219 lines
18 KiB
XML
219 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.363.6482" ID-GBIF-Dataset="36e5551d-3881-45a2-810a-386e517c86fc" ID-PMC="PMC3867181" ID-Pensoft-Pub="1313-2970-363-1" ID-PubMed="24363601" ID-ZBK="780695A9CB2E4FECA0560BA6F456185E" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-363-1" ModsDocOrigin="ZooKeys 363" ModsDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" checkinTime="1451246560986" checkinUser="pensoft" docAuthor="Mathis, Wayne N. & Zatwarnicki, Tadeusz" docDate="2013" docId="268EB4C55DB3E66557CFAEF72096AF3C" docLanguage="en" docName="ZooKeys 363: 1-161" docOrigin="ZooKeys 363" docSource="http://dx.doi.org/10.3897/zookeys.363.6482" docTitle="Hydrochasma aquia Mathis & Zatwarnicki" docType="treatment" docVersion="3" lastPageNumber="78" masterDocId="FFF2FFCA0333765855689824FFCDB839" masterDocTitle="A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="76" updateTime="1668157177207" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>363</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>161</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.363.6482</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.363.6482</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-363-1</mods:identifier>
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<mods:identifier type="ZBK">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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<mods:identifier type="ZooBank">780695A9CB2E4FECA0560BA6F456185E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050163" LSID="urn:lsid:plazi:treatment:268EB4C55DB3E66557CFAEF72096AF3C" httpUri="http://treatment.plazi.org/id/268EB4C55DB3E66557CFAEF72096AF3C" lastPageId="77" lastPageNumber="78" pageId="75" pageNumber="76">
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<subSubSection pageId="75" pageNumber="76" type="nomenclature">
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<paragraph pageId="75" pageNumber="76">
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26.
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<taxonomicName LSID="http://species-id.net/wiki/Hydrochasma_aquia" authority="Mathis & Zatwarnicki" class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma aquia" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="aquia">Hydrochasma aquia Mathis & Zatwarnicki</taxonomicName>
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Figs 144-148
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</paragraph>
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</subSubSection>
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<subSubSection pageId="75" pageNumber="76" type="reference_group">
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<paragraph pageId="75" pageNumber="76">
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma aquia" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="aquia">Hydrochasma aquia</taxonomicName>
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Mathis & Zatwarnicki 2010: 110 [United States. Virginia. Stafford: Aquia Harbour, Lions Park (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.45">38°27'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.388336">77°23.3'W</geoCoordinate>
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); HT ♂, USNM].
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="diagnosis">
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<paragraph pageId="75" pageNumber="76">Diagnosis.</paragraph>
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<paragraph lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
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This species is distinguished from congeners by the following combination of characters: Small shore flies, body length 1.50-1.95 mm.
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<pageBreakToken pageId="76" pageNumber="77" start="start">Head</pageBreakToken>
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: Antenna mostly dark gray; parafacial silvery white, concolorous with facial coloration. Gena moderately high, height usually slightly higher than length of basal flagellomere; gena-to-eye ratio 0.28-0.30. Thorax: Wing hyaline; costal vein ratio 0.65-0.72; M vein ratio 0.56-0.60. Hindtibia with only pale, yellowish setae apicoventrally. Abdomen: Tergites lacking wedge-like marking laterally but tergites 2-4 with narrow to moderately wide medial area extensively dark slate gray to black; tergite 5 light gray to silvery gray with undifferentiated posterior margin, uniformly colored; medial coloration on tergites 1-4 narrow, sometimes only a stripe, slightly darker than color of lateral margins. Male terminalia (Fig. 144-147): Combined structures generally elongate, in posterior view (Fig. 144) height more than twice width, dorsal portion semi-rectangular, ventral portion narrowed; epandrium with dorsal arch above cerci attenuate, not connected, dorsal half more or less roundly rectangular, slightly higher than wide, thereafter ventrally tapered to narrow, almost parallel-sided processes, apex with taper more angulate, dorsal 2/3 bearing setulae, in lateral view (Fig. 145) mostly parallel sided with anterior margin with a midlength shallow prominence, dorsal half shallowly curved anteriorly, apical half parallel sided, straight, apex moderately narrowly rounded; cerci moderately short, height not quite twice width (Fig. 144), narrowly attached ventrally tenuously with epandrium; aedeagus in lateral view (Fig. 147) very elongate, sclerotized portion very narrow, with length almost 10
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<normalizedToken originalValue="×">x</normalizedToken>
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or more width, anteroapical, membranous portion only slightly expanded, width not much more than lateral width of hypandrium, membranous portion with numerous cuticular spicules, in ventral view (Fig. 146) also narrow and very elongate, almost parallel sided basally, membranous apical portion becoming wider apically, apex rounded; phallapodeme in lateral view (Fig. 147) narrow, elongate, unevenly clavate toward hypandrium, keel slightly developed, apical width, including keel, slightly less than greatest aedeagal width, in ventral view (Fig. 146) with hypandrial end narrowly linear as a T, with arms of T short and curved, thereafter toward base of aedeagus bar-like, slightly expanding; gonite in lateral view (Fig. 147) very narrow, elongate, very shallowly sinuous, in ventral view (Fig. 146) narrow, bar-like with apices toward base of aedeagus shallowly curved medially; hypandrium in lateral view (Fig. 147) elongate, moderately shallow, width more or less uniform and slightly less than 1/4 length, nearly straight, in ventral view (Fig. 146) generally deeply V-shaped with posterior margin very deeply V-shaped, anterior margin moderately narrow, bluntly rounded.
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</paragraph>
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<caption pageId="76" pageNumber="77">
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<paragraph pageId="76" pageNumber="77">
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Figures 144-147.
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma aquia" order="Diptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="aquia">Hydrochasma aquia</taxonomicName>
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Mathis and Zatwarnicki (USA. Virginia. Stafford: Aquia Harbour, Lions Park) 144 epandrium and cerci, posterior view 145 same, lateral view 146 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 147 same, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="76" pageNumber="77" type="type material">
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<paragraph pageId="76" pageNumber="77">Type material.</paragraph>
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<paragraph pageId="76" pageNumber="77">
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The holotype male is labeled "USA. V[IRGINI]A. Stafford: Aquia Harbour, Lions Park (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.45">38°27'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.388336">77°23.3'W</geoCoordinate>
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), 21 Jul 2008, D. & W. N. Mathis/USNM ENT 00118100 [plastic bar code label]/HOLOTYPE ♂
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma aquia" order="Diptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="aquia">Hydrochasma aquia</taxonomicName>
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W. Mathis & T. Zatwarnicki USNM [red]." The holotype is double mounted (minuten in a block of plastic), is in excellent condition, and is deposited in the USNM. Sixteen paratypes (15♂, 1♀; USNM) bear the same label data as the holotype. Other paratypes are as follows:
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</paragraph>
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<paragraph pageId="76" pageNumber="77">
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VIRGINIA. Culpeper: Lake Pelham (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.463333">38°27.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.045">78°02.7'W</geoCoordinate>
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), 28 Apr 2006, D. and W. N. Mathis (1♀; USNM). Fairfax: Great Falls (quarry;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.985">38°59.1'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.246666">77°14.8'W</geoCoordinate>
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; 50 m), 13 Jun 2007, D. and W. N. Mathis (3♂; USNM); Turkey Run (mouth;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.965">38°57.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.15667">77°09.4'W</geoCoordinate>
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), 25 May-7 Sep 2006, 2007, 2008, D. and W. N. Mathis (5♂, 1♀; USNM). Henry: Martinsville Reservoir (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.745">36°44.7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-79.87">79°52.2'W</geoCoordinate>
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), 17 May 2005, D. and W. N. Mathis (1♀; USNM). Madison: Criglersville (1.6 km W;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.47333">38°28.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.331665">78°19.9'W</geoCoordinate>
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; 185 m; Robinson River), 1 Jul 2005, D. and W. N. Mathis (1♂, 1♀; USNM). Patrick: Meadows of Dan (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.736668">36°44.2'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.38167">80°22.9'W</geoCoordinate>
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), 18 May 2005, D. and W. N. Mathis (2♂; USNM); Woolwine (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="36.79">36°47.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-80.278336">80°16.7'W</geoCoordinate>
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; 300 m), 17 May 2005, D. and W. N. Mathis (1♂, 1♀; USNM). Spotsylvania: Rappahannock River (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.31333">38°18.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.541664">77°32.5'W</geoCoordinate>
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), 3 Jul 2007, D. and W. N. Mathis (1♂, 1♀; USNM). Stafford: Aquia Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.485">38°29.1'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.39667">77°23.8'W</geoCoordinate>
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), 6 Jun 2005, D. and W. N. Mathis (1♂; USNM); Falmouth (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.32">38°19.2'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.46833">77°28.1'W</geoCoordinate>
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; Rappahannock River; 9 m), 26 May-6 Jun 2007, D. and W. N. Mathis (2♂, 2♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="76" pageNumber="77" type="type locality">
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<paragraph pageId="76" pageNumber="77">Type locality.</paragraph>
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<paragraph pageId="76" pageNumber="77">
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United States. Virginia. Stafford: Aquia Harbour, Lions Park (
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="38.45">38°27'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.388336">77°23.3'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="other specimens examined">
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<paragraph pageId="76" pageNumber="77">Other specimens examined.</paragraph>
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<paragraph pageId="76" pageNumber="77">
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Nearctic. UNITED STATES. DELAWARE. Kent: Woodland Beach (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="39.331665">39°19.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-75.471664">75°28.3'W</geoCoordinate>
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; beach), 18 May 2006, D. and W. N. Mathis (1♂; USNM).
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</paragraph>
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<paragraph pageId="76" pageNumber="77">
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DISTRICT OF COLUMBIA. Rock Creek, Milkhouse Ford (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.965">38°57.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-77.04833">77°02.9'W</geoCoordinate>
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), 18 May 2007, D. and W. N. Mathis (1♀; USNM).
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</paragraph>
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<paragraph pageId="76" pageNumber="77">
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IOWA. Louisa: Oakville (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="42.163334">42°09.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-91.013336">91°0.8'W</geoCoordinate>
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), 10 Aug 1960, D. L. Deonier (12♂, 15♀; USNM).
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</paragraph>
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<paragraph pageId="77" pageNumber="78">
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<pageBreakToken pageId="77" pageNumber="78" start="start">WEST</pageBreakToken>
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VIRGINIA. Hardy: Mathias (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.876667">38°52.6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-78.86667">78°52'W</geoCoordinate>
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; 465 m), 13 Jul 2007, D. and W. N. Mathis (1♀; USNM); Trout Pond (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="38.956665">38°57.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-78.736664">78°44.2'W</geoCoordinate>
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; 595 m), 13 Jul 2007, D. and W. N. Mathis (1♀; USNM). Mercer: Ceres (Kee Reservoir;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.306667">37°18.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.17333">81°10.4'W</geoCoordinate>
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; 757 m), 24 Sep 2007, D. and W. N. Mathis (2♂; USNM). Summers: Hinton (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="37.696667">37°41.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-80.88333">80°53'W</geoCoordinate>
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; New River; 427 m), 26 Sep 2007, D. and W. N. Mathis (1♂; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="distribution">
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<paragraph pageId="77" pageNumber="78">Distribution</paragraph>
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<paragraph pageId="77" pageNumber="78">(Fig. 148). Nearctic: United States (Delaware, District of Columbia, Iowa, Virginia, West Virginia).</paragraph>
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<caption pageId="77" pageNumber="78">
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<paragraph pageId="77" pageNumber="78">
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Figure 148. Distribution of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma aquia" order="Diptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="aquia">Hydrochasma aquia</taxonomicName>
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Mathis and Zatwarnicki.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="etymology">
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<paragraph pageId="77" pageNumber="78">Etymology.</paragraph>
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<paragraph pageId="77" pageNumber="78">
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The species epithet, aquia, is a noun in apposition and is the name for the general area, including Aquia Creek and Aquia Harbour, where the type series of this species was collected. Aquia is the transliterated name of a Native American village (Powhatan Confederacy) located near mouth of Aquia Creek in present day Stafford County, Virginia. The name, which was originally spelled phonetically as
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<normalizedToken originalValue="“Quiyough,”">"Quiyough,"</normalizedToken>
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apparently means gulls, a common bird on the creek.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="77" pageNumber="78" type="remarks">
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<paragraph pageId="77" pageNumber="78">Remarks.</paragraph>
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<paragraph pageId="77" pageNumber="78">
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This is a widespread species that is relatively common in the Nearctic Region. Externally, this species is very similar and evidently closely related to
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma andeum" order="Diptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="andeum">Hydrochasma andeum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma avanae" order="Diptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="avanae">Hydrochasma avanae</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Ephydridae" genus="Hydrochasma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrochasma annae" order="Diptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="annae">Hydrochasma annae</taxonomicName>
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but can be distinguished from these species by having a hindtibia that bears only yellowish setulae ventrally near the apex, a tapered ventral epandrial process, and by having a more delicately developed hypandrium that is deeply U-shaped in ventral view (Fig. 146).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |