treatments-xml/data/26/22/9F/26229F1D1672DE63FF1E646F6C39ADE5.xml

<document id="C54B7274FB364D433804B021C6491B98" ID-DOI="10.11646/zootaxa.5016.4.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222684" ID-ZooBank="3AC9F0AA-B258-428E-BB2E-64842D743AD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629383827173" checkinUser="plazi" docAuthor="Li, Qi-Qi, Wang, Yan &amp; Yin, Zi-Wei" docDate="2021" docId="26229F1D1672DE63FF1E646F6C39ADE5" docLanguage="en" docName="zootaxa.5016.4.9.pdf" docOrigin="Zootaxa 5016 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ctenisomorphus yinggelingensis Li &amp; Wang &amp; Yin 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="593" masterDocId="DA1BE7651673DE66FF89606A6913A800" masterDocTitle="First species of Ctenisomorphus Raffray, 1890 from China, with comments on Largeyeus J. - K. Li, 1993 (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="596" masterPageNumber="588" pageNumber="589" updateTime="1699070735244" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8B13272C08E807DA65B66C235EFE7AEF">First species of Ctenisomorphus Raffray, 1890 from China, with comments on Largeyeus J. - K. Li, 1993 (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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<mods:namePart id="9E3EB26E73860DE4765806CA4AB5D8E7">Li, Qi-Qi</mods:namePart>
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<mods:affiliation id="F47DA42F8757048D9C7555924DDCCA24">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China. &amp; 409429668 @ qq. com; https: // orcid. org / 0000 - 0003 - 0651 - 8401</mods:affiliation>
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<mods:namePart id="001AD3F9C7F79BA6F568A47F87EC7BBF">Wang, Yan</mods:namePart>
<mods:affiliation id="97510331557DFBE783F44EC80E77C0FC">Yinggeling Branch of Hainan Tropical Rainforest National Park Administration, Yacha Town, Baisha Li Autonomous County, Hainan Province 572800, China.</mods:affiliation>
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<mods:namePart id="141192CC5A9F363FF1E8466A25A577B2">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="F0E4290EE92DD8F1457A320E157F67AE">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China.</mods:affiliation>
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<heading id="F57C99671672DE67FF1E646F6B95AC20" bold="true" box="[151,646,1029,1056]" fontSize="11" level="1" pageId="1" pageNumber="589" reason="1">
<emphasis id="9CFFF2191672DE67FF1E646F6B95AC20" bold="true" box="[151,646,1029,1056]" pageId="1" pageNumber="589">
<taxonomicName id="698B55881672DE67FF1E646F6B33AC20" ID-CoL="8XYXV" authority="Li &amp; Wang &amp; Yin, 2021" authorityName="Li &amp; Wang &amp; Yin" authorityYear="2021" box="[151,544,1029,1056]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="589" phylum="Arthropoda" rank="species" species="yinggelingensis" status="sp. nov.">
<emphasis id="9CFFF2191672DE67FF1E646F6B33AC20" bold="true" box="[151,544,1029,1056]" italics="true" pageId="1" pageNumber="589">Ctenisomorphus yinggelingensis</emphasis>
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<paragraph id="AE342E0B1672DE67FF1E6440681FAC44" blockId="1.[151,646,1029,1092]" box="[151,268,1066,1092]" pageId="1" pageNumber="589">
(
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, 
<figureCitation id="36B0328E1672DE67FF7D64406810AC44" box="[244,259,1066,1092]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="1" pageNumber="589">2</figureCitation>
)
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<paragraph id="AE342E0B1672DE67FF1E64186D5AACD4" blockId="1.[151,1437,1137,2029]" pageId="1" pageNumber="589">
<emphasis id="9CFFF2191672DE67FF1E64186856AC8B" bold="true" box="[151,325,1137,1164]" pageId="1" pageNumber="589">Type material</emphasis>
(6 exx.). 
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<emphasis id="9CFFF2191672DE67FE41641B6BC8AC8B" bold="true" box="[456,731,1137,1163]" pageId="1" pageNumber="589">
<typeStatus id="713090A91672DE67FE41641B6B78AC8B" box="[456,619,1137,1163]" pageId="1" pageNumber="589" type="holotype">HOLOTYPE</typeStatus>
: 
<collectingCountry id="D69C6E9B1672DE67FDF7641B6BC8AC8B" box="[638,731,1137,1163]" name="China" pageId="1" pageNumber="589">CHINA</collectingCountry>
</emphasis>
: 
<specimenCount id="B88DE5821672DE67FD7B64186A1AAC8C" box="[754,777,1138,1164]" pageId="1" pageNumber="589" type="male">♂</specimenCount>
</materialsCitation>
, ‘ 
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<collectingCountry id="D69C6E9B1672DE67FCA864186A7EAC8C" box="[801,877,1138,1164]" name="China" pageId="1" pageNumber="589">China</collectingCountry>
: 
<collectingRegion id="6C4FE0E91672DE67FCF764186AC1AC8C" box="[894,978,1138,1164]" country="China" name="Hainan" pageId="1" pageNumber="589">Hainan</collectingRegion>
, 
<collectingCounty id="475556871672DE67FC6C64186D7AAC8C" box="[997,1129,1138,1164]" pageId="1" pageNumber="589">Baisha Co.</collectingCounty>
, Budao Vill., 
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, 
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, 
<geoCoordinate id="CBBF48CC1672DE67FEA564FC68AEACB0" box="[300,445,1174,1200]" degrees="109" direction="east" minutes="23" orientation="longitude" pageId="1" pageNumber="589" precision="15" seconds="21" value="109.38917">109°23'21&quot;E</geoCoordinate>
, 285~ 
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<elevation id="25A6C9381672DE67FD8D64FC6B43ACB0" box="[516,592,1174,1201]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="1" pageNumber="589" unit="m" value="365.0">365 m</elevation>
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, 
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<collectingDate id="CA71F1231672DE67FDD664FC6BC4ACB0" box="[607,727,1174,1200]" pageId="1" pageNumber="589" value="2018-04-19">19.iv.2018</collectingDate>
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, mixed leaf litter, sifted, 
<collectorName id="037E4BDD1672DE67FC7464FC6D83ACB0" box="[1021,1168,1174,1200]" pageId="1" pageNumber="589">Jiang Ri-Xin</collectorName>
leg. (
<emphasis id="9CFFF2191672DE67FB5F64FD6C9FACAE" bold="true" box="[1238,1420,1175,1198]" pageId="1" pageNumber="589">布łh,NJ哥岭</emphasis>
)’ (
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)
</materialsCitation>
. 
<materialsCitation id="1EE324561672DE67FE8A64D36D56ACD4" box="[259,1093,1209,1236]" collectingDate="2018-04-19" collectionCode="SNUC" collectorName="Jiang Ri-Xin" country="China" county="Baisha Co." elevation="365" latitude="19.05" location="Yinggeling" longLatPrecision="21" longitude="109.38917" pageId="1" pageNumber="589" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
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: 
<collectingCountry id="D69C6E9B1672DE67FE3E64D36B0AACD3" box="[439,537,1209,1235]" name="China" pageId="1" pageNumber="589">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="B88DE5821672DE67FDAD64D06B43ACD4" box="[548,592,1210,1236]" pageId="1" pageNumber="589" type="male">1 ♂</specimenCount>
, 
<specimenCount id="B88DE5821672DE67FDD564D06B8FACD4" box="[604,668,1210,1236]" pageId="1" pageNumber="589" type="female">4 ♀♀</specimenCount>
, same label data as holotype (
<collectionCode id="C89AB6CE1672DE67FC6564D06D2EACD4" box="[1004,1085,1210,1236]" pageId="1" pageNumber="589">SNUC</collectionCode>
)
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.
</paragraph>
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<subSubSection id="E6917D801672DE67FF4E64B76BD7ADD0" pageId="1" pageNumber="589" type="diagnosis">
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<emphasis id="9CFFF2191672DE67FF4E64B76851ACF7" bold="true" box="[199,322,1245,1271]" pageId="1" pageNumber="589">Diagnosis.</emphasis>
<emphasis id="9CFFF2191672DE67FEC364B46897ACF8" box="[330,388,1246,1272]" italics="true" pageId="1" pageNumber="589">Male</emphasis>
. Body length approximately 2.0 mm. Maxillary palpomeres 3 and 4 with lateral extensions shorter than their respective mesal parts. Antennomers 3 slightly elongate, 4 as long as broad, antennomeres 8 2.2– 2.3 times as long as antennomeres 9. Elytra with discal striae extending from outer basal foveae to approximately apical 4/5 of elytral length. Aedeagus with median lobe almost symmetric dorso-ventrally; endophallus composed of elongate cluster of small spines; parameres short and roundly triangular, each with two apical setae. 
<emphasis id="9CFFF2191672DE67FAC865046C86AD88" box="[1345,1429,1390,1416]" italics="true" pageId="1" pageNumber="589">Female</emphasis>
. Body length approximately 
<quantity id="697383EE1672DE67FE5F65F86B72ADAC" box="[470,609,1426,1452]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="1" pageNumber="589" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1–2.2 mm</quantity>
, antennomeres 2–7 each more or less elongate, 8 as long as broad, 9–11 elongate, gradually becoming longer and broader.
</paragraph>
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<subSubSection id="E6917D801672DE63FF4E65B369D9AD9D" lastPageId="5" lastPageNumber="593" pageId="1" pageNumber="589" type="description">
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<emphasis id="9CFFF2191672DE67FF4E65B3684AADF3" bold="true" box="[199,345,1497,1523]" pageId="1" pageNumber="589">Description.</emphasis>
[a female specimen was disarticulated to check the foveal pattern] 
<emphasis id="9CFFF2191672DE67FBDE65B06D82ADF4" box="[1111,1169,1498,1524]" italics="true" pageId="1" pageNumber="589">Male</emphasis>
. Body (
<figureCitation id="36B0328E1672DE67FB6465B06C56ADF4" box="[1261,1349,1498,1524]" captionStart="FIGURE 1" captionStartId="2.[151,250,1525,1549]" captionTargetBox="[153,1432,236,1496]" captionTargetId="figure-52@2.[151,1435,234,1498]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Ctenisomorphus yinggelingensis sp. nov. A. Male. B. Female. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222686" httpUri="https://zenodo.org/record/5222686/files/figure.png" pageId="1" pageNumber="589">Fig. 1A</figureCitation>
) length 
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; color reddish-brown, tarsi and mouthparts lighter. Postantennal areas, basolateral areas of head, base of pronotum, posterior margin of elytra and tergite 1 (IV), and bases of tergite 1 and sternites 2 (IV) and 3 (V) covered with broad squamous setae, other parts of the body usually with spatulate setae.
</paragraph>
<paragraph id="AE342E0B1672DE64FF4E66006D9DA8D5" blockId="1.[151,1437,1137,2029]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="590" pageId="1" pageNumber="589">
Head (
<figureCitation id="36B0328E1672DE67FE9A6600687FAE84" box="[275,364,1642,1668]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="1" pageNumber="589">Fig. 2A</figureCitation>
) rounded at base, slightly broader than long, length from anterior margin of frontal rostrum to head base 
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, width across eyes 
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; apical portion of frons sharply sloping, almost vertical to plane of vertex, anterior margins of frons and clypeus each distinctly carinate and elevated. Eyes greatly prominent, each composed of approximately 40 moderately large facets. With C-shaped setose sulcus anterior to each eye. Venter with small gular foveae (posterior tentorial pits) in oval opening, lacking median carina. Maxillary palpus (
<figureCitation id="36B0328E1672DE67FAB766906C87AF14" box="[1342,1428,1786,1812]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="1" pageNumber="589">Fig. 2B</figureCitation>
) with palpomeres 2–4 greatly extended laterally, each with coalesced setae at apex of extension; palpomere 1 minute, palpomere 2 pedunculate for basal half, apical half broadened, palpomere 3 broad, subtriangular, jointed with palpomere 4 at mesal margin of latter segment, palpomere 4 elongate, broadest bat basal 1/3. Antennae 
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long (antennomeres 3–11 sexually dimorphic), male antennomeres as follow: 1 thick, subcylindrical, 2 much shorter and slightly narrower than 1, 3 (
<figureCitation id="36B0328E1672DE67FD8067C46B73AFC8" box="[521,608,1966,1992]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="1" pageNumber="589">Fig. 2C</figureCitation>
) slightly elongate, 4–7 of similar width, successively shorter, 4 approximately as long as broad, 5–7 each transverse, club formed by apical 4 antennomeres (
<figureCitation id="36B0328E1672DE67FBD567B86DA5AFEC" box="[1116,1206,2002,2028]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="1" pageNumber="589">Fig. 2D</figureCitation>
), 8 greatly elongate, slightly curved, approximately 2.2–2.3 times as long as 9, 9 approximately as broad as 8, 10 obliquely narrowed at base, much longer than 9, 11 longer than 10, with truncate base and round, narrowed apex.
</paragraph>
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<paragraph id="AE342E0B1671DE64FF1E659F6DC3AE0C" blockId="2.[151,1232,1525,1549]" box="[151,1232,1525,1549]" pageId="2" pageNumber="590">
<emphasis id="9CFFF2191671DE64FF1E659F6800AE0D" bold="true" box="[151,275,1525,1549]" pageId="2" pageNumber="590">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="698B55881671DE64FE03659F6BC7AE0D" authority="Li &amp; Wang &amp; Yin, 2021" authorityName="Li &amp; Wang &amp; Yin" authorityYear="2021" box="[394,724,1525,1549]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="590" phylum="Arthropoda" rank="species" species="yinggelingensis" status="sp. nov.">
<emphasis id="9CFFF2191671DE64FE03659F6BC7AE0D" box="[394,724,1525,1549]" italics="true" pageId="2" pageNumber="590">Ctenisomorphus yinggelingensis</emphasis>
</taxonomicName>
<emphasis id="9CFFF2191671DE64FD53659F6A59AE0D" bold="true" box="[730,842,1525,1549]" pageId="2" pageNumber="590">
<taxonomicNameLabel id="87CC4F621671DE64FD53659F6A38AE0D" box="[730,811,1525,1549]" pageId="2" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Male. 
<emphasis id="9CFFF2191671DE64FC18659F6ABBAE0D" bold="true" box="[913,936,1525,1549]" pageId="2" pageNumber="590">B.</emphasis>
Female. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="AE342E0B1671DE64FF4E66566AD1AEC1" blockId="2.[151,1437,1595,2018]" pageId="2" pageNumber="590">
Pronotum (
<figureCitation id="36B0328E1671DE64FECC6651688DAE55" box="[325,414,1595,1621]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="2" pageNumber="590">Fig. 2A</figureCitation>
) slightly longer than wide, length along midline 
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, maximum width 
<quantity id="697383EE1671DE64FABB66516C8BAE56" box="[1330,1432,1595,1622]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="590" unit="mm" value="0.37">0.37 mm</quantity>
, widest at basal 1/3, sides rounded; disc slightly convex, glabrous, with small median antebasal fovea covered by squamous setae, lacking lateral antebasal foveae. Prosternum with anterior part slightly longer than coxal part, with small lateral procoxal foveae; margins of coxal cavities weakly carinate.
</paragraph>
<paragraph id="AE342E0B1671DE64FF4E66A16B57AF51" blockId="2.[151,1437,1595,2018]" pageId="2" pageNumber="590">
Elytra much broader than long, length along suture 
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, maximum width 
<quantity id="697383EE1671DE64FB1166A16C11AEE6" box="[1176,1282,1739,1766]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="590" unit="mm" value="0.74">0.74 mm</quantity>
; each elytron (
<figureCitation id="36B0328E1671DE64FF29668569E0AF09" box="[160,243,1775,1801]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="2" pageNumber="590">Fig. 2E</figureCitation>
) with two large, setose basal foveae, lacking subbasal fovea; humeri slightly prominent but not denticulate; sutural stria broad and deep, thin discal stria extending from outer basal foveae to approximately apical 4/5 of elytral length; with broad posterolateral cleft.
</paragraph>
<paragraph id="AE342E0B1671DE64FF4E67316B67AFE1" blockId="2.[151,1437,1595,2018]" pageId="2" pageNumber="590">Mesoventrite short, confluent with metaventrite, with broad and setose subbasal sulcus separated by thick ridge at middle, with single setose median mesoventral fovea at posterior end of ridge, lacking lateral mesoventral foveae. Metaventrite broadly impressed along middle, with large, setose median metaventral and lateral mesocoxal foveae; posterior margin with deep split at middle.</paragraph>
<paragraph id="AE342E0B1670DE65FF4E60FD6B7EA8B1" blockId="3.[151,1436,151,393]" box="[199,621,151,177]" pageId="3" pageNumber="591">Legs elongate, lacking modifications.</paragraph>
<paragraph id="AE342E0B1670DE65FF4E60D16D67A989" blockId="3.[151,1436,151,393]" pageId="3" pageNumber="591">
Abdomen slightly narrower than elytra, widest at lateral margins of tergite 1 (IV), length of abdomen along midline 
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, maximum width 
<quantity id="697383EE1670DE65FDEF60B56BDFA8F9" box="[614,716,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="591" unit="mm" value="0.71">0.71 mm</quantity>
, with long segments IV and V and short VI–VIII. Tergite 1 (IV) slightly shorter than tergite 2 (V) along middle, tergites 1 and 2 each with broad, setose basal sulcus, tergite 3–5 (VI–VIII) successively shorter, tergites 1–4 (IV–VII) each with accompanying paratergites, 1–5 (IV–VIII) each with one pair of basolateral foveae. Sternite 2 (IV) and 3 (V) with broad, setose basal sulcus, sternite 2 (IV) with large basolateral sockets, sternites 3–5 (V–VII) each with one pair of basolateral foveae.
</paragraph>
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<paragraph id="AE342E0B1670DE65FF1E67036DF2AFC9" blockId="3.[151,1437,1897,1994]" pageId="3" pageNumber="591">
<emphasis id="9CFFF2191670DE65FF1E67036806AF81" bold="true" box="[151,277,1897,1921]" pageId="3" pageNumber="591">FIGURE 2.</emphasis>
Diagnostic characters of 
<taxonomicName id="698B55881670DE65FDAB67036A7CAF81" authority="Li &amp; Wang &amp; Yin, 2021" authorityName="Li &amp; Wang &amp; Yin" authorityYear="2021" box="[546,879,1897,1921]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="591" phylum="Arthropoda" rank="species" species="yinggelingensis" status="sp. nov.">
<emphasis id="9CFFF2191670DE65FDAB67036A7CAF81" box="[546,879,1897,1921]" italics="true" pageId="3" pageNumber="591">Ctenisomorphus yinggelingensis</emphasis>
</taxonomicName>
<emphasis id="9CFFF2191670DE65FCFE67036AD9AF81" bold="true" box="[887,970,1897,1921]" pageId="3" pageNumber="591">
<taxonomicNameLabel id="87CC4F621670DE65FCFE67036AD9AF81" box="[887,970,1897,1921]" pageId="3" pageNumber="591" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 
<emphasis id="9CFFF2191670DE65FB9367036D20AF81" bold="true" box="[1050,1075,1897,1921]" pageId="3" pageNumber="591">A.</emphasis>
Head, pronotum, and elytral base. 
<emphasis id="9CFFF2191670DE65FF1E67E769BDAFA5" bold="true" box="[151,174,1933,1957]" pageId="3" pageNumber="591">B.</emphasis>
Left maxillary palpus, in ventral view. 
<emphasis id="9CFFF2191670DE65FDCA67E76B4FAFA5" bold="true" box="[579,604,1933,1957]" pageId="3" pageNumber="591">C.</emphasis>
Antennomeres 1–4. 
<emphasis id="9CFFF2191670DE65FCB867E76A59AFA5" bold="true" box="[817,842,1933,1957]" pageId="3" pageNumber="591">D.</emphasis>
Antennomeres 7–11. 
<emphasis id="9CFFF2191670DE65FBA067E76D53AFA5" bold="true" box="[1065,1088,1933,1957]" pageId="3" pageNumber="591">E.</emphasis>
Right elytron. 
<emphasis id="9CFFF2191670DE65FB5567E76C01AFA5" bold="true" box="[1244,1298,1933,1957]" pageId="3" pageNumber="591">F–H.</emphasis>
Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H.
</paragraph>
</caption>
<caption id="FAF47E831677DE62FF1E67FD6D6BAFB0" ID-DOI="http://doi.org/10.5281/zenodo.5222690" ID-Zenodo-Dep="5222690" box="[151,1144,1943,1968]" httpUri="https://zenodo.org/record/5222690/files/figure.png" pageId="4" pageNumber="592" startId="4.[151,250,1943,1967]" targetBox="[240,1348,182,1918]" targetPageId="4">
<paragraph id="AE342E0B1677DE62FF1E67FD6D6BAFB0" blockId="4.[151,1144,1943,1968]" box="[151,1144,1943,1968]" pageId="4" pageNumber="592">
<emphasis id="9CFFF2191677DE62FF1E67FD6800AFAF" bold="true" box="[151,275,1943,1967]" pageId="4" pageNumber="592">FIGURE 3.</emphasis>
J.-K. 
<bibRefCitation id="CA1A53FA1677DE62FED867F268A6AFB0" author="Li, J. - K." box="[337,437,1944,1968]" pageId="4" pageNumber="592" pagination="151 - 167" refId="ref4215" refString="Li, J. - K. (1993) The soil beetles in Anhui Province. In: Li, J. - K. &amp; Chen, P. (Eds.), Studies on fauna and ecogeography of soil animal [sic]. Press of Northeast Normal University, Changchun, Jilin, pp. 151 - 167. [in Chinese, English subtitles]" type="book chapter" year="1993">Li (1993)</bibRefCitation>
’s original description and figures, with captions translated to English.
</paragraph>
</caption>
<paragraph id="AE342E0B1677DE63FF4E67B46D0DA8D5" blockId="4.[199,1436,2014,2040]" lastBlockId="5.[151,1437,151,790]" lastPageId="5" lastPageNumber="593" pageId="4" pageNumber="592">
Aedeagus (
<figureCitation id="36B0328E1677DE62FECD67B468AAAFF8" box="[324,441,2014,2040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1897,1921]" captionTargetBox="[152,1436,417,1869]" captionTargetId="figure-134@3.[151,1436,417,1874]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic characters of Ctenisomorphus yinggelingensis sp. nov., male. A. Head, pronotum, and elytral base. B. Left maxillary palpus, in ventral view. C. Antennomeres 1–4. D. Antennomeres 7–11. E. Right elytron. F–H. Aedeagus, in dorsal (F), lateral (G), and ventral (H) view. Scale bars: 0.3 mm in A, D, E; 0.2 mm in C; 0.1 mm in B, F–H." figureDoi="http://doi.org/10.5281/zenodo.5222688" httpUri="https://zenodo.org/record/5222688/files/figure.png" pageId="4" pageNumber="592">Fig. 2F–H</figureCitation>
) 
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long, median lobe dorsoventrally almost symmetric, apical portion greatly bent ventrally, gradually narrowing apically to form round apex; endophallus composed of numerous small spine-like structures; parameres short, roundly triangular, each with two long setae at apex.
</paragraph>
<paragraph id="AE342E0B1676DE63FF4E60B56828A989" blockId="5.[151,1437,151,790]" pageId="5" pageNumber="593">
<emphasis id="9CFFF2191676DE63FF4E60B56808A8F9" box="[199,283,223,249]" italics="true" pageId="5" pageNumber="593">Female</emphasis>
. Similar to male in external morphology; antennae shorter; each eye composed of approximately 33 facets. Measurements (as for male): body length 
<quantity id="697383EE1676DE63FD1661696A39A91D" box="[671,810,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="5" pageNumber="593" unit="mm" value="2.15" valueMax="2.2" valueMin="2.1">2.1–2.2 mm</quantity>
; length/width of head 0.35–0.36/ 
<quantity id="697383EE1676DE63FB2961686C56A91D" box="[1184,1349,258,285]" metricMagnitude="-4" metricUnit="m" metricValue="4.1499999999999995" metricValueMax="4.2" metricValueMin="4.1" pageId="5" pageNumber="593" unit="mm" value="0.415" valueMax="0.42" valueMin="0.41">0.41–0.42 mm</quantity>
, pronotum 0.40–0.41/ 
<quantity id="697383EE1676DE63FECB614D68F4A941" box="[322,487,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="5" pageNumber="593" unit="mm" value="0.375" valueMax="0.38" valueMin="0.37">0.37–0.38 mm</quantity>
, elytra 0.76–0.80/ 
<quantity id="697383EE1676DE63FD3B614D6A4BA941" box="[690,856,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="6.2" metricValueMin="6.0" pageId="5" pageNumber="593" unit="mm" value="0.61" valueMax="0.62" valueMin="0.6">0.60–0.62 mm</quantity>
; abdomen 0.65–0.71/ 
<quantity id="697383EE1676DE63FBC0614C6DE3A941" box="[1097,1264,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="5" pageNumber="593" unit="mm" value="0.775" valueMax="0.8" valueMin="0.75">0.75–0.80 mm</quantity>
; antennae (
<figureCitation id="36B0328E1676DE63FAE6614D69ABA965" captionStart="FIGURE 1" captionStartId="2.[151,250,1525,1549]" captionTargetBox="[153,1432,236,1496]" captionTargetId="figure-52@2.[151,1435,234,1498]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Ctenisomorphus yinggelingensis sp. nov. A. Male. B. Female. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222686" httpUri="https://zenodo.org/record/5222686/files/figure.png" pageId="5" pageNumber="593">Fig. 1B</figureCitation>
) lacking modifications, length 
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, antennomere 7 longer than 6 and 8, 8 shortest, 9–11 enlarged, forming clubs.
</paragraph>
<paragraph id="AE342E0B1676DE63FF4E61F86D6AAB15" blockId="5.[151,1437,151,790]" pageId="5" pageNumber="593">
<emphasis id="9CFFF2191676DE63FF4E61F868A2A9AD" bold="true" box="[199,433,402,429]" pageId="5" pageNumber="593">Comparative notes.</emphasis>
Separation of members of 
<taxonomicName id="698B55881676DE63FD6A61F96A88A9AD" authorityName="Raffray" authorityYear="1890" box="[739,923,403,429]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="9CFFF2191676DE63FD6A61F96A88A9AD" box="[739,923,403,429]" italics="true" pageId="5" pageNumber="593">Ctenisomorphus</emphasis>
</taxonomicName>
usually relies on the relative lengths (proportions) of the male antennomeres (especially antennomeres 8–11), the form of the maxillary palpi, as well as the configuration of the aedeagus. At present seven species of the genus have been known. The male of the new species is most close to that of 
<taxonomicName id="698B55881676DE63FE0961956B7DAA1A" authority="(Jeannel)" baseAuthorityName="Jeannel" box="[384,622,511,538]" class="Insecta" family="Staphylinidae" genus="Ctenistidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="9CFFF2191676DE63FE0961956880AA19" box="[384,403,511,537]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FE2961956B13AA18" box="[416,512,511,536]" italics="true" pageId="5" pageNumber="593">coomani</emphasis>
(Jeannel)
</taxonomicName>
distributed in 
<collectingCountry id="D69C6E9B1676DE63FC8661956A66AA19" box="[783,885,511,537]" name="Thailand" pageId="5" pageNumber="593">Thailand</collectingCountry>
, 
<collectingCountry id="D69C6E9B1676DE63FCF661956ACDAA19" box="[895,990,511,537]" name="Vietnam" pageId="5" pageNumber="593">Vietnam</collectingCountry>
and 
<collectingCountry id="D69C6E9B1676DE63FB9D61956D92AA19" box="[1044,1153,511,537]" name="Myanmar" pageId="5" pageNumber="593">Myanmar</collectingCountry>
by sharing a similar general appearance, and can be separated from the latter only by subtle differences: in the new species antennomeres 3 are slightly elongate, antennomeres 4 are approximately as broad as long, and antennomeres 8 are relatively shorter in relation to antennomeres 9, while both antennomeres 3 and 
<quantity id="697383EE1676DE63FCEB62016A9CAA85" box="[866,911,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="593" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="698B55881676DE63FC1162016D09AA84" box="[920,1050,619,645]" class="Insecta" family="Staphylinidae" genus="Ctenistidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="9CFFF2191676DE63FC1162016AB8AA85" box="[920,939,619,645]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FC3362016D09AA84" box="[954,1050,619,644]" italics="true" pageId="5" pageNumber="593">coomani</emphasis>
</taxonomicName>
are transverse, and antennomeres 8 are relatively longer in relation to antennomeres 9. In addition the shape of the apical portion of the aedeagus, and the position and form of the endophallus of the new species are different from the illustration given in 
<bibRefCitation id="CA1A53FA1676DE63FAC162D969F7AAF1" author="Jeannel, R." pageId="5" pageNumber="593" pagination="5 - 32" refId="ref4058" refString="Jeannel, R. (1957) Sur quelques Pselaphides du Tonkin recueillis par le Pere A. de Cooman. Revue Francaise d'Entomologie, 24 (1), 5 - 32." type="journal article" year="1957">Jeannel (1957)</bibRefCitation>
, when rotating to a same angle for comparison. 
<taxonomicName id="698B55881676DE63FC8162BD6D65AAF1" authority="Li &amp; Wang &amp; Yin, 2021" authorityName="Li &amp; Wang &amp; Yin" authorityYear="2021" box="[776,1142,727,753]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="yinggelingensis" status="sp. nov.">
<emphasis id="9CFFF2191676DE63FC8162BD6D65AAF1" box="[776,1142,727,753]" italics="true" pageId="5" pageNumber="593">Ctenisomorphus yinggelingensis</emphasis>
</taxonomicName>
<emphasis id="9CFFF2191676DE63FBF762BD6DCAAAF1" bold="true" box="[1150,1241,727,753]" pageId="5" pageNumber="593">
<taxonomicNameLabel id="87CC4F621676DE63FBF762BD6DCAAAF1" box="[1150,1241,727,753]" pageId="5" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from the other congeners except for 
<taxonomicName id="698B55881676DE63FDA662916A2AAB16" authority="(Reitter)" baseAuthorityName="Reitter" baseAuthorityYear="1900" box="[559,825,763,790]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="heterocerus">
<emphasis id="9CFFF2191676DE63FDA662916B51AB15" box="[559,578,763,789]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FDC662916BC1AB15" box="[591,722,763,789]" italics="true" pageId="5" pageNumber="593">heterocerus</emphasis>
(Reitter)
</taxonomicName>
by the following characters:
</paragraph>
<paragraph id="AE342E0B1676DE63FF1E63296D5EAB81" blockId="5.[151,1436,835,1293]" pageId="5" pageNumber="593">
― from 
<taxonomicName id="698B55881676DE63FF7763296BBAAB5D" authority="Besuchet &amp; Cuccodoro" authorityName="Besuchet &amp; Cuccodoro" authorityYear="2011" box="[254,681,835,861]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="fortipalpis">
<emphasis id="9CFFF2191676DE63FF7763296802AB5D" box="[254,273,835,861]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FEAB63296884AB5D" box="[290,407,835,861]" italics="true" pageId="5" pageNumber="593">fortipalpis</emphasis>
Besuchet &amp; Cuccodoro
</taxonomicName>
distributed in 
<collectingCountry id="D69C6E9B1676DE63FCDC63296A96AB5D" box="[853,901,835,861]" name="Iran" pageId="5" pageNumber="593">Iran</collectingCountry>
, 
<collectingCountry id="D69C6E9B1676DE63FC1A63296D9DAB5D" box="[915,1166,835,861]" name="United Arab Emirates" pageId="5" pageNumber="593">United Arab Emirates</collectingCountry>
by the relatively much shorter maxillary palpomeres 3 and 4, and different structure of the endophallus;
</paragraph>
<paragraph id="AE342E0B1676DE63FF1E63E169EDAC11" blockId="5.[151,1436,835,1293]" pageId="5" pageNumber="593">
― from 
<taxonomicName id="698B55881676DE63FF7463E16B5FABA5" authority="Peyerimhoff" authorityName="Peyerimhoff" authorityYear="1929" box="[253,588,907,933]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="garamanticus">
<emphasis id="9CFFF2191676DE63FF7463E16803ABA5" box="[253,272,907,933]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FEA963E168A9ABA4" box="[288,442,907,932]" italics="true" pageId="5" pageNumber="593">garamanticus</emphasis>
Peyerimhoff
</taxonomicName>
distributed in 
<collectingCountry id="D69C6E9B1676DE63FD7D63E16A58ABA5" box="[756,843,907,933]" name="Algeria" pageId="5" pageNumber="593">Algeria</collectingCountry>
, and the Afrotropical region by the relatively much shorter antennae and legs, shorter antennomeres 
<quantity id="697383EE1676DE63FD6663C56A0FABC9" box="[751,796,943,969]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="5" pageNumber="593" unit="in" value="9.0">9 in</quantity>
relation to antennomeres 8 (antennomeres 9 longer than half length of antennomeres 
<quantity id="697383EE1676DE63FD8063B96B26ABED" box="[521,565,979,1005]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="593" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="698B55881676DE63FDB563B96BE4ABEC" authorityName="Peyerimhoff" authorityYear="1929" box="[572,759,979,1005]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="garamanticus">
<emphasis id="9CFFF2191676DE63FDB563B96B5CABED" box="[572,591,979,1005]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FDD463B96BE4ABEC" box="[605,759,979,1004]" italics="true" pageId="5" pageNumber="593">garamanticus</emphasis>
</taxonomicName>
), and transverse antennomeres 7 (elongate in 
<taxonomicName id="698B55881676DE63FB7463B969FFAC10" authorityName="Peyerimhoff" authorityYear="1929" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="garamanticus">
<emphasis id="9CFFF2191676DE63FB7463B96C03ABED" box="[1277,1296,979,1005]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FA9763B969FFAC10" italics="true" pageId="5" pageNumber="593">garamanticus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="AE342E0B1676DE63FF1E64716B0AAC59" blockId="5.[151,1436,835,1293]" pageId="5" pageNumber="593">
― from 
<taxonomicName id="698B55881676DE63FF7F647168C6AC36" authority="(Jeannel)" baseAuthorityName="Jeannel" baseAuthorityYear="1960" box="[246,469,1051,1078]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="9CFFF2191676DE63FF7F6471681AAC35" box="[246,265,1051,1077]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FE9F64716874AC35" box="[278,359,1051,1077]" italics="true" pageId="5" pageNumber="593">indicus</emphasis>
(Jeannel)
</taxonomicName>
distributed in 
<collectingRegion id="6C4FE0E91676DE63FDFC64716A02AC35" box="[629,785,1051,1077]" country="India" name="Uttar Pradesh" pageId="5" pageNumber="593">Uttar Pradesh</collectingRegion>
, northern 
<collectingCountry id="D69C6E9B1676DE63FC0964716AA9AC35" box="[896,954,1051,1077]" name="India" pageId="5" pageNumber="593">India</collectingCountry>
by the transverse antennomeres 4–7 (about as long as wide in 
<taxonomicName id="698B55881676DE63FE1E64556B1BAC59" baseAuthorityName="Jeannel" baseAuthorityYear="1960" box="[407,520,1087,1113]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="9CFFF2191676DE63FE1E645568B9AC59" box="[407,426,1087,1113]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FE3E64556B1BAC59" box="[439,520,1087,1113]" italics="true" pageId="5" pageNumber="593">indicus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="AE342E0B1676DE63FF1E64096DE1ACA1" blockId="5.[151,1436,835,1293]" pageId="5" pageNumber="593">
― from 
<taxonomicName id="698B55881676DE63FF71640968FFAC7E" authority="(Jeannel)" baseAuthorityName="Jeannel" baseAuthorityYear="1959" box="[248,492,1123,1150]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="lindbergi">
<emphasis id="9CFFF2191676DE63FF7164096818AC7D" box="[248,267,1123,1149]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FE916409686DAC7D" box="[280,382,1123,1149]" italics="true" pageId="5" pageNumber="593">lindbergi</emphasis>
(Jeannel)
</taxonomicName>
distributed in Central Asia by the slightly elongate antennomeres 3 (transverse in 
<taxonomicName id="698B55881676DE63FA0A6409683EACA1" baseAuthorityName="Jeannel" baseAuthorityYear="1959" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="lindbergi">
<emphasis id="9CFFF2191676DE63FA0A64096C85AC7D" box="[1411,1430,1123,1149]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FF4E64ED683EACA1" box="[199,301,1159,1185]" italics="true" pageId="5" pageNumber="593">lindbergi</emphasis>
</taxonomicName>
), and antennomeres 10 being much shorter than 8 (about as long as 
<quantity id="697383EE1676DE63FBA164ED6D40ACA1" box="[1064,1107,1159,1185]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="593" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="698B55881676DE63FBD364ED6DF2ACA1" baseAuthorityName="Jeannel" baseAuthorityYear="1959" box="[1114,1249,1159,1185]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="lindbergi">
<emphasis id="9CFFF2191676DE63FBD364ED6D7EACA1" box="[1114,1133,1159,1185]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FBF264ED6DF2ACA1" box="[1147,1249,1159,1185]" italics="true" pageId="5" pageNumber="593">lindbergi</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="AE342E0B1676DE63FF1E64C16811AD0D" blockId="5.[151,1436,835,1293]" pageId="5" pageNumber="593">
― from 
<taxonomicName id="698B55881676DE63FF7464C168C2ACC6" authority="(Raffray)" baseAuthorityName="Raffray" box="[253,465,1195,1222]" class="Insecta" family="Staphylinidae" genus="Ctenistes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="major">
<emphasis id="9CFFF2191676DE63FF7464C16803ACC5" box="[253,272,1195,1221]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FEA964C16870ACC4" box="[288,355,1195,1220]" italics="true" pageId="5" pageNumber="593">major</emphasis>
(Raffray)
</taxonomicName>
distributed in the Middle East and Africa by the four-segmented antennal clubs and transverse antennomeres 5–7 (
<taxonomicName id="698B55881676DE63FD9564A56B6CACE8" box="[540,639,1231,1257]" class="Insecta" family="Staphylinidae" genus="Ctenistes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="major">
<emphasis id="9CFFF2191676DE63FD9564A56B3CACE9" box="[540,559,1231,1257]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FDB564A56B6CACE8" box="[572,639,1231,1256]" italics="true" pageId="5" pageNumber="593">major</emphasis>
</taxonomicName>
has two-segmented antennal clubs, and slightly elongate antennomeres 5–7).
</paragraph>
<paragraph id="AE342E0B1676DE63FF4E655169D9AD9D" blockId="5.[151,1436,1339,1509]" pageId="5" pageNumber="593">
Judging from the original description (
<bibRefCitation id="CA1A53FA1676DE63FDF265516A1EAD55" author="Reitter, E." box="[635,781,1339,1365]" pageId="5" pageNumber="593" pagination="49 - 59" refId="ref5105" refString="Reitter, E. (1900) Beitrag zur Coleopteren-Fauna des russischen Reiches. Deutsche Entomologische Zeitschrift, 1, 49 - 59. https: // doi. org / 10.1002 / mmnd. 48019000104" type="journal article" year="1900">Reitter 1900</bibRefCitation>
), 
<taxonomicName id="698B55881676DE63FCA865516AD5AD55" baseAuthorityName="Reitter" baseAuthorityYear="1900" box="[801,966,1339,1365]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="species" species="heterocerus">
<emphasis id="9CFFF2191676DE63FCA865516A27AD55" box="[801,820,1339,1365]" italics="true" pageId="5" pageNumber="593">C</emphasis>
. 
<emphasis id="9CFFF2191676DE63FCCA65516AD5AD55" box="[835,966,1339,1365]" italics="true" pageId="5" pageNumber="593">heterocerus</emphasis>
</taxonomicName>
of 
<collectingCountry id="D69C6E9B1676DE63FC6465516D78AD55" box="[1005,1131,1339,1365]" name="Uzbekistan" pageId="5" pageNumber="593">Uzbekistan</collectingCountry>
is deemed to be a member of 
<taxonomicName id="698B55881676DE63FF3F6535687DAD79" authorityName="Raffray" authorityYear="1890" box="[182,366,1375,1401]" class="Insecta" family="Staphylinidae" genus="Ctenisomorphus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="9CFFF2191676DE63FF3F6535687DAD79" box="[182,366,1375,1401]" italics="true" pageId="5" pageNumber="593">Ctenisomorphus</emphasis>
</taxonomicName>
, but its separation from the other members of the genus is based solely on its geographic isolation.
</paragraph>
</subSubSection>
<subSubSection id="E6917D801676DE63FF4E65CC6B01ADC1" box="[199,530,1446,1473]" pageId="5" pageNumber="593" type="distribution">
<paragraph id="AE342E0B1676DE63FF4E65CC6B01ADC1" blockId="5.[151,1436,1339,1509]" box="[199,530,1446,1473]" pageId="5" pageNumber="593">
<emphasis id="9CFFF2191676DE63FF4E65CC6872ADC0" bold="true" box="[199,353,1446,1472]" pageId="5" pageNumber="593">Distribution.</emphasis>
<collectingCountry id="D69C6E9B1676DE63FEEE65CD68BDADC1" box="[359,430,1447,1473]" name="China" pageId="5" pageNumber="593">China</collectingCountry>
: 
<collectingRegion id="6C4FE0E91676DE63FE3065CD6B1DADC1" box="[441,526,1447,1473]" country="China" name="Hainan" pageId="5" pageNumber="593">Hainan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="E6917D801676DE63FF4E65A16C39ADE5" box="[199,1322,1483,1509]" pageId="5" pageNumber="593" type="etymology">
<paragraph id="AE342E0B1676DE63FF4E65A16C39ADE5" blockId="5.[151,1436,1339,1509]" box="[199,1322,1483,1509]" pageId="5" pageNumber="593">
<emphasis id="9CFFF2191676DE63FF4E65A1685DADE5" bold="true" box="[199,334,1483,1509]" pageId="5" pageNumber="593">Etymology.</emphasis>
The species is named after Yinggeling Nature Reserve, 
<typeStatus id="713090A91676DE63FC4B65A16AE1ADE5" box="[962,1010,1483,1509]" pageId="5" pageNumber="593">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>