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<document id="30A72395689F9DA2BC2A3BDA645D360A" ID-CLB-Dataset="4184" ID-DOI="10.5852/ejt.2021.768.1501" ID-GBIF-Dataset="1361fbca-2ed1-4f60-85cc-1a3f62f73d79" ID-ISSN="2118-9773" ID-Zenodo-Dep="5535911" ID-ZooBank="58117DB6-B3F3-41BF-81B3-8AF089708922" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632918358292" checkinUser="felipe" docAuthor="Annisaqois, Manikmayang &amp; Wägele, J. Wolfgang" docDate="2021" docId="1A6987B821141F0BFDCD6024FC6EC4D3" docLanguage="en" docName="ejt-768_1-52.pdf" docOrigin="European Journal of Taxonomy 768" docSource="http://dx.doi.org/10.5852/ejt.2021.768.1501" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pendanthura bangkaensis Annisaqois &amp; Wägele 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="26" masterDocId="E650FFC021001F12FF846470FFA2C33A" masterDocTitle="Morphology and Taxonomy of Isopoda Anthuroidea (Crustacea) from Sulawesi with description of six new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="21" updateTime="1699458108869" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B1F537C2F04E00053AA57F032FC72A5B">Morphology and Taxonomy of Isopoda Anthuroidea (Crustacea) from Sulawesi with description of six new species</mods:title>
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<mods:namePart id="4C425DCCA82EA013A5B1B6847AD52280">Annisaqois, Manikmayang</mods:namePart>
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<mods:affiliation id="011CA593617D90070B35620474B41870">Faculty of Fisheries and Marine Sciences, Sam Ratulangi University, Jln Kampus UNSRAT - Bahu, Manado 95115, Indonesia.</mods:affiliation>
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<mods:namePart id="3DAF7E4C5CD2CC7F3FD47A4ED1981883">Wägele, J. Wolfgang</mods:namePart>
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<mods:affiliation id="FA8C8595D7EF6A68ED12EC6BFECFECCF">Zoological Research Museum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany.</mods:affiliation>
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<paragraph id="927F36AE21141F06FDCD6024FC49C755" blockId="20.[358,1230,1108,1206]" box="[585,1003,1108,1135]" pageId="20" pageNumber="21">
<heading id="C93781C221141F06FDCD6024FC49C755" box="[585,1003,1108,1135]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="21" reason="2">
<taxonomicName id="55C04D2D21141F06FDCD6024FC2BC755" ID-CoL="8QCD5" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[585,905,1108,1135]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21141F06FDCD6024FC2BC755" bold="true" box="[585,905,1108,1135]" italics="true" pageId="20" pageNumber="21">Pendanthura bangkaensis</emphasis>
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<taxonomicNameLabel id="BB8757C721141F06FC156025FC49C755" box="[913,1003,1109,1135]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="0AFB2A2B21141F06FD5C60ECFCF9C78C" box="[728,859,1180,1206]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="21.[189,232,1963,1989]" captionStartId-1="22.[189,232,1963,1989]" captionStartId-2="23.[189,232,1963,1989]" captionStartId-3="24.[189,232,1963,1989]" captionTargetBox-0="[253,1356,265,1891]" captionTargetBox-2="[208,1336,273,1895]" captionTargetBox-3="[223,1335,685,1908]" captionTargetId-0="figure-12@21.[204,1364,262,1902]" captionTargetId-1="figure-15@22.[186,1399,299,1904]" captionTargetId-2="figure-12@23.[204,1374,265,1915]" captionTargetId-3="figure-61@24.[192,1389,641,1923]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="Fig. 14. Pendanthura bangkaensis sp. nov., holotype, ♀, in dorsal and lateral view (3.4 mm) (MZB Iso 104). Abbreviations: A1 = antenna 1; A2 = antenna 2; Md = mandible; Mx = maxilla; Mxp = maxilliped." captionText-1="Fig. 15. Pendanthura bangkaensis sp. nov., ♀, holotype (MZB Iso 104). Abbreviations: P1–P7 = pereopods 1–7." captionText-2="Fig. 16. Pendanthura bangkaensis sp. nov., ♀, holotype (MZB Iso 104). Abbreviations: Plp 1–Plp 5 = pleopods 1–5." captionText-3="Fig. 17. Pendanthura bangkaensis sp. nov., ♀, holotype (MZB Iso 104). Abbreviations: UEx = exopod of uropod; UEn = endopod of uropod; Tel = telson." figureDoi-0="http://doi.org/10.5281/zenodo.5535965" figureDoi-1="http://doi.org/10.5281/zenodo.5538795" figureDoi-2="http://doi.org/10.5281/zenodo.5535969" figureDoi-3="http://doi.org/10.5281/zenodo.5535971" httpUri-0="https://zenodo.org/record/5535965/files/figure.png" httpUri-1="https://zenodo.org/record/5538795/files/figure.png" httpUri-2="https://zenodo.org/record/5535969/files/figure.png" httpUri-3="https://zenodo.org/record/5535971/files/figure.png" pageId="20" pageNumber="21">Figs 14–17</figureCitation>
</paragraph>
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<subSubSection id="DADA652521141F06FF3960ADFE27C4D2" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="927F36AE21141F06FF3960ADFE6BC7CD" blockId="20.[189,457,1245,1271]" box="[189,457,1245,1271]" pageId="20" pageNumber="21">
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<emphasis id="A0B4EABC21141F06FF3960ADFE6BC7CD" bold="true" box="[189,457,1245,1271]" pageId="20" pageNumber="21">Differential diagnosis</emphasis>
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<paragraph id="927F36AE21141F06FF39617DFC50C57E" blockId="20.[187,1400,1293,1604]" pageId="20" pageNumber="21">
Species of 
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<emphasis id="A0B4EABC21141F06FEC4617DFE79C61D" box="[320,475,1293,1319]" italics="true" pageId="20" pageNumber="21">Pendanthura</emphasis>
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with dorsally pigmented body, distinct lateral eyes with few large ommatidia, mandible palp with 1 tiny article and a long apical seta, pereopod 1 carpus triangular with protruding edge distally, propodus lower half protruding, forming a rounded corner near palm center, with 14 setae in mesial row on distal half of propodus, and 5 longer setae more proximally; pereopods 2–3 merus triangular, dorsal edge reaching nearly to propodus, uropod sympod elongate, endopod subovate, medially as long as laterally, about 0.3 sympod length, exopod twice length of endopod, long oval, margin with many short feathered setae; margins of endopod, exopod and tip of telson weakly crenulated. Telson tongue-shaped, with a pair of statocyst proximally, rounded apex bearing 3 pairs of setae, and on dorsal surface another 2 pairs of longer setae and 1 pair of short setae.
</paragraph>
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Few species of 
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<emphasis id="A0B4EABC21141F06FEFF621BFDB4C5BF" box="[379,534,1643,1669]" italics="true" pageId="20" pageNumber="21">Pendanthura</emphasis>
</taxonomicName>
share with the new species the distinct projection near the center of the palm of P1 propodus. This feature also occurs in 
<taxonomicName id="55C04D2D21141F06FC8062FFFBEEC593" authority="Kensley, 1979" authorityName="Kensley" authorityYear="1979" box="[772,1100,1678,1705]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="rarotonga">
<emphasis id="A0B4EABC21141F06FC8062FFFC3BC592" box="[772,921,1679,1704]" italics="true" pageId="20" pageNumber="21">P. rarotonga</emphasis>
<bibRefCitation id="F6514B5F21141F06FC2462FEFBEEC593" author="Kensley B." box="[928,1100,1678,1705]" pageId="20" pageNumber="21" pagination="814 - 836" refId="ref13186" refString="Kensley B. 1979. New species of anthurideans from the Cook and Fiji Islands (Crustacea: Isopoda: Anthuridea). Proceedings of the Biological Society of Washington 92 (4): 814 - 836." type="journal article" year="1979">Kensley, 1979</bibRefCitation>
</taxonomicName>
. 
<bibRefCitation id="F6514B5F21141F06FBDD62FEFB5CC593" author="Muller H. G." box="[1113,1278,1678,1705]" pageId="20" pageNumber="21" pagination="353 - 371" refId="ref13331" refString="Muller H. G. 1992. Anthuridae from coral reefs at Bora Bora and Moorea, Society Islands, with description of three new species (Crustacea: Isopoda). Senckenbergiana Biologica 72: 353 - 371." type="journal article" year="1992">Müller (1992)</bibRefCitation>
described a specimen of 
<taxonomicName id="55C04D2D21141F06FEF462C3FDA5C5F6" authorityName="Kensley" authorityYear="1979" box="[368,519,1715,1740]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="rarotonga">
<emphasis id="A0B4EABC21141F06FEF462C3FDA5C5F6" box="[368,519,1715,1740]" italics="true" pageId="20" pageNumber="21">P. rarotonga</emphasis>
</taxonomicName>
from Bora Bora with 1 article and 2 short setae of the mandibular palp, and a pereopod 1 palm more convex in its lower half and with a sharper protrusion than in Kensley’s material (1979). Müller’s species from Bora Bora is probably not the same as 
<taxonomicName id="55C04D2D21141F06FBDD628AFB4CC429" authorityName="Kensley" authorityYear="1979" box="[1113,1262,1786,1811]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="rarotonga">
<emphasis id="A0B4EABC21141F06FBDD628AFB4CC429" box="[1113,1262,1786,1811]" italics="true" pageId="20" pageNumber="21">P. rarotonga</emphasis>
</taxonomicName>
of 
<bibRefCitation id="F6514B5F21141F06FA936289FEAEC40D" author="Kensley B." pageId="20" pageNumber="21" pagination="814 - 836" refId="ref13186" refString="Kensley B. 1979. New species of anthurideans from the Cook and Fiji Islands (Crustacea: Isopoda: Anthuridea). Proceedings of the Biological Society of Washington 92 (4): 814 - 836." type="journal article" year="1979">Kensley (1979)</bibRefCitation>
, which occurs on Rarotonga Island. In 
<taxonomicName id="55C04D2D21141F06FD74636CFB85C40C" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[752,1063,1820,1846]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21141F06FD74636CFB85C40C" box="[752,1063,1820,1846]" italics="true" pageId="20" pageNumber="21">Pendanthura bangkaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721141F06FBB7636DFB32C40D" box="[1075,1168,1821,1847]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
from 
<collectingRegion id="5004F84C21141F06FB5A636CFAEAC40C" box="[1246,1352,1820,1846]" country="Indonesia" name="Sulawesi Utara" pageId="20" pageNumber="21">Sulawesi</collectingRegion>
the palm projection of the female is similar, but the mesial row of setae has more setae (14 instead of 8). Also, in 
<taxonomicName id="55C04D2D21141F06FEA46314FE11C447" authorityName="Kensley" authorityYear="1979" box="[288,435,1892,1917]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="rarotonga">
<emphasis id="A0B4EABC21141F06FEA46314FE11C447" box="[288,435,1892,1917]" italics="true" pageId="20" pageNumber="21">P. rarotonga</emphasis>
</taxonomicName>
the telson bears dorsally 4–7 pairs of setae, in the 
<collectingRegion id="5004F84C21141F06FC7B6314FBCBC444" box="[1023,1129,1892,1918]" country="Indonesia" name="Sulawesi Utara" pageId="20" pageNumber="21">Sulawesi</collectingRegion>
material there are only 2 pairs of long setae and a pair of short seta dorsolaterally. A similar but larger P1 palm protrusion also occurs in 
<taxonomicName id="55C04D2D21141F06FEA863DCFD56C4FF" authority="Menzies &amp; Glynn, 1968" authorityName="Menzies &amp; Glynn" authorityYear="1968" box="[300,756,1963,1989]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tanaiformis">
<emphasis id="A0B4EABC21141F06FEA863DCFE70C4FF" box="[300,466,1963,1989]" italics="true" pageId="20" pageNumber="21">P. tanaiformis</emphasis>
Menzies &amp; Glynn, 1968
</taxonomicName>
, a Caribbean species. Uropod details of this species are still undescribed.
</paragraph>
</subSubSection>
<caption id="C6BF662621151F07FF3963DBFAD5C4D2" ID-DOI="http://doi.org/10.5281/zenodo.5535965" ID-Zenodo-Dep="5535965" httpUri="https://zenodo.org/record/5535965/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1963,1989]" targetBox="[253,1356,265,1891]" targetPageId="21">
<paragraph id="927F36AE21151F07FF3963DBFAD5C4D2" blockId="21.[189,1399,1963,2025]" pageId="21" pageNumber="22">
<emphasis id="A0B4EABC21151F07FF3963DBFEB8C4FF" bold="true" box="[189,282,1963,1989]" pageId="21" pageNumber="22">Fig. 14.</emphasis>
<taxonomicName id="55C04D2D21151F07FEA563DBFDF7C4FF" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[289,597,1963,1989]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21151F07FEA563DBFDF7C4FF" box="[289,597,1963,1989]" italics="true" pageId="21" pageNumber="22">Pendanthura bangkaensis</emphasis>
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<taxonomicNameLabel id="BB8757C721151F07FDD963DCFD18C4FC" box="[605,698,1964,1990]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀, in dorsal and lateral view (3.4 mm) (MZB Iso 104). Abbreviations: A1 = antenna 1; A2 = antenna 2; Md = mandible; Mx = maxilla; Mxp = maxilliped.
</paragraph>
</caption>
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<paragraph id="927F36AE21161F04FF3963DBFED2C4D2" blockId="22.[189,1399,1963,2024]" pageId="22" pageNumber="23">
<emphasis id="A0B4EABC21161F04FF3963DBFE82C4FF" bold="true" box="[189,288,1963,1989]" pageId="22" pageNumber="23">Fig. 15.</emphasis>
<taxonomicName id="55C04D2D21161F04FEA863DBFDC4C4FF" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[300,614,1963,1989]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21161F04FEA863DBFDC4C4FF" box="[300,614,1963,1989]" italics="true" pageId="22" pageNumber="23">Pendanthura bangkaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721161F04FDF763DCFD77C4FC" box="[627,725,1964,1990]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype (MZB Iso 104). Abbreviations: P1–P7 = pereopods 1–7.
</paragraph>
</caption>
<caption id="C6BF662621171F05FF3963DBFEC3C4D3" ID-DOI="http://doi.org/10.5281/zenodo.5535969" ID-Zenodo-Dep="5535969" httpUri="https://zenodo.org/record/5535969/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1963,1989]" targetBox="[208,1336,273,1895]" targetPageId="23">
<paragraph id="927F36AE21171F05FF3963DBFEC3C4D3" blockId="23.[189,1400,1963,2025]" pageId="23" pageNumber="24">
<emphasis id="A0B4EABC21171F05FF3963DBFEB9C4FF" bold="true" box="[189,283,1963,1989]" pageId="23" pageNumber="24">Fig. 16.</emphasis>
<taxonomicName id="55C04D2D21171F05FEA763DBFDF5C4FF" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[291,599,1963,1989]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21171F05FEA763DBFDF5C4FF" box="[291,599,1963,1989]" italics="true" pageId="23" pageNumber="24">Pendanthura bangkaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721171F05FDE563DCFD1CC4FC" box="[609,702,1964,1990]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype (MZB Iso 104). Abbreviations: Plp 1–Plp 5 = pleopods 1–5.
</paragraph>
</caption>
<subSubSection id="DADA652521181F0AFF396579FD72C26B" box="[189,720,265,338]" pageId="24" pageNumber="25" type="etymology">
<paragraph id="927F36AE21181F0AFF396579FEE6C219" blockId="24.[189,324,265,291]" box="[189,324,265,291]" pageId="24" pageNumber="25">
<heading id="C93781C221181F0AFF396579FEE6C219" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="A0B4EABC21181F0AFF396579FEE6C219" bold="true" box="[189,324,265,291]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21181F0AFF396547FD72C26B" blockId="24.[189,720,311,338]" box="[189,720,311,338]" pageId="24" pageNumber="25">
The species epithet refers to the 
<typeStatus id="4D7B880C21181F0AFDBD6548FDCEC268" box="[569,620,312,338]" pageId="24" pageNumber="25">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="DADA652521181F0BFF396508FB5CC254" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="927F36AE21181F0AFF396508FE08C2A8" blockId="24.[189,426,376,402]" box="[189,426,376,402]" pageId="24" pageNumber="25">
<heading id="C93781C221181F0AFF396508FE08C2A8" bold="true" box="[189,426,376,402]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="A0B4EABC21181F0AFF396508FE08C2A8" bold="true" box="[189,426,376,402]" pageId="24" pageNumber="25">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21181F0BFF6E65D7FB5CC254" blockId="24.[189,1398,423,520]" lastBlockId="25.[189,1398,265,366]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<materialsCitation id="22A83CF321181F0AFF6E65D7FB5CC132" ID-GBIF-Occurrence="3359244308" collectingDate="2019-09" collectionCode="MZB" collectorName="M. Annisaqois &amp; J. W. Wagele" country="Indonesia" elevation="2" latitude="1.7394556" location="Bangka Island" longLatPrecision="1" longitude="125.144775" municipality="Coral Eye Resort" pageId="24" pageNumber="25" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="A0B4EABC21181F0AFF6E65D7FEFEC2FB" bold="true" box="[234,348,423,449]" pageId="24" pageNumber="25">
<typeStatus id="4D7B880C21181F0AFF6E65D7FEFEC2FB" box="[234,348,423,449]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="EAD7763E21181F0AFF3965BAFEF9C2DE" box="[189,347,458,484]" name="Indonesia" pageId="24" pageNumber="25">INDONESIA</collectingCountry>
•brooding 
<specimenCount id="84C6FD2721181F0AFE5E65BBFE52C2DF" box="[474,496,459,485]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
(
<quantity id="55389B4B21181F0AFE7F65BAFDFAC2DF" box="[507,600,458,485]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="24" pageNumber="25" unit="mm" value="3.4">3.4 mm</quantity>
); 
<location id="971F607521181F0AFDEC65BAFCB2C2DE" LSID="urn:lsid:plazi:treatment:1A6987B821141F0BFDCD6024FC6EC4D3:971F607521181F0AFDEC65BAFCB2C2DE" box="[616,784,458,484]" country="Indonesia" latitude="1.7394556" longLatPrecision="1" longitude="125.144775" municipality="Coral Eye Resort" name="Bangka Island" pageId="24" pageNumber="25">Bangka Island</location>
, 
<locationDeviation id="83CD2F8C21181F0AFC9C65BAFC5DC2DF" box="[792,1023,458,485]" pageId="24" pageNumber="25">off Coral Eye Resort</locationDeviation>
; 
<geoCoordinate id="F7F4506921181F0AFB8A65B9FB14C2DF" box="[1038,1206,457,485]" degrees="1" direction="north" minutes="44" orientation="latitude" pageId="24" pageNumber="25" precision="1" seconds="22.04" value="1.7394556">1°44′22.04″N</geoCoordinate>
, 
<geoCoordinate id="F7F4506921181F0AFB3A65B9FAD3C2DF" box="[1214,1393,457,485]" degrees="125" direction="east" minutes="8" orientation="longitude" pageId="24" pageNumber="25" precision="1" seconds="41.19" value="125.144775">125°8′41.19″ E</geoCoordinate>
; 
<elevation id="19EDD19D21181F0AFF39659EFEF1C13D" box="[189,339,493,520]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="24" pageNumber="25" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">
<quantity id="55389B4B21181F0AFF39659EFEA5C132" box="[189,263,494,520]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="24" pageNumber="25" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
depth
</elevation>
; in coral rubble; 
<collectorName id="3F35537821181F0AFD9E659EFD67C13D" box="[538,709,493,520]" pageId="24" pageNumber="25">M. Annisaqois</collectorName>
and 
<collectorName id="3F35537821181F0AFD7A659EFC30C13D" box="[766,914,493,520]" pageId="24" pageNumber="25">J.W. Wägele</collectorName>
leg.; 
<date id="E67E106E21181F0AFC57659EFBEEC132" box="[979,1100,494,520]" pageId="24" pageNumber="25" value="2019-09">
<collectingDate id="F63AE98621181F0AFC57659EFBEEC132" box="[979,1100,494,520]" pageId="24" pageNumber="25" value="2019-09">Sep. 2019</collectingDate>
</date>
; 
<collectionCode id="F4D1AE6B21181F0AFBDC659EFB3AC132" box="[1112,1176,494,520]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="24" pageNumber="25" type="Museum">MZB</collectionCode>
Iso 104.
</materialsCitation>
<materialsCitation id="22A83CF321191F0BFF6E6579FB5BC254" ID-GBIF-Occurrence="3359244310" collectingDate="2019-09" collectionCode="MZB" collectorName="M. Annisaqois &amp; J. W. Wagele" country="Indonesia" elevation="2" latitude="1.7394556" location="Bangka Island" longLatPrecision="1" longitude="125.144775" municipality="Coral Eye Resort" pageId="25" pageNumber="26" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis id="A0B4EABC21191F0BFF6E6579FEC5C219" bold="true" box="[234,359,265,291]" pageId="25" pageNumber="26">
<typeStatus id="4D7B880C21191F0BFF6E6579FEC5C219" box="[234,359,265,291]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="EAD7763E21191F0BFF39655EFEF9C272" box="[189,347,302,328]" name="Indonesia" pageId="25" pageNumber="26">INDONESIA</collectingCountry>
• 
<specimenCount id="84C6FD2721191F0BFEEB655EFE0DC272" box="[367,431,302,328]" pageId="25" pageNumber="26" type="female">2 ♀♀</specimenCount>
(
<quantity id="55389B4B21191F0BFE38655EFDEFC272" box="[444,589,302,328]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="25" pageNumber="26" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.8–3.4 mm</quantity>
); 
<location id="971F607521191F0BFDDB655EFCAAC272" LSID="urn:lsid:plazi:treatment:1A6987B821141F0BFDCD6024FC6EC4D3:971F607521191F0BFDDB655EFCAAC272" box="[607,776,302,328]" country="Indonesia" latitude="1.7394556" longLatPrecision="1" longitude="125.144775" municipality="Coral Eye Resort" name="Bangka Island" pageId="25" pageNumber="26">Bangka Island</location>
, 
<locationDeviation id="83CD2F8C21191F0BFC95655EFBA2C272" box="[785,1024,302,328]" pageId="25" pageNumber="26">off Coral Eye Resort</locationDeviation>
; 
<geoCoordinate id="F7F4506921191F0BFB8E655CFB11C272" box="[1034,1203,300,328]" degrees="1" direction="north" minutes="44" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="22.04" value="1.7394556">1°44′22.04″ N</geoCoordinate>
, 
<geoCoordinate id="F7F4506921191F0BFB38655CFAD3C272" box="[1212,1393,300,328]" degrees="125" direction="east" minutes="8" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="41.19" value="125.144775">125°8′41.19″ E</geoCoordinate>
; 
<elevation id="19EDD19D21191F0BFF396523FEF1C257" box="[189,339,339,366]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="25" pageNumber="26" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">
<quantity id="55389B4B21191F0BFF396523FEA5C254" box="[189,263,339,366]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="25" pageNumber="26" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
depth
</elevation>
; in coral rubble; 
<collectorName id="3F35537821191F0BFD9E6524FD67C257" box="[538,709,339,366]" pageId="25" pageNumber="26">M. Annisaqois</collectorName>
and 
<collectorName id="3F35537821191F0BFD7A6524FC30C257" box="[766,914,339,366]" pageId="25" pageNumber="26">J.W. Wägele</collectorName>
leg.; 
<date id="E67E106E21191F0BFC576523FBEEC257" box="[979,1100,339,365]" pageId="25" pageNumber="26" value="2019-09">
<collectingDate id="F63AE98621191F0BFC576523FBEEC257" box="[979,1100,339,365]" pageId="25" pageNumber="26" value="2019-09">Sep. 2019</collectingDate>
</date>
; 
<collectionCode id="F4D1AE6B21191F0BFBDC6524FB3AC254" box="[1112,1176,340,366]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="25" pageNumber="26" type="Museum">MZB</collectionCode>
Iso 105
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="C6BF662621181F0AFF3963DBFCB6C4D2" ID-DOI="http://doi.org/10.5281/zenodo.5535971" ID-Zenodo-Dep="5535971" httpUri="https://zenodo.org/record/5535971/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1963,1989]" targetBox="[223,1335,685,1908]" targetPageId="24">
<paragraph id="927F36AE21181F0AFF3963DBFCB6C4D2" blockId="24.[189,1398,1963,2025]" pageId="24" pageNumber="25">
<emphasis id="A0B4EABC21181F0AFF3963DBFEB8C4FF" bold="true" box="[189,282,1963,1989]" pageId="24" pageNumber="25">Fig. 17.</emphasis>
<taxonomicName id="55C04D2D21181F0AFEA563DBFDF7C4FF" authority="Annisaqois &amp; Wägele, 2021" authorityName="Annisaqois &amp; Wägele" authorityYear="2021" box="[289,597,1963,1989]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bangkaensis" status="sp. nov.">
<emphasis id="A0B4EABC21181F0AFEA563DBFDF7C4FF" box="[289,597,1963,1989]" italics="true" pageId="24" pageNumber="25">Pendanthura bangkaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BB8757C721181F0AFDD963DCFD1BC4FC" box="[605,697,1964,1990]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, ♀, holotype (MZB Iso 104). Abbreviations: UEx = exopod of uropod; UEn = endopod of uropod; Tel = telson.
</paragraph>
</caption>
<subSubSection id="DADA652521191F0BFF3965E6FAC5C6DB" pageId="25" pageNumber="26" type="description">
<paragraph id="927F36AE21191F0BFF3965E6FE43C28B" blockId="25.[189,481,406,433]" box="[189,481,406,433]" pageId="25" pageNumber="26">
<heading id="C93781C221191F0BFF3965E6FE43C28B" bold="true" box="[189,481,406,433]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0B4EABC21191F0BFF3965E6FE43C28B" bold="true" box="[189,481,406,433]" pageId="25" pageNumber="26">
Description of 
<typeStatus id="4D7B880C21191F0BFEF165E7FE43C28B" box="[373,481,407,433]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21191F0BFF3965B8FB4BC16B" blockId="25.[188,1399,455,594]" pageId="25" pageNumber="26">BODY. About 8.1 times as long as greatest width, dorsally pigmented, small cuticular scales on body and part of appendages. Cephalothorax in dorsal view wider than long, a pronounced rounded rostrum between insertion of antennae, a pair of compound eyes with few ommatidia. Body proportions: C&lt;1&gt;2&gt;3&gt;4&gt;5&gt;6&gt;7. Pleonites 1–5 fused, together wider than long, length about as pereonite 6.</paragraph>
<paragraph id="927F36AE21191F0BFF39660BFB07C06A" blockId="25.[189,1400,635,848]" pageId="25" pageNumber="26">ANTENNAE. Antenna 1 peduncle article 1 longest, about 2.3 times as long as wide, articles 2 and 3 progressively narrower, article 2 as long as article 3; flagellum of 2 articles, last article with 3 aesthetascs and 5 setae. Antenna 2 with very short flagellum, basal flagellar article slightly as long as wide. Mandible palp with 1 tiny article and a long apical seta. Maxilla 1 distally with 7 pointed teeth. Maxilliped palp of only 1 article, roughly rectangular, center of apex with concavity, distally with 5 simple setae and 5 medial setae on ventral palp surface, endite short, sub-triangular, apex bearing 1 seta.</paragraph>
<paragraph id="927F36AE21191F0BFF396709FDDCC749" blockId="25.[188,1401,889,1139]" pageId="25" pageNumber="26">PEREOPODS. Pereopod 1 subchelate, basis about 1.3 times as long as greatest width, merus dorsal part trapezoidal, carpus triangular with protruding edge distally, propodus lower half protruding, forming a rounded corner near palm center, with 14 setae in mesial row on distal half of propodus, and 5 longer setae more proximally. Propodus of pereopods 2–6 distally with 1 sensory spine on palm. Pereopods 2–3 not subchelate, merus triangular, dorsal edge reaching nearly to propodus. Pereopod 2 propodus with cuticular scales laterally. Pereopods 4–6 merus nearly triangular, slightly as long as carpus. Pereopod 7 propodus as long as in pereopods 2–6.</paragraph>
<paragraph id="927F36AE21191F0BFF3960EDFAC5C6DB" blockId="25.[188,1400,1179,1505]" pageId="25" pageNumber="26">EXOPODS. Exopod of pleopod 1 operculiform, distal margin with 16 feathered setae, endopod about ½ of exopod width, margin with 9 feathered setae. Exopod of pleopods 2–5 as long as endopod. Pleopods 2–3 exopod distally with 6 feathered setae, endopod pleopod 2 with 6 feathered setae and 9 feathered setae on pleopod 3. Exopod of pleopods 4–5 with 5 feathered setae and endopod with 10 feathered setae, respectively. Uropod sympod elongate, distal margin with 3 feathered setae, endopod subovate, medially as long as laterally, about 0.3 sympod length. Exopod of uropod twice length of endopod and elongate oval, margin with 26 short feathered setae and 7 simple setae. Margins of endopod, exopod and tip of telson weakly crenulated. Telson tongue-shaped, with a pair of statocyst proximally, rounded apex bearing 3 pairs of setae, and on dorsal surface another 2 pairs of longer setae and 1 pair of short setae.</paragraph>
</subSubSection>
<subSubSection id="DADA652521191F0BFF39627AFC6EC4D3" pageId="25" pageNumber="26" type="discussion">
<paragraph id="927F36AE21191F0BFF39627AFE8DC51E" blockId="25.[189,303,1546,1572]" box="[189,303,1546,1572]" pageId="25" pageNumber="26">
<heading id="C93781C221191F0BFF39627AFE8DC51E" bold="true" box="[189,303,1546,1572]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0B4EABC21191F0BFF39627AFE8DC51E" bold="true" box="[189,303,1546,1572]" pageId="25" pageNumber="26">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="927F36AE21191F0BFF39624BFC95C435" blockId="25.[187,1399,1595,1808]" pageId="25" pageNumber="26">
This species belongs to the 
<taxonomicName id="55C04D2D21191F0BFD9A624BFD1BC56F" authorityName="Menzies &amp; Glynn" authorityYear="1968" box="[542,697,1595,1621]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFD9A624BFD1BC56F" box="[542,697,1595,1621]" italics="true" pageId="25" pageNumber="26">Pendanthura</emphasis>
</taxonomicName>
group of genera discussed by 
<bibRefCitation id="F6514B5F21191F0BFBC5624CFB41C56C" author="Poore G. C. B." box="[1089,1251,1595,1622]" pageId="25" pageNumber="26" pagination="63 - 173" refId="ref13754" refString="Poore G. C. B. 2001. Families and genera of Isopoda Anthuridea. Crustacean Issues 13: 63 - 173" type="journal article" year="2001">Poore (2001)</bibRefCitation>
. Within the family 
<taxonomicName id="55C04D2D21191F0BFE956210FE3AC540" authorityName="Leach" authorityYear="1814" box="[273,408,1632,1658]" class="Malacostraca" family="Anthuridae" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="family">Anthuridae</taxonomicName>
, 
<taxonomicName id="55C04D2D21191F0BFE226210FC09C540" authority="Botosaneanu &amp; Stock, 1981" authorityName="Botosaneanu &amp; Stock" authorityYear="1981" box="[422,939,1632,1659]" class="Malacostraca" family="Anthuridae" genus="Stygocyathura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFE226210FDF3C540" box="[422,593,1632,1658]" italics="true" pageId="25" pageNumber="26">Stygocyathura</emphasis>
Botosaneanu &amp; Stock, 1981
</taxonomicName>
, 
<taxonomicName id="55C04D2D21191F0BFC3F6210FAD3C540" authority="Norman &amp; Stebbing, 1886" authorityName="Norman &amp; Stebbing" authorityYear="1886" box="[955,1393,1632,1659]" class="Malacostraca" family="Anthuridae" genus="Cyathura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFC3F6210FB89C540" box="[955,1067,1632,1658]" italics="true" pageId="25" pageNumber="26">Cyathura</emphasis>
Norman &amp; Stebbing, 1886
</taxonomicName>
, 
<taxonomicName id="55C04D2D21191F0BFF3B62F6FD25C5A5" authority="Menzies &amp; Glynn, 1986" authorityName="Menzies &amp; Glynn" authorityYear="1986" box="[191,647,1669,1696]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFF3B62F6FEF8C59A" box="[191,346,1670,1696]" italics="true" pageId="25" pageNumber="26">Pendanthura</emphasis>
Menzies &amp; Glynn, 1986
</taxonomicName>
and 
<taxonomicName id="55C04D2D21191F0BFD4262F6FB23C59A" authority="Poore &amp; Kensley, 1981" authorityName="Poore &amp; Kensley" authorityYear="1981" box="[710,1153,1670,1696]" class="Malacostraca" family="Anthuridae" genus="Sauranthura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFD4262F6FCFEC59A" box="[710,860,1670,1696]" italics="true" pageId="25" pageNumber="26">Sauranthura</emphasis>
Poore &amp; Kensley, 1981
</taxonomicName>
possess a shortened pleon, shortest in 
<taxonomicName id="55C04D2D21191F0BFE0262DBFD83C5FF" authorityName="Menzies &amp; Glynn" authorityYear="1986" box="[390,545,1707,1733]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFE0262DBFD83C5FF" box="[390,545,1707,1733]" italics="true" pageId="25" pageNumber="26">Pendanthura</emphasis>
</taxonomicName>
, and they lack sensory spines distally on the carpus of P4–7. 
<taxonomicName id="55C04D2D21191F0BFB5E62DBFE70C5D0" authority="Menzies &amp; Glynn, 1968" authorityName="Menzies &amp; Glynn" authorityYear="1968" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFB5E62DBFAD7C5FF" box="[1242,1397,1707,1733]" italics="true" pageId="25" pageNumber="26">Pendanthura</emphasis>
Menzies &amp; Glynn, 1968
</taxonomicName>
is characterized by a pronounced rostrum, a maxilliped palp of only 1 rectangular article, and a very short uniarticulate mandibular palp.
</paragraph>
<paragraph id="927F36AE21191F0BFF396349FC6EC4D3" blockId="25.[189,1399,1849,2025]" pageId="25" pageNumber="26">
There are 10 named species that can be assigned to the genus, and there is a description of an unnamed species from 
<collectingRegion id="5004F84C21191F0BFEE5632EFE48C442" box="[353,490,1886,1912]" country="Indonesia" name="Kalimantan Utara" pageId="25" pageNumber="26">Kalimantan</collectingRegion>
(see 
<bibRefCitation id="F6514B5F21191F0BFDAA632FFD4AC443" author="Negoescu I." box="[558,744,1887,1913]" pageId="25" pageNumber="26" pagination="177 - 251" refId="ref13493" refString="Negoescu I. 1997. Isopoda Anthuridea. Results of the Zoological Expedition organized by &quot; Grigore Antipa &quot; Museum in the Indonesia Archipelago (1991). 1. Peracarida (Crustacea). Travaux du Museum national d'histoire naturelle &quot; Grigore Antipa &quot; 38: 177 - 251." type="journal article" year="1997">Negoescu 1997</bibRefCitation>
). Species of 
<taxonomicName id="55C04D2D21191F0BFC0F632EFB84C442" authorityName="Menzies &amp; Glynn" authorityYear="1968" box="[907,1062,1886,1912]" class="Malacostraca" family="Anthuridae" genus="Pendanthura" kingdom="Animalia" order="Isopoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="A0B4EABC21191F0BFC0F632EFB84C442" box="[907,1062,1886,1912]" italics="true" pageId="25" pageNumber="26">Pendanthura</emphasis>
</taxonomicName>
have been found mostly in tropical seas. Most characters useful to distinguish the species can be found in the pereopod 1 propodus palm shape. The palm is generally convex, usually more so in its proximal half. The shape varies somewhat between the sexes, and males have a denser set of setae.
</paragraph>
</subSubSection>
</treatment>
</document>