198 lines
32 KiB
XML
198 lines
32 KiB
XML
<document id="39A1A7E2B1C4806D8B4BE4801E576824" ID-DOI="10.5281/zenodo.183391" ID-GBIF-Dataset="46dd9791-f59e-4019-b1cc-30f1ddf8629f" ID-ISSN="1175-5326" ID-Zenodo-Dep="183391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459905098680" checkinUser="plazi" docAuthor="Guinot, Danièle" docDate="2008" docId="20528B6EFFD2FFA9D8E5F946FE31FADE" docLanguage="en" docName="zt01850p026.pdf" docOrigin="Zootaxa 1850" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Dynomeninae Ortmann 1892" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="DC6BF316FFD5FFA0D872FFE9FFCFFFE4" masterDocTitle="A re-evaluation of the Dynomenidae Ortmann, 1892 (Crustacea, Decapoda, Brachyura, Podotremata), with the recognition of four subfamilies" masterLastPageNumber="26" masterPageNumber="1" pageNumber="8" updateTime="1698228993237" updateUser="plazi">
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<mods:title id="C7927C7EBC3D5628D5027DFB3A0E8BF1">A re-evaluation of the Dynomenidae Ortmann, 1892 (Crustacea, Decapoda, Brachyura, Podotremata), with the recognition of four subfamilies</mods:title>
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<mods:namePart id="CA01BD72960BE5600F0A893C9BBA103B">Guinot, Danièle</mods:namePart>
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<mods:title id="606C2CFB3B58A7479C472E7C1D74D4D1">Zootaxa</mods:title>
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<mods:date id="0DC44EBF377775FA9DA9F5FD35C57CAB">2008</mods:date>
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<mods:number id="3A90747C249A282EBF02C5572FAADBB3">1850</mods:number>
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<treatment id="20528B6EFFD2FFA9D8E5F946FE31FADE" ID-DOI="http://doi.org/10.5281/zenodo.6234453" ID-GBIF-Taxon="119373363" ID-Zenodo-Dep="6234453" LSID="urn:lsid:plazi:treatment:20528B6EFFD2FFA9D8E5F946FE31FADE" httpUri="http://treatment.plazi.org/id/20528B6EFFD2FFA9D8E5F946FE31FADE" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
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<subSubSection id="E0E169F3FFD2FFA7D8E5F946FD82F908" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="A8443A78FFD2FFA7D8E5F946FDC6F92D" blockId="7.[151,589,1711,1772]" box="[151,521,1711,1737]" pageId="7" pageNumber="8">
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<heading id="F30C8D14FFD2FFA7D8E5F946FDC6F92D" bold="true" box="[151,521,1711,1737]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<taxonomicName id="6FFB41FBFFD2FFA7D8E5F946FDC6F92D" ID-CoL="847F5" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[151,521,1711,1737]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Dynomeninae">
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<emphasis id="9A8FE66AFFD2FFA7D8E5F946FDC6F92D" bold="true" box="[151,521,1711,1737]" pageId="7" pageNumber="8">
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||
Dynomeninae
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<bibRefCitation id="CC6A4789FFD2FFA7D939F946FDC6F92D" author="Ortmann" box="[331,521,1711,1737]" pageId="7" pageNumber="26" refString="Ortmann, A. (1892) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen, V. Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher (Systematik), 6, 532 - 588." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="A8443A78FFD2FFA7D8E5F93BFD82F908" blockId="7.[151,589,1711,1772]" box="[151,589,1746,1772]" pageId="7" pageNumber="8">
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(
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<figureCitation id="30C026FDFFD2FFA7D8EDF93BFEC5F908" box="[159,266,1746,1772]" captionStart="FIGURE 1" captionStartId="4.[151,255,1602,1626]" captionTargetBox="[283,1303,425,1571]" captionTargetId="figure@4.[283,1304,425,1574]" captionTargetPageId="4" captionText="FIGURE 1. Representative species of the four subfamilies of Dynomenidae, overall view. A, Acanthodromiinae n. subfam.: Acanthodromia margarita (Alcock, 1899), Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay & Ng 2005: 18); B, Paradynomeninae n. subfam.: Paradynomene tuberculata Sakai, 1963, Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay & Ng 2004: 4); C, Metadynomeninae n. subfam.: Metadynomene tanensis (Yokoya, 1933) with its undulating tomentum, Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay & Ng 2005: 25); D, Dynomeninae Ortmann, 1892: Hirsutodynomene vespertilio McLay & Ng, 2005, Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay & Ng 2005: 21). E, Paradynomene tuberculata Sakai, 1963, New Caledonia, frontal view showing the characteristic “ face ” (ZRC). (All photographs courtesy of P. K. L. Ng)." httpUri="https://zenodo.org/record/183392/files/figure.png" pageId="7" pageNumber="8">Figures 1</figureCitation>
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D, 3C, D, 4A–C, E, 5A–H)
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</paragraph>
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</subSubSection>
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<subSubSection id="E0E169F3FFD2FFA8D8E5F8CBFE7AFF3E" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="A8443A78FFD2FFA7D8E5F8CBFBF8F880" blockId="7.[151,1436,1826,2012]" pageId="7" pageNumber="8">
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<emphasis id="9A8FE66AFFD2FFA7D8E5F8CBFEECF8D8" bold="true" box="[151,291,1826,1852]" pageId="7" pageNumber="8">
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<typeStatus id="774084DAFFD2FFA7D8E5F8CBFF1BF8D8" box="[151,212,1826,1852]" pageId="7" pageNumber="8">Type</typeStatus>
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genus
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</emphasis>
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.
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<taxonomicName id="6FFB41FBFFD2FFA7D941F8CAFDB1F8D8" authority="Desmarest, 1823" authorityName="Desmarest" authorityYear="1823" box="[307,638,1826,1852]" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="9A8FE66AFFD2FFA7D941F8CAFE7FF8D8" box="[307,432,1827,1852]" italics="true" pageId="7" pageNumber="8">Dynomene</emphasis>
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Desmarest, 1823
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</taxonomicName>
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(
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<taxonomicName id="6FFB41FBFFD2FFA7DAFDF8CAFC32F8D8" authority="Latreille, 1812" authorityName="Latreille" authorityYear="1812" box="[655,1021,1826,1852]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hispida">
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<emphasis id="9A8FE66AFFD2FFA7DAFDF8CAFC8BF8D8" box="[655,836,1827,1852]" italics="true" pageId="7" pageNumber="8">Cancer hispida</emphasis>
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<bibRefCitation id="CC6A4789FFD2FFA7DB3EF8CBFC32F8D8" author="Latreille" box="[844,1021,1826,1852]" pageId="7" pageNumber="25" refString="Latreille, P. A. (1812) Crustaces et Insectes. In: Milbert, J., Voyage pittoresque a l'Ile-de-France, au Cap de Bonne- Esperance et a l'Ile de Teneriffe. Vol. 2. Crustaces. A. Nepveu, Paris, 270 - 280." type="book chapter" year="1812">Latreille, 1812</bibRefCitation>
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</taxonomicName>
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:
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<typeStatus id="774084DAFFD2FFA7DC7BF8CBFBF4F8D8" box="[1033,1083,1826,1852]" pageId="7" pageNumber="8">type</typeStatus>
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species by monotypy by subsequent designation of H.
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<bibRefCitation id="CC6A4789FFD2FFA7D9BBF8A3FD04F880" author="Milne" box="[457,715,1866,1892]" pageId="7" pageNumber="25" refString="Milne Edwards, H. (1837) Histoire naturelle des Crustaces comprenant l'anatomie, la physiologie et la classification de ces animaux. Vol. 2. Librairie Encyclopedique de Roret, Paris, 531 pp." type="book" year="1837">Milne Edwards, 1837</bibRefCitation>
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: 180;
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<bibRefCitation id="CC6A4789FFD2FFA7DB63F8A3FC55F880" author="ICZN" box="[785,922,1866,1892]" pageId="7" pageNumber="25" refString="ICZN (1999) International Code of Zoological Nomenclature. International Commission of Zoological Nomenclature. Fourth Edition. Adopted by the XXI General Assembly of the International Union of Biological Sciences. International Trust for Zoological Nomenclature, in association with the British Museum (Natural History), London, 338 pp." type="book" year="1999">
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<collectionCode id="CEEAA2BDFFD2FFA7DB63F8A3FC9AF880" box="[785,853,1866,1892]" pageId="7" pageNumber="8">ICZN</collectionCode>
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1999
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</bibRefCitation>
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, Art. 67.2.2).
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</paragraph>
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<paragraph id="A8443A78FFD2FFA7D8B4F89BFE73F838" blockId="7.[151,1436,1826,2012]" pageId="7" pageNumber="8">
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The authorship of
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<taxonomicName id="6FFB41FBFFD2FFA7D9EFF89AFDB8F868" box="[413,631,1907,1932]" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hispida">
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<emphasis id="9A8FE66AFFD2FFA7D9EFF89AFDB8F868" box="[413,631,1907,1932]" italics="true" pageId="7" pageNumber="8">Dynomene hispida</emphasis>
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</taxonomicName>
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, usually credited to Guérin or
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<bibRefCitation id="CC6A4789FFD2FFA7DBAEF89BFAC8F868" author="Guerin-Meneville" box="[988,1287,1906,1932]" pageId="7" pageNumber="24" refString="Guerin-Meneville, F. E. (1832) Iconographie du Regne Animal de G. Cuvier (1829 - 1844). J. - B. Baillere, Paris, 48 pp. [Livraison 22, published 14 July, 1832]." type="book chapter" year="1832">Guérin-Méneville (1832)</bibRefCitation>
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, is incorrect, the first author to describe this species was actually Latreille (
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<emphasis id="9A8FE66AFFD2FFA7DB15F872FCB1F850" box="[871,894,1947,1972]" italics="true" pageId="7" pageNumber="8">in</emphasis>
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Milbert 1812) (see
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<bibRefCitation id="CC6A4789FFD2FFA7DC16F873FAE0F850" author="Cleva" box="[1124,1327,1946,1972]" pageId="7" pageNumber="24" refString="Cleva, R., Guinot D. & Albenga, L. (2007) Annotated catalogue of brachyuran type specimens (Crustacea, Decapoda, Brachyura) deposited in the Museum national d'Histoire naturelle, Paris. Part I. Podotremata. Zoosystema, 29 (2), 229 - 279." type="journal article" year="2007">
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Cleva
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<emphasis id="9A8FE66AFFD2FFA7DCC2F872FB25F850" box="[1200,1258,1947,1972]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
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2007
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</bibRefCitation>
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: 246, fig. 14C; Ng
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<emphasis id="9A8FE66AFFD2FFA7D970F82AFEF2F838" box="[258,317,1987,2012]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
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2008: 37).
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</paragraph>
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<paragraph id="A8443A78FFDDFFA8D8B4FF7EFE7AFF3E" blockId="8.[151,1436,151,218]" pageId="8" pageNumber="9">
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<emphasis id="9A8FE66AFFDDFFA8D8B4FF7EFE1CFF55" bold="true" box="[198,467,151,177]" pageId="8" pageNumber="9">Other genus included</emphasis>
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.
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<taxonomicName id="6FFB41FBFFDDFFA8D991FF71FC97FF56" authority="McLay, 1999" authorityName="McLay" authorityYear="1999" box="[483,856,152,178]" class="Malacostraca" family="Dynomenidae" genus="Hirsutodynomene" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="9A8FE66AFFDDFFA8D991FF71FD7CFF55" box="[483,691,152,177]" italics="true" pageId="8" pageNumber="9">Hirsutodynomene</emphasis>
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<bibRefCitation id="CC6A4789FFDDFFA8DAC9FF71FC97FF56" author="McLay" box="[699,856,152,178]" pageId="8" pageNumber="25" refString="McLay, C. L. (1999) Crustacea Decapoda: Revision of the Family Dynomenidae. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, Volume 20. Memoires du Museum national d'Histoire naturelle, 180, 427 - 569." type="journal article" year="1999">McLay, 1999</bibRefCitation>
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</taxonomicName>
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(
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<typeStatus id="774084DAFFDDFFA8DB15FF71FC54FF56" box="[871,923,152,178]" pageId="8" pageNumber="9">type</typeStatus>
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species by original designation:
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<taxonomicName id="6FFB41FBFFDDFFA8DD6DFF71FE66FF3E" authority="Rathbun, 1917" authorityName="Rathbun" authorityYear="1917" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinosa">
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<emphasis id="9A8FE66AFFDDFFA8DD6DFF71FF3EFF3D" italics="true" pageId="8" pageNumber="9">Dynomene spinosa</emphasis>
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Rathbun, 1917
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<caption id="FC846AF0FFDDFFA8D8E5FAA5FDF4F978" httpUri="https://zenodo.org/record/183394/files/figure.png" pageId="8" pageNumber="9" targetBox="[283,1303,274,1311]" targetPageId="8">
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<paragraph id="A8443A78FFDDFFA8D8E5FAA5FDF4F978" blockId="8.[151,1437,1356,1692]" pageId="8" pageNumber="9">
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<emphasis id="9A8FE66AFFDDFFA8D8E5FAA5FEDBFA80" bold="true" box="[151,276,1356,1380]" pageId="8" pageNumber="9">FIGURE 3</emphasis>
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.
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<taxonomicName id="6FFB41FBFFDDFFA8D957FAA4FE24FA80" box="[293,491,1357,1380]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Metadynomeninae">Metadynomeninae</taxonomicName>
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<emphasis id="9A8FE66AFFDDFFA8D986FAA5FDA1FA80" bold="true" box="[500,622,1356,1380]" pageId="8" pageNumber="9">n. subfam.</emphasis>
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(A, B) and
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<taxonomicName id="6FFB41FBFFDDFFA8DA87FAA4FC45FA80" box="[757,906,1357,1380]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Dynomeninae">Dynomeninae</taxonomicName>
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Ortmann (C, D), views of thoracic sternum and abdomen (A, C, D) and ventral surface of abdomen (B).
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<emphasis id="9A8FE66AFFDDFFA8DB70FA86FCDAFA63" bold="true" box="[770,789,1391,1415]" pageId="8" pageNumber="9">A</emphasis>
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,
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<taxonomicName id="6FFB41FBFFDDFFA8DB56FA99FB17FA62" authority="Yokoya, 1933" authorityName="Yokoya" authorityYear="1933" box="[804,1240,1391,1415]" class="Malacostraca" family="Dynomenidae" genus="Metadynomene" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tanensis">
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||
<emphasis id="9A8FE66AFFDDFFA8DB56FA99FBE9FA63" box="[804,1062,1392,1415]" italics="true" pageId="8" pageNumber="9">Metadynomene tanensis</emphasis>
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(Yokoya, 1933)
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</taxonomicName>
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, male 12,5 x 13,5 mm, New Caledonia, SMIB 2 (MNHN-B25583): abdominal holding by P2 coxa;
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<emphasis id="9A8FE66AFFDDFFA8DC67FA7BFBE8FA4E" bold="true" box="[1045,1063,1426,1450]" pageId="8" pageNumber="9">B</emphasis>
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,
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<taxonomicName id="6FFB41FBFFDDFFA8DC44FA7BFF00FA28" authority="McLay, 1999" authorityName="McLay" authorityYear="1999" class="Malacostraca" family="Dynomenidae" genus="Metadynomene" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crosnieri">
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||
<emphasis id="9A8FE66AFFDDFFA8DC44FA7BFA8EFA4D" box="[1078,1345,1426,1449]" italics="true" pageId="8" pageNumber="9">Metadynomene crosnieri</emphasis>
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McLay, 1999
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</taxonomicName>
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, holotype, male, 23.2 x 22.7 mm, western Indian Ocean, îles Glorieuses, Benthedi Exp. (MNHN-B22510): biramous vestigial pleopods 3–5; detail at right;
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<emphasis id="9A8FE66AFFDDFFA8DAD6FA3EFD78FA0B" bold="true" box="[676,695,1495,1519]" pageId="8" pageNumber="9">C</emphasis>
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||
,
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<taxonomicName id="6FFB41FBFFDDFFA8DAB7FA31FC42FA0B" box="[709,909,1496,1519]" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hispida">
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<emphasis id="9A8FE66AFFDDFFA8DAB7FA31FC42FA0B" box="[709,909,1496,1519]" italics="true" pageId="8" pageNumber="9">Dynomene hispida</emphasis>
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</taxonomicName>
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(Latreille,
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<emphasis id="9A8FE66AFFDDFFA8DC75FA31FBD3FA0B" box="[1031,1052,1496,1519]" italics="true" pageId="8" pageNumber="9">in</emphasis>
|
||
Milbert 1812), male 6.9 x 8.8 mm, New Caledonia (MNHN-B22091): abdominal holding mechanism by sternal structure;
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||
<emphasis id="9A8FE66AFFDDFFA8DC2CFA13FBBEF9F6" bold="true" box="[1118,1137,1530,1554]" pageId="8" pageNumber="9">D</emphasis>
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,
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<taxonomicName id="6FFB41FBFFDDFFA8DCF0FA13FE85F9D0" authority="Rathbun, 1911" authorityName="Rathbun" authorityYear="1911" class="Malacostraca" family="Dynomenidae" genus="Hirsutodynomene" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinosa">
|
||
<emphasis id="9A8FE66AFFDDFFA8DCF0FA13FA53F9F5" box="[1154,1436,1530,1553]" italics="true" pageId="8" pageNumber="9">Hirsutodynomene spinosa</emphasis>
|
||
(Rathbun, 1911)
|
||
</taxonomicName>
|
||
, male, 14.3 x 16.4 mm, western Indian Ocean, îles Glorieuses (MNHN-B6899): abdominal holding mechanism by sternal structure (see Bouchard 2000: fig. 24A). a3–a6, abdominal somites 3–6; cx1–cx4, coxae of P1–P4; e4–e6, episternites 4–6; pl3–pl5, vestigial pleopods on abdominal somites 3–5; pr, coxal projection; s, shield; t, telson; u, uropod; 4, sternite 4. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E0E169F3FFDDFFA9D8B4F926FEFDFB26" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
|
||
<paragraph id="A8443A78FFDDFFA9D8B4F926FEFDFB26" blockId="8.[151,1436,1743,2010]" lastBlockId="9.[151,1437,152,1338]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<emphasis id="9A8FE66AFFDDFFA8D8B4F926FEF0F90D" bold="true" box="[198,319,1743,1769]" pageId="8" pageNumber="9">Diagnosis</emphasis>
|
||
. Carapace wider or much wider than long, broadly rounded in outline, with “xanthoid” facies; dorsal surface moderately convex, may be areolate, smooth or granulate, may be laterally spiny, sparsely covered with short or long setae. Cervical groove as broad V, usually pronounced, never reaching, thus not forming notch with lateral carapace margin, interrupted between elongated gastric pits, or not interrupted; branchial groove faint, generally indistinct laterally or, rarely, joining lateral border of carapace; branchiocardiac variously marked. Anterolateral margin beginning slightly below infraorbital border, well defined, armed with distinct teeth, rarely by granules only. Posterior margin concave. Frontal margin broadly triangular, continuous above orbit; supraorbital margin gently oblique, may be notched, usually ornamented; infraorbital margin usually irregular, may be toothed, granular, notched. Orbits well defined, directed more or less obliquely; eyestalks rather short. Antenna with narrow urinal article beaked medially; second article with exopod firmly fixed. Proepistome very narrow. Anterior border of endostome not prominently raised. Mxp3 not firmly operculiform; basis separated from ischium by incomplete suture. Branchiostegite decalcified. Thoracic sternum slightly tilted posteriorly, wide, not entirely covered laterally by abdomen at level of P2–P3 coxae, a small portion of episternites 5, 6 remaining exposed when male abdomen folded. Sternites 1, 2 fused as pentagonal shield, variously pointed at anterior end; sternite 3 represented by narrow band at base of shield, delimited posteriorly by change in level (
|
||
<figureCitation id="30C026FDFFDCFFA9DA09FE31FD10FE16" box="[635,735,472,498]" captionStart="FIGURE 3" captionStartId="8.[151,255,1356,1380]" captionTargetBox="[283,1303,274,1311]" captionTargetId="figure@8.[283,1304,274,1333]" captionTargetPageId="8" captionText="FIGURE 3. Metadynomeninae n. subfam. (A, B) and Dynomeninae Ortmann (C, D), views of thoracic sternum and abdomen (A, C, D) and ventral surface of abdomen (B). A, Metadynomene tanensis (Yokoya, 1933), male 12, 5 x 13, 5 mm, New Caledonia, SMIB 2 (MNHN-B 25583): abdominal holding by P 2 coxa; B, Metadynomene crosnieri McLay, 1999, holotype, male, 23.2 x 22.7 mm, western Indian Ocean, îles Glorieuses, Benthedi Exp. (MNHN-B 22510): biramous vestigial pleopods 3 – 5; detail at right; C, Dynomene hispida (Latreille, in Milbert 1812), male 6.9 x 8.8 mm, New Caledonia (MNHN-B 22091): abdominal holding mechanism by sternal structure; D, Hirsutodynomene spinosa (Rathbun, 1911), male, 14.3 x 16.4 mm, western Indian Ocean, îles Glorieuses (MNHN-B 6899): abdominal holding mechanism by sternal structure (see Bouchard 2000: fig. 24 A). a 3 – a 6, abdominal somites 3 – 6; cx 1 – cx 4, coxae of P 1 – P 4; e 4 – e 6, episternites 4 – 6; pl 3 – pl 5, vestigial pleopods on abdominal somites 3 – 5; pr, coxal projection; s, shield; t, telson; u, uropod; 4, sternite 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/183394/files/figure.png" pageId="9" pageNumber="10">Figure 3</figureCitation>
|
||
C, 4A); suture 4/5 indistinct but limit between sternites 4, 5 marked transversely by setiferous, convex, line, may be salient, ornamented; sternites 4–8 mostly fused into single wide plate. Sterno-coxal depressions deep. Female sutures 7/8 ending well apart, inserted in ridged groove along internal border of sterno-coxal depression for P3; spermathecal aperture slightly posterior to coxal female gonopore, tiny, below prominence. Sterno-abdominal depression wide, shallow. Male abdomen with all somites free, wide, not entirely filling sterno-abdominal depression, leaving exposed thoracic sternites 1–3, mostly (
|
||
<figureCitation id="30C026FDFFDCFFA9D942FD21FE5BFD06" box="[304,404,712,738]" captionStart="FIGURE 3" captionStartId="8.[151,255,1356,1380]" captionTargetBox="[283,1303,274,1311]" captionTargetId="figure@8.[283,1304,274,1333]" captionTargetPageId="8" captionText="FIGURE 3. Metadynomeninae n. subfam. (A, B) and Dynomeninae Ortmann (C, D), views of thoracic sternum and abdomen (A, C, D) and ventral surface of abdomen (B). A, Metadynomene tanensis (Yokoya, 1933), male 12, 5 x 13, 5 mm, New Caledonia, SMIB 2 (MNHN-B 25583): abdominal holding by P 2 coxa; B, Metadynomene crosnieri McLay, 1999, holotype, male, 23.2 x 22.7 mm, western Indian Ocean, îles Glorieuses, Benthedi Exp. (MNHN-B 22510): biramous vestigial pleopods 3 – 5; detail at right; C, Dynomene hispida (Latreille, in Milbert 1812), male 6.9 x 8.8 mm, New Caledonia (MNHN-B 22091): abdominal holding mechanism by sternal structure; D, Hirsutodynomene spinosa (Rathbun, 1911), male, 14.3 x 16.4 mm, western Indian Ocean, îles Glorieuses (MNHN-B 6899): abdominal holding mechanism by sternal structure (see Bouchard 2000: fig. 24 A). a 3 – a 6, abdominal somites 3 – 6; cx 1 – cx 4, coxae of P 1 – P 4; e 4 – e 6, episternites 4 – 6; pl 3 – pl 5, vestigial pleopods on abdominal somites 3 – 5; pr, coxal projection; s, shield; t, telson; u, uropod; 4, sternite 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/183394/files/figure.png" pageId="9" pageNumber="10">Figure 3</figureCitation>
|
||
C) or totally (
|
||
<figureCitation id="30C026FDFFDCFFA9DA40FD21FD5BFD06" box="[562,660,712,738]" captionStart="FIGURE 3" captionStartId="8.[151,255,1356,1380]" captionTargetBox="[283,1303,274,1311]" captionTargetId="figure@8.[283,1304,274,1333]" captionTargetPageId="8" captionText="FIGURE 3. Metadynomeninae n. subfam. (A, B) and Dynomeninae Ortmann (C, D), views of thoracic sternum and abdomen (A, C, D) and ventral surface of abdomen (B). A, Metadynomene tanensis (Yokoya, 1933), male 12, 5 x 13, 5 mm, New Caledonia, SMIB 2 (MNHN-B 25583): abdominal holding by P 2 coxa; B, Metadynomene crosnieri McLay, 1999, holotype, male, 23.2 x 22.7 mm, western Indian Ocean, îles Glorieuses, Benthedi Exp. (MNHN-B 22510): biramous vestigial pleopods 3 – 5; detail at right; C, Dynomene hispida (Latreille, in Milbert 1812), male 6.9 x 8.8 mm, New Caledonia (MNHN-B 22091): abdominal holding mechanism by sternal structure; D, Hirsutodynomene spinosa (Rathbun, 1911), male, 14.3 x 16.4 mm, western Indian Ocean, îles Glorieuses (MNHN-B 6899): abdominal holding mechanism by sternal structure (see Bouchard 2000: fig. 24 A). a 3 – a 6, abdominal somites 3 – 6; cx 1 – cx 4, coxae of P 1 – P 4; e 4 – e 6, episternites 4 – 6; pl 3 – pl 5, vestigial pleopods on abdominal somites 3 – 5; pr, coxal projection; s, shield; t, telson; u, uropod; 4, sternite 4. Scale bars: 1 mm." httpUri="https://zenodo.org/record/183394/files/figure.png" pageId="9" pageNumber="10">Figure 3</figureCitation>
|
||
D) sternite 4, and episternites 5, 6; first abdominal somite dorsally visible, narrow, slightly wider than somite 2. Somites 3–5 of males with vestigial, rudimentary pleopods, biramous, rarely uniramous. Pl1 vestigial in females. Uropods sexually dimorphic, immobile, rather large, occupying variable length of abdominal somite 6, not filling whole length of somite
|
||
<quantity id="6F03979DFFDCFFA9DCFBFCA9FB77FCBE" box="[1161,1208,832,858]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="10" unit="in" value="6.0">6 in</quantity>
|
||
males and females (
|
||
<figureCitation id="30C026FDFFDCFFA9D8EDFC81FECAFC66" box="[159,261,872,898]" captionStart="FIGURE 5" captionStartId="15.[151,255,1445,1469]" captionTargetBox="[285,1309,322,1417]" captionTargetId="figure@15.[274,1314,314,1421]" captionTargetPageId="15" captionText="FIGURE 5. Uropods in the four dynomenid subfamilies (dorsal surface, unless mentioned otherwise). A – H, Dynomeninae Ortmann, 1892; I – L, Metadynomeninae n. subfam.; M – N, Paradynomeninae n. subfam.; O – P, Acanthodromiinae n. subfam. A – B, Dynomene hispida (Latreille, in Milbert 1812): A, male, New Caledonia (MNHN-B 22091); B, female (After McLay 1999: fig. 18 g). C, Dynomene pilumnoides Alcock, 1900, female (After McLay 1999: fig. 21 g). D, Dynomene praedator A. Milne-Edwards, 1879, male (After McLay 1999: fig. 19 g). E, Dynomene pugnatrix De Man, 1889, male (After McLay 1999: fig. 22 g). F, Dynomene filholi Bouvier, 1894, male (After McLay 1999: fig. 20 g). G, Hirsutodynomene spinosa (Rathbun, 1911), male (After McLay 1999: fig. 23 g). H, Hirsutodynomene ursula (Stimpson, 1860), female (After McLay 1999: fig. 24 g). I – J, Metadynomene tanensis (Yokoya, 1933): I, male, New Caledonia, dorsal surface (MNHN-B 25583); J, female, ventral surface (After McLay 1999: fig. 27 g). K – L, Metadynomene crosnieri McLay, 1999, male, holotype, western Indian Ocean, îles Glorieuses (MNHN-B 22510): dorsal (K) and ventral (L) surfaces; M – N, Paradynomene sp., New Caledonia (MNHN-B 24779); M, male: ventral surface; N, female: dorsal surface (After McLay 1999: fig. 32 g). O, Acanthodromia erinacea A. Milne-Edwards, 1880, female, Puerto Rico (USNM 124263); P, Acanthodromia margarita (Alcock, 1899), female (After McLay 1999: fig. 31 f)." httpUri="https://zenodo.org/record/183396/files/figure.png" pageId="9" pageNumber="10">Figure 5</figureCitation>
|
||
A, C–H) in contrast to females in some species (
|
||
<figureCitation id="30C026FDFFDCFFA9DB24FC81FC72FC66" box="[854,957,872,898]" captionStart="FIGURE 5" captionStartId="15.[151,255,1445,1469]" captionTargetBox="[285,1309,322,1417]" captionTargetId="figure@15.[274,1314,314,1421]" captionTargetPageId="15" captionText="FIGURE 5. Uropods in the four dynomenid subfamilies (dorsal surface, unless mentioned otherwise). A – H, Dynomeninae Ortmann, 1892; I – L, Metadynomeninae n. subfam.; M – N, Paradynomeninae n. subfam.; O – P, Acanthodromiinae n. subfam. A – B, Dynomene hispida (Latreille, in Milbert 1812): A, male, New Caledonia (MNHN-B 22091); B, female (After McLay 1999: fig. 18 g). C, Dynomene pilumnoides Alcock, 1900, female (After McLay 1999: fig. 21 g). D, Dynomene praedator A. Milne-Edwards, 1879, male (After McLay 1999: fig. 19 g). E, Dynomene pugnatrix De Man, 1889, male (After McLay 1999: fig. 22 g). F, Dynomene filholi Bouvier, 1894, male (After McLay 1999: fig. 20 g). G, Hirsutodynomene spinosa (Rathbun, 1911), male (After McLay 1999: fig. 23 g). H, Hirsutodynomene ursula (Stimpson, 1860), female (After McLay 1999: fig. 24 g). I – J, Metadynomene tanensis (Yokoya, 1933): I, male, New Caledonia, dorsal surface (MNHN-B 25583); J, female, ventral surface (After McLay 1999: fig. 27 g). K – L, Metadynomene crosnieri McLay, 1999, male, holotype, western Indian Ocean, îles Glorieuses (MNHN-B 22510): dorsal (K) and ventral (L) surfaces; M – N, Paradynomene sp., New Caledonia (MNHN-B 24779); M, male: ventral surface; N, female: dorsal surface (After McLay 1999: fig. 32 g). O, Acanthodromia erinacea A. Milne-Edwards, 1880, female, Puerto Rico (USNM 124263); P, Acanthodromia margarita (Alcock, 1899), female (After McLay 1999: fig. 31 f)." httpUri="https://zenodo.org/record/183396/files/figure.png" pageId="9" pageNumber="10">Figure 5</figureCitation>
|
||
B). Abdominal holding structure sternal (but only restricting lateral movements of abdomen); in males a small tubercle on episternite 5 facing either uropod margin or abdomen margin (location dependent on size of uropod and its extension along somite 6); tubercle lost in mature females. Chelipeds equal, either stout or slender; fingers largely gaping at base, dactylus strongly curved. P2–P4 conspicuously ornamented with spines or granules; dactyli with 4–6 spines on inferior margin. P5 sexually dimorphic, reduced; basis and ischium free, not fused to merus; dactylus rudimentary, with obsolete subchelate mechanism. Coxa of P5 modified in males, extended to enclose penis. Gonopod 1 stout, forming half-rolled tube, with apical plate. G2 needle-like, with varying number of subterminal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E0E169F3FFDCFFA9D8B4FB26FE31FADE" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="A8443A78FFDCFFA9D8B4FB26FE31FADE" blockId="9.[151,1437,152,1338]" pageId="9" pageNumber="10">
|
||
<emphasis id="9A8FE66AFFDCFFA9D8B4FB26FEF7FB0D" bold="true" box="[198,312,1231,1257]" pageId="9" pageNumber="10">Remarks</emphasis>
|
||
. The subfamily comprises only two genera:
|
||
<taxonomicName id="6FFB41FBFFDCFFA9DB39FB39FC07FB0D" box="[843,968,1232,1257]" class="Malacostraca" family="Dynomenidae" genus="Dynomene" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9A8FE66AFFDCFFA9DB39FB39FC07FB0D" box="[843,968,1232,1257]" italics="true" pageId="9" pageNumber="10">Dynomene</emphasis>
|
||
</taxonomicName>
|
||
, with
|
||
<bibRefCitation id="CC6A4789FFDCFFA9DC66FB39FB01FB0E" author="McLay" box="[1044,1230,1232,1258]" pageId="9" pageNumber="25" refString="McLay, C. L. (2001 b) Dynomenidae and Dromiidae (Decapoda: Brachyura) from Guam, Philippine Islands, Tonga and Samoa. Zoosystema, 23 (4), 807 - 856." type="journal article" year="2001" yearSuffix="b">McLay (2001b)</bibRefCitation>
|
||
recently describing two new species from
|
||
<collectingCountry id="D0EC7AE8FFDCFFA9D9A7FB11FDEFFAF6" box="[469,544,1272,1298]" name="Guam" pageId="9" pageNumber="10">Guam</collectingCountry>
|
||
, and
|
||
<taxonomicName id="6FFB41FBFFDCFFA9DA12FB11FCFFFAF5" box="[608,816,1272,1297]" class="Malacostraca" family="Dynomenidae" genus="Hirsutodynomene" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9A8FE66AFFDCFFA9DA12FB11FCFFFAF5" box="[608,816,1272,1297]" italics="true" pageId="9" pageNumber="10">Hirsutodynomene</emphasis>
|
||
</taxonomicName>
|
||
, with
|
||
<bibRefCitation id="CC6A4789FFDCFFA9DB0CFB11FBB8FAF6" author="McLay" box="[894,1143,1272,1298]" pageId="9" pageNumber="25" refString="McLay, C. L & Ng, P. K. L. (2005) On a collection of Dromiidae and Dynomenidae from the Philippines, with the description of a new species of Hirsutodynomene McLay, 1999 (Crustacea: Decapoda: Brachyura). Zootaxa, 1029, 1 - 30." type="journal article" year="2005">McLay & Ng (2005)</bibRefCitation>
|
||
adding one new species from the
|
||
<collectingCountry id="D0EC7AE8FFDCFFA9D970FAC9FE49FADE" box="[258,390,1312,1338]" name="Philippines" pageId="9" pageNumber="10">Philippines</collectingCountry>
|
||
(
|
||
<tableCitation id="E5790FC3FFDCFFA9D9E7FAC9FE22FADE" box="[405,493,1312,1338]" captionStart="TABLE 1" captionStartId="1.[151,239,391,415]" captionText="TABLE 1. List of extant genera and species currently placed in the family Dynomenidae Ortmann, 1892 (after Ng et al. 2008: 37). Generic names in square parentheses are the genera in which the species were originally described" pageId="9" pageNumber="10">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |