captionTargetId-1="figure-365@11.[178,1085,212,800]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." captionText-1="Figs 48-51: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Yunnan (48-49), Taiwan (50) and from Hunan (51): (48) aedeagus in lateral view; (49-51) aedeagus in dorsal view. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5326117" figureDoi-1="http://doi.org/10.5281/zenodo.5326119" httpUri-0="https://zenodo.org/record/5326117/files/figure.png" httpUri-1="https://zenodo.org/record/5326119/files/figure.png" pageId="11" pageNumber="1190">Figs 39-51</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E8A06567FF86FFF8FF73F6B912BC30F3" pageId="11" pageNumber="1190" type="reference_group">
<paragraph id="A00536ECFF86FFF8FF73F6B912CC37A9" blockId="11.[178,686,956,1004]" box="[178,686,956,978]" pageId="11" pageNumber="1190">
<taxonomicName id="67BA4D6FFF86FFF8FF73F6B912F837A9" ID-CoL="6QLMT" authority="LI, TANG &amp; ZHU 2007: 261" authorityName="LI, TANG &amp; ZHU" authorityPageNumber="261" authorityYear="2007" box="[178,666,956,978]" class="Insecta" family="Staphylinidae" genus="Lobrathium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1190" phylum="Arthropoda" rank="species" species="bimaculatum">
<emphasis id="92CEEAFEFF86FFF8FF73F6B911F837AA" box="[178,410,956,977]" italics="true" pageId="11" pageNumber="1190">Lobrathium bimaculatum</emphasis>
LI, TANG &amp; ZHU 2007:
<specimenCount id="B6BCFD65FF86FFF8FDB9F6BB12F837A9" box="[632,666,958,978]" pageId="11" pageNumber="1190" type="generic">261</specimenCount>
</taxonomicName>
f.
</paragraph>
<paragraph id="A00536ECFF86FFF8FF73F6D212583797" blockId="11.[178,686,956,1004]" box="[178,570,982,1004]" pageId="11" pageNumber="1190">
<taxonomicName id="67BA4D6FFF86FFF8FF73F6D212543797" authority=": ASSING (2010 b)" authorityName=": ASSING" authorityYear="2010" box="[178,566,982,1004]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1190" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="92CEEAFEFF86FFF8FF73F6D211F13790" box="[178,403,982,1004]" italics="true" pageId="11" pageNumber="1190">Tetartopeus bimaculatus</emphasis>
:
<bibRefCitation id="C42B4B1DFF86FFF8FE5EF6DD12543797" author="ASSING V" box="[415,566,984,1004]" pageId="11" pageNumber="1190" pagination="301 - 361" refId="ref8378" refString="ASSING V. (2010 b): On the Lathrobiina of Taiwan (Coleoptera: Staphylinidae: Paederinae). - Beitrage zur Entomologie, Keltern 60 (2): 301 - 361." type="journal article" year="2010">ASSING (2010b)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="A00536ECFF86FFF8FF73F6F812BC30F3" blockId="11.[178,1086,1021,1360]" pageId="11" pageNumber="1190">
C o m m e n t: The illustrations of the male sexual characters provided in the original description, which is based on a single male
<typeStatus id="7F01884EFF86FFF8FD45F11F12BF304B" box="[644,733,1050,1072]" pageId="11" pageNumber="1190" type="holotype">holotype</typeStatus>
from
<collectingRegion id="627EF80EFF86FFF8FCDFF11F131A304B" box="[798,888,1050,1072]" country="China" name="Guizhou" pageId="11" pageNumber="1190">Guizhou</collectingRegion>
, are schematic and unsuitable for an identification of this species. It was only after Liang Tang (
<collectingRegion id="627EF80EFF86FFF8FC0EF13214503036" box="[975,1074,1079,1101]" country="China" name="Shanghai" pageId="11" pageNumber="1190">Shanghai</collectingRegion>
), one of the authors, kindly sent photographs of the primary and secondary sexual characters of the
<typeStatus id="7F01884EFF86FFF8FEDCF177111430F3" box="[285,374,1138,1160]" pageId="11" pageNumber="1190" type="holotype">holotype</typeStatus>
that an interpretation was possible.
</paragraph>
</subSubSection>
<subSubSection id="E8A06567FF86FFFFFF73F1911290308F" lastPageId="12" lastPageNumber="1191" pageId="11" pageNumber="1190" type="description">
<paragraph id="A00536ECFF86FFF8FF73F19113E9312B" blockId="11.[178,1086,1021,1360]" pageId="11" pageNumber="1190">
<materialsCitation id="10D23CB1FF86FFF8FF73F191118C30BB" collectingDate="1990-04-11" collectorName="Smetana" country="Taiwan" elevation="1100" location="Taiwan" pageId="11" pageNumber="1190" specimenCount="65">
M a t e r i a l e x a m i n e d:
<collectingCountry id="D8AD767CFF86FFF8FE38F191125C30D3" box="[505,574,1172,1192]" name="Taiwan" pageId="11" pageNumber="1190">Taiwan</collectingCountry>
:
<specimenCount id="B6BCFD65FF86FFF8FD87F19112E530D3" box="[582,647,1172,1192]" pageId="11" pageNumber="1190" type="generic">65 exs.</specimenCount>
, Ilan Hsien, Chyr Duan,
<quantity id="67429B09FF86FFF8FCA0F19113C430D3" box="[865,934,1172,1192]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="1190" unit="m" value="1100.0">
<elevation id="2B97D1DFFF86FFF8FCA0F19113C430D3" box="[865,934,1172,1192]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="1190" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="D404102CFF86FFF8FC6EF191147630D3" box="[943,1044,1172,1192]" pageId="11" pageNumber="1190" value="1990-04-11">
<collectingDate id="C440E9C4FF86FFF8FC6EF191147630D3" box="[943,1044,1172,1192]" pageId="11" pageNumber="1190" value="1990-04-11">11.IV.1990</collectingDate>
</date>
, leg.
<collectorName id="0D4F533AFF86FFF8FF1EF1A9114830BB" box="[223,298,1196,1216]" pageId="11" pageNumber="1190">Smetana</collectorName>
(cSme, cAss, cSch)
</materialsCitation>
;
<materialsCitation id="10D23CB1FF86FFF8FE3FF1A913E9312B" collectingDate="1993-11-04" collectingDateMax="2009-08-29" collectingDateMin="1993-11-04" collectionCode="NHMW" collectorName="Smetana &amp; Reitter &amp; O. Scheerpeltz &amp; SW Huitong &amp; Guangping &amp; Schillhammer &amp; Baoshan Pref. &amp; Gaoligong Shan &amp; Kambaiti &amp; Wrase" country="China" county="Ilan Hsien" elevation="1110" latitude="25.183332" location="Tianmu Shan" longLatPrecision="1247" longitude="98.333336" municipality="Lake" pageId="11" pageNumber="1190" specimenCount="7" stateProvince="Zhejiang" typeStatus="holotype">
1,
<collectingCounty id="49644E60FF86FFF8FDEDF1A912ED30BB" box="[556,655,1196,1216]" pageId="11" pageNumber="1190">Ilan Hsien</collectingCounty>
, Shen-Mi
<collectingMunicipality id="4061AC96FF86FFF8FD36F1A9134730BB" box="[759,805,1196,1216]" pageId="11" pageNumber="1190">Lake</collectingMunicipality>
,
<geoCoordinate id="C58E502BFF86FFF8FCF5F1A9131C30BB" box="[820,894,1196,1216]" degrees="24" direction="north" minutes="23" orientation="latitude" pageId="11" pageNumber="1190" precision="925" value="24.383333">24°23'N</geoCoordinate>
,
<geoCoordinate id="C58E502BFF86FFF8FC4CF1A913BD30BB" box="[909,991,1196,1216]" degrees="121" direction="east" minutes="44" orientation="longitude" pageId="11" pageNumber="1190" precision="925" value="121.73333">121°44'E</geoCoordinate>
,
<quantity id="67429B09FF86FFF8FC2FF1A9145B30BB" box="[1006,1081,1196,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="11" pageNumber="1190" unit="m" value="1110.0">
<elevation id="2B97D1DFFF86FFF8FC2FF1A9145B30BB" box="[1006,1081,1196,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="11" pageNumber="1190" unit="m" value="1110.0">1110 m</elevation>
</quantity>
,
<date id="D404102CFF86FFF8FF1EF1C1115030A3" box="[223,306,1220,1240]" pageId="11" pageNumber="1190" value="1995-05-09">
<collectingDate id="C440E9C4FF86FFF8FF1EF1C1115030A3" box="[223,306,1220,1240]" pageId="11" pageNumber="1190" value="1995-05-09">9.V.1995</collectingDate>
</date>
, leg.
<collectorName id="0D4F533AFF86FFF8FEAFF1C111D930A3" box="[366,443,1220,1240]" pageId="11" pageNumber="1190">Smetana</collectorName>
&quot; (cSme).
<collectingCountry id="D8AD767CFF86FFF8FDDEF1C1123530A3" box="[543,599,1220,1240]" name="China" pageId="11" pageNumber="1190">China</collectingCountry>
: 1,
<collectingRegion id="627EF80EFF86FFF8FD54F1C1128730A3" box="[661,741,1220,1240]" country="China" name="Zhejiang" pageId="11" pageNumber="1190">Zhejiang</collectingRegion>
,
<location id="A5656037FF86FFF8FD35F1C1131130A3" LSID="urn:lsid:plazi:treatment:281387FAFF86FFFFFF73F6901290308F:A5656037FF86FFF8FD35F1C1131130A3" box="[756,883,1220,1240]" country="China" county="Ilan Hsien" latitude="25.183332" longLatPrecision="1247" longitude="98.333336" municipality="Lake" name="Tianmu Shan" pageId="11" pageNumber="1190" stateProvince="Zhejiang">Tianmu Shan</location>
, leg.
<collectorName id="0D4F533AFF86FFF8FC6EF1C1138830A3" box="[943,1002,1220,1240]" pageId="11" pageNumber="1190">Reitter</collectorName>
[&quot;
<typeStatus id="7F01884EFF86FFF8FBC2F1C1145F30A3" box="[1027,1085,1220,1240]" pageId="11" pageNumber="1190">Typus</typeStatus>
<taxonomicName id="67BA4D6FFF86FFF8FF1EF1D91124308B" box="[223,326,1244,1264]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1190" phylum="Arthropoda" rank="genus">Lathrobium</taxonomicName>
tienmuschanensis
<collectorName id="0D4F533AFF86FFF8FE2EF1D91216308B" box="[495,628,1244,1264]" pageId="11" pageNumber="1190">O. Scheerpeltz</collectorName>
&quot; (
<collectionCode id="C6ABAE29FF86FFF8FD48F1D912B4308B" box="[649,726,1244,1264]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="1190" type="Museum">NHMW</collectionCode>
); 1,
<collectingCountry id="D8AD767CFF86FFF8FCD5F1D91328308B" box="[788,842,1244,1264]" name="China" pageId="11" pageNumber="1190">China</collectingCountry>
, SW-Hunan,
<collectorName id="0D4F533AFF86FFF8FC07F1D91458308B" box="[966,1082,1244,1264]" pageId="11" pageNumber="1190">SW Huitong</collectorName>
,
<date id="D404102CFF86FFF8FF1EF1F1115B3173" box="[223,313,1268,1288]" pageId="11" pageNumber="1190" value="1993-11-07">
<collectingDate id="C440E9C4FF86FFF8FF1EF1F1115B3173" box="[223,313,1268,1288]" pageId="11" pageNumber="1190" value="1993-11-07">7.XI.1993</collectingDate>
</date>
,
<collectorName id="0D4F533AFF86FFF8FE85F1F111C73173" box="[324,421,1268,1288]" pageId="11" pageNumber="1190">Guangping</collectorName>
env.,
<quantity id="67429B09FF86FFF8FE1FF1F1127B3173" box="[478,537,1268,1288]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="1190" unit="m" value="400.0">
<elevation id="2B97D1DFFF86FFF8FE1FF1F1127B3173" box="[478,537,1268,1288]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="1190" unit="m" value="400.0">400 m</elevation>
</quantity>
, leg.
<collectorName id="0D4F533AFF86FFF8FD8AF1F112A73173" box="[587,709,1268,1288]" pageId="11" pageNumber="1190">Schillhammer</collectorName>
(
<collectionCode id="C6ABAE29FF86FFF8FD12F1F113423173" box="[723,800,1268,1288]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="1190" type="Museum">NHMW</collectionCode>
);
<specimenCount id="B6BCFD65FF86FFF8FCF0F1F113053173" box="[817,871,1268,1288]" pageId="11" pageNumber="1190" type="generic">7 exs.</specimenCount>
[2 teneral], SW-Hunan,
<collectorName id="0D4F533AFF86FFF8FF1EF0091136315B" box="[223,340,1292,1312]" pageId="11" pageNumber="1190">SW Huitong</collectorName>
,
<collectorName id="0D4F533AFF86FFF8FEA0F00911A0315B" box="[353,450,1292,1312]" pageId="11" pageNumber="1190">Guangping</collectorName>
env.,
<quantity id="67429B09FF86FFF8FE3EF009125F315B" box="[511,573,1292,1312]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="1190" unit="m" value="350.0">
<elevation id="2B97D1DFFF86FFF8FE3EF009125F315B" box="[511,573,1292,1312]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="1190" unit="m" value="350.0">350 m</elevation>
</quantity>
,
<date id="D404102CFF86FFF8FD8BF00912C7315B" box="[586,677,1292,1312]" pageId="11" pageNumber="1190" value="1993-11-04">
<collectingDate id="C440E9C4FF86FFF8FD8BF00912C7315B" box="[586,677,1292,1312]" pageId="11" pageNumber="1190" value="1993-11-04">4.XI.1993</collectingDate>
</date>
, leg.
<collectorName id="0D4F533AFF86FFF8FD1AF0091337315B" box="[731,853,1292,1312]" pageId="11" pageNumber="1190">Schillhammer</collectorName>
(
<collectionCode id="C6ABAE29FF86FFF8FCA4F00913D0315B" box="[869,946,1292,1312]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="1190" type="Museum">NHMW</collectionCode>
); 1,
<collectingRegion id="627EF80EFF86FFF8FC33F0091458315B" box="[1010,1082,1292,1312]" country="China" name="Yunnan" pageId="11" pageNumber="1190">Yunnan</collectingRegion>
,
<collectorName id="0D4F533AFF86FFF8FF1EF02111003143" box="[223,354,1316,1336]" pageId="11" pageNumber="1190">Baoshan Pref.</collectorName>
,
<collectorName id="0D4F533AFF86FFF8FEAEF02112663143" box="[367,516,1316,1336]" pageId="11" pageNumber="1190">Gaoligong Shan</collectorName>
, road to
<collectorName id="0D4F533AFF86FFF8FDA0F02112D03143" box="[609,690,1316,1336]" pageId="11" pageNumber="1190">Kambaiti</collectorName>
pass,
<locationDeviation id="B1B72FCEFF86FFF8FD32F02113833143" box="[755,993,1316,1336]" location="Tengchong" pageId="11" pageNumber="1190">
<quantity id="67429B09FF86FFF8FD32F02113203143" box="[755,834,1316,1336]" metricMagnitude="4" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="1190" unit="km" value="22.5">22.5 km</quantity>
NW Tengchong
</locationDeviation>
,
<geoCoordinate id="C58E502BFF86FFF8FC31F02114583143" box="[1008,1082,1316,1336]" degrees="25" direction="north" minutes="11" orientation="latitude" pageId="11" pageNumber="1190" precision="925" value="25.183332">25°11'N</geoCoordinate>
,
<geoCoordinate id="C58E502BFF86FFF8FF1EF0391145312B" box="[223,295,1340,1360]" degrees="98" direction="east" minutes="20" orientation="longitude" pageId="11" pageNumber="1190" precision="925" value="98.333336">98°20'E</geoCoordinate>
,
<quantity id="67429B09FF86FFF8FEEEF0391110312B" box="[303,370,1340,1360]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="11" pageNumber="1190" unit="m" value="1930.0">
<elevation id="2B97D1DFFF86FFF8FEEEF0391110312B" box="[303,370,1340,1360]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="11" pageNumber="1190" unit="m" value="1930.0">1930 m</elevation>
</quantity>
, creek bank with vegetation,
<date id="D404102CFF86FFF8FDAEF0391280312B" box="[623,738,1340,1360]" pageId="11" pageNumber="1190" value="2009-08-29">
<collectingDate id="C440E9C4FF86FFF8FDAEF0391280312B" box="[623,738,1340,1360]" pageId="11" pageNumber="1190" value="2009-08-29">29.VIII.2009</collectingDate>
</date>
, leg.
<collectorName id="0D4F533AFF86FFF8FCCEF0391325312B" box="[783,839,1340,1360]" pageId="11" pageNumber="1190">Wrase</collectorName>
(cAss).
</materialsCitation>
</paragraph>
<paragraph id="A00536ECFF86FFF8FF73F05A11AD319F" blockId="11.[178,1086,1374,1508]" pageId="11" pageNumber="1190">
D i a g n o s i s: Body length
<quantity id="67429B09FF86FFF8FE35F05B120E310F" box="[500,620,1374,1396]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.8" metricValueMin="5.8" pageId="11" pageNumber="1190" unit="mm" value="6.3" valueMax="6.8" valueMin="5.8">5.8-6.8mm</quantity>
;lengthofforebody
<quantity id="67429B09FF86FFF8FC85F05B13DC310F" box="[836,958,1374,1396]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="11" pageNumber="1190" unit="mm" value="3.45" valueMax="3.7" valueMin="3.2">3.2-3.7 mm</quantity>
. Coloration: body blackish, postero-external angles of elytra with yellowish to reddish-yellow spot of variable size; middle and hind legs yellowish; forelegs in mature specimens almost always somewhat darker; antennae dark-brown, with the basal 1-2 and the apical 1-3 antennomeres paler reddish.
</paragraph>
<paragraph id="A00536ECFF86FFF8FF73F0F6131B323F" blockId="11.[178,1085,1523,1604]" pageId="11" pageNumber="1190">
Elytra polymorphic, approximately as long as pronotum or nearly so, rarely shorter, in specimens from mainland
<collectingCountry id="D8AD767CFF86FFF8FE06F3151264325D" box="[455,518,1552,1574]" name="China" pageId="11" pageNumber="1190">China</collectingCountry>
, and approximately 0.9 times as long as pronotum in material from
<collectingCountry id="D8AD767CFF86FFF8FE83F32B11EC323F" box="[322,398,1582,1604]" name="Taiwan" pageId="11" pageNumber="1190">Taiwan</collectingCountry>
(
<figureCitation id="38812A69FF86FFF8FE5DF32B1185323F" box="[412,487,1582,1604]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="11" pageNumber="1190">Fig. 39</figureCitation>
); punctation dense and usually defined.
</paragraph>
<paragraph id="A00536ECFF81FFFFFF06F5DE13A635DB" blockId="12.[178,1085,218,416]" pageId="12" pageNumber="1191">
: posterior margin of tergite VIII convexly produced in the middle (
<figureCitation id="38812A69FF81FFFFFC40F5DF13AD348B" box="[897,975,218,240]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Fig. 40</figureCitation>
); posterior margin of sternite VII weakly concave in the middle; sternite VIII without distinct clusters of black setae, posterior margin with pronounced V-shaped incision (
<figureCitation id="38812A69FF81FFFFFC7AF41014533550" box="[955,1073,277,299]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Figs 41-42</figureCitation>
); aedeagus (
<figureCitation id="38812A69FF81FFFFFEDEF43711F13533" box="[287,403,306,328]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Figs 43-45</figureCitation>
,
<figureCitation id="38812A69FF81FFFFFE60F43711BF3533" box="[417,477,306,328]" captionStart="Figs 48-51" captionStartId="11.[178,218,810,832]" captionTargetBox="[182,1073,212,796]" captionTargetId="figure-365@11.[178,1085,212,800]" captionTargetPageId="11" captionText="Figs 48-51: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Yunnan (48-49), Taiwan (50) and from Hunan (51): (48) aedeagus in lateral view; (49-51) aedeagus in dorsal view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326119" httpUri="https://zenodo.org/record/5326119/files/figure.png" pageId="12" pageNumber="1191">48-51</figureCitation>
) 0.9-1.0 mm long; ventral process usually weakly angled approximately in the midde (lateral view), moderately broad basally and very slender apically in ventral view, apex of ventral process distinctly hooked (lateral view); dorsal plate apically convex (dorsal view); internal sac with large sclerotized spines.
</paragraph>
<paragraph id="A00536ECFF81FFFFFF06F4B5119D367B" blockId="12.[178,1085,431,512]" pageId="12" pageNumber="1191">
: posterior margin of tergite VIII produced and distinctly angled in the middle (
<figureCitation id="38812A69FF81FFFFFBD5F4AA10AD3598" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Fig. 46</figureCitation>
); sternite VIII distinctly longer than tergite VIII, posterior margin produced, in the middle almost truncate (
<figureCitation id="38812A69FF81FFFFFE66F4EF1196367B" box="[423,500,490,512]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Fig. 47</figureCitation>
).
</paragraph>
<paragraph id="A00536ECFF81FFFFFF73F71513A437FE" blockId="12.[178,1085,527,901]" pageId="12" pageNumber="1191">
I n t r a s p e c i f i c v a r i a t i o n: The length of the elytra, the size and colour of the pale posterior spot on the elytra, and even the shape of the ventral process of the aedeagus are subject to rather pronounced variation. The elytra are distinctly shorter in specimens from
<collectingCountry id="D8AD767CFF81FFFFFE9CF76211CB3606" box="[349,425,615,637]" name="Taiwan" pageId="12" pageNumber="1191">Taiwan</collectingCountry>
and from Yunnan than in material from the remaining Chinese localities. The elytral spots are darker (pale reddish) and much larger (occupying approximately one third of the elytra surface) in the specimen from Yunnan, whereas in the material seen from other localities they are pale yellow and confined to the extreme postero-lateral angles. The ventral process of the aedeagus is subapically slightly more slender in males from
<collectingCountry id="D8AD767CFF81FFFFFE61F7FC118E3774" box="[416,492,761,783]" name="Taiwan" pageId="12" pageNumber="1191">Taiwan</collectingCountry>
(
<figureCitation id="38812A69FF81FFFFFE3DF7FC12283774" box="[508,586,761,783]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Fig. 44</figureCitation>
) and less distinctly angled in the middle in the male from Yunnan (
<figureCitation id="38812A69FF81FFFFFE4FF61211823756" box="[398,480,791,813]" captionStart="Figs 48-51" captionStartId="11.[178,218,810,832]" captionTargetBox="[182,1073,212,796]" captionTargetId="figure-365@11.[178,1085,212,800]" captionTargetPageId="11" captionText="Figs 48-51: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Yunnan (48-49), Taiwan (50) and from Hunan (51): (48) aedeagus in lateral view; (49-51) aedeagus in dorsal view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326119" httpUri="https://zenodo.org/record/5326119/files/figure.png" pageId="12" pageNumber="1191">Fig. 48</figureCitation>
). However, since no additional distinguishing characters were found and in view of the similar general morphology of the aedeagus, particularly the internal structures in lateral (
<figureCitation id="38812A69FF81FFFFFDCCF65412E3371C" box="[525,641,849,871]" captionStart="Figs 39-47" captionStartId="10.[178,218,1431,1453]" captionTargetBox="[184,1076,212,1413]" captionTargetId="figure-79@10.[178,1085,212,1422]" captionTargetPageId="10" captionText="Figs 39-47: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Taiwan (39-41, 44-47) and from Hunan (42-43): (39) forebody; (40) male tergite VIII; (41-42) male sternite VIII; (43-44) aedeagus in lateral view; (45) aedeagus in ventral view; (46) female tergite VIII; (47) female sternite VIII. Scale bars: 39: 1.0 mm; 40-47: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326117" httpUri="https://zenodo.org/record/5326117/files/figure.png" pageId="12" pageNumber="1191">Figs 43-44</figureCitation>
,
<figureCitation id="38812A69FF81FFFFFD4EF65412CB371C" box="[655,681,849,871]" captionStart="Figs 48-51" captionStartId="11.[178,218,810,832]" captionTargetBox="[182,1073,212,796]" captionTargetId="figure-365@11.[178,1085,212,800]" captionTargetPageId="11" captionText="Figs 48-51: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Yunnan (48-49), Taiwan (50) and from Hunan (51): (48) aedeagus in lateral view; (49-51) aedeagus in dorsal view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326119" httpUri="https://zenodo.org/record/5326119/files/figure.png" pageId="12" pageNumber="1191">48</figureCitation>
) and in dorsal view (
<figureCitation id="38812A69FF81FFFFFC52F654146A371C" box="[915,1032,849,871]" captionStart="Figs 48-51" captionStartId="11.[178,218,810,832]" captionTargetBox="[182,1073,212,796]" captionTargetId="figure-365@11.[178,1085,212,800]" captionTargetPageId="11" captionText="Figs 48-51: Tetartopeus bimaculatus (LI, TANG &amp; ZHU) from Yunnan (48-49), Taiwan (50) and from Hunan (51): (48) aedeagus in lateral view; (49-51) aedeagus in dorsal view. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5326119" httpUri="https://zenodo.org/record/5326119/files/figure.png" pageId="12" pageNumber="1191">Figs 49-51</figureCitation>
), the observed differences are attributed to intra- rather than interspecific variation.
</paragraph>
<paragraph id="A00536ECFF81FFFFFF73F69113D33022" blockId="12.[178,1086,916,1113]" pageId="12" pageNumber="1191">
C o m p a r a t i v e n o t e s: Both in external and sexual characters,
<taxonomicName id="67BA4D6FFF81FFFFFC67F690145C37D2" authorityName=": ASSING" authorityYear="2010" box="[934,1086,916,938]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1191" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="92CEEAFEFF81FFFFFC67F690145C37D2" box="[934,1086,916,938]" italics="true" pageId="12" pageNumber="1191">T. bimaculatus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="67BA4D6FFF81FFFFFEB0F6B711BF37BD" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[369,477,945,967]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1191" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="92CEEAFEFF81FFFFFEB0F6B711BF37BD" box="[369,477,945,967]" italics="true" pageId="12" pageNumber="1191">T. pallipes</emphasis>
</taxonomicName>
, but distinguished by the usually somewhat darker coloration of the forelegs (in
<taxonomicName id="67BA4D6FFF81FFFFFE61F6CA126E379F" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[416,524,975,996]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1191" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="92CEEAFEFF81FFFFFE61F6CA126E379F" box="[416,524,975,996]" italics="true" pageId="12" pageNumber="1191">T. pallipes</emphasis>
</taxonomicName>
of similar coloration as the yellowish middle and hind legs), the morphology of the aedeagus (larger; ventral process stouter, bent in the middle, and apically distinctly hooked), and by the slightly different shape of the posterior margin of the female sternite VIII (
<taxonomicName id="67BA4D6FFF81FFFFFE35F12212023040" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[500,608,1062,1084]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1191" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="92CEEAFEFF81FFFFFE35F12212023040" box="[500,608,1062,1084]" italics="true" pageId="12" pageNumber="1191">T. pallipes</emphasis>
</taxonomicName>
: more convex in the middle). Specimens from
<collectingCountry id="D8AD767CFF81FFFFFF73F146109C3022" box="[178,254,1091,1113]" name="Taiwan" pageId="12" pageNumber="1191">Taiwan</collectingCountry>
are additionally distinguished from
<taxonomicName id="67BA4D6FFF81FFFFFDACF14112BA3022" baseAuthorityName="Sharp" baseAuthorityYear="1889" box="[621,728,1092,1113]" class="Insecta" family="Staphylinidae" genus="Tetartopeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1191" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="92CEEAFEFF81FFFFFDACF14112BA3022" box="[621,728,1092,1113]" italics="true" pageId="12" pageNumber="1191">T. pallipes</emphasis>
</taxonomicName>
by the shorter elytra.
</paragraph>
<paragraph id="A00536ECFF81FFFFFF73F16C1290308F" blockId="12.[178,1085,1129,1268]" pageId="12" pageNumber="1191">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The species is currently known from mainland
<collectingCountry id="D8AD767CFF81FFFFFE69F183118630E7" box="[424,484,1158,1180]" name="China" pageId="12" pageNumber="1191">China</collectingCountry>
(
<collectingRegion id="627EF80EFF81FFFFFE36F183123030E7" box="[503,594,1158,1180]" country="China" name="Guizhou" pageId="12" pageNumber="1191">Guizhou</collectingRegion>
,
<collectingRegion id="627EF80EFF81FFFFFDA3F18312DD30E7" box="[610,703,1158,1180]" country="China" name="Zhejiang" pageId="12" pageNumber="1191">Zhejiang</collectingRegion>
,
<collectingRegion id="627EF80EFF81FFFFFD0FF182137630E7" box="[718,788,1159,1180]" country="China" name="Hunan" pageId="12" pageNumber="1191">Hunan</collectingRegion>
,
<collectingRegion id="627EF80EFF81FFFFFCE2F182131730E7" box="[803,885,1159,1180]" country="China" name="Yunnan" pageId="12" pageNumber="1191">Yunnan</collectingRegion>
) and
<collectingCountry id="D8AD767CFF81FFFFFC76F183146430E7" box="[951,1030,1158,1180]" name="Taiwan" pageId="12" pageNumber="1191">Taiwan</collectingCountry>
. The material from
<collectingCountry id="D8AD767CFF81FFFFFE83F1A611EC30C2" box="[322,398,1187,1209]" name="Taiwan" pageId="12" pageNumber="1191">Taiwan</collectingCountry>
was collected at the edge of a pond by pushing
<emphasis id="92CEEAFEFF81FFFFFCB9F1A113D430C2" box="[888,950,1188,1209]" italics="true" pageId="12" pageNumber="1191">Carex</emphasis>
tufts into the water (SMETANA pers. comm.). The male from Yunnan was found on a vegetated stream bank. Two specimens collected in November are teneral.
</paragraph>
</subSubSection>
</treatment>
</document>
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