472 lines
65 KiB
XML
472 lines
65 KiB
XML
<document id="12EF9DE9D651958CFA2DB4333A90D496" ID="10.11646/zootaxa.4213.1.1" ID-CLB-Dataset="36398" ID-DOI="10.11646/zootaxa.4213.1.1" ID-GBIF-Dataset="e68c6438-b109-46ea-a01a-e941191fda8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="253033" ID-ZooBank="9C76B987-3897-4666-87EF-62EB5BF5CF04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484873202414" checkinUser="plazi" docAuthor="Framenau, Volker W. & Baehr, Barbara C." docDate="2016" docId="0B32B23C7B119F70BEF83E29FEDCFA8F" docLanguage="en" docName="zootaxa.4213.1.1.pdf" docOrigin="Zootaxa 4213 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tasmanicosa Roewer 1959" docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="F70BCA447B199F7CBE6F3F45FFE6FFBF" masterDocTitle="Revision of the Australian Union-Jack wolf spiders, genus Tasmanicosa (Araneae, Lycosidae, Lycosinae)" masterLastPageNumber="82" masterPageNumber="1" pageNumber="9" updateTime="1698435957925" updateUser="ExternalLinkService">
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<mods:titleInfo id="C4EFEE57D18830D3E49DBDBF52C413EB">
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<mods:title id="5927BD64EEAF074839E02D227566C68A">Revision of the Australian Union-Jack wolf spiders, genus Tasmanicosa (Araneae, Lycosidae, Lycosinae)</mods:title>
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<mods:namePart id="78F6DAE60C401235F4BD56DD5FC0E074">Framenau, Volker W.</mods:namePart>
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<mods:namePart id="8FAA2039F24E71C51A3491F7282AD99C">Baehr, Barbara C.</mods:namePart>
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<mods:typeOfResource id="3E2D82E551B788CEFFC63A5403299393">text</mods:typeOfResource>
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<mods:title id="39E830DC52F908A00A78D2A1E8C67E41">Zootaxa</mods:title>
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<mods:part id="79803CD5F52185A28854E5BF6A9369AD">
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<mods:date id="DC304C7CE1C282D34722A3A10F796E0D">2016</mods:date>
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<mods:detail id="DA4F5C74CC575A8679F420CAA93B046A" type="volume">
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<mods:number id="763F3081A087D10FB6ED987FB2F5F159">4213</mods:number>
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<mods:identifier id="90E5FAA866AF67C8B4B359F3099C7EE6" type="CLB-Dataset">36398</mods:identifier>
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<mods:identifier id="D8274A57BD03001B75D7493C33B7302D" type="DOI">10.11646/zootaxa.4213.1.1</mods:identifier>
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<treatment id="0B32B23C7B119F70BEF83E29FEDCFA8F" ID-DOI="http://doi.org/10.5281/zenodo.5676921" ID-GBIF-Taxon="127711497" ID-Zenodo-Dep="5676921" LSID="urn:lsid:plazi:treatment:0B32B23C7B119F70BEF83E29FEDCFA8F" httpUri="http://treatment.plazi.org/id/0B32B23C7B119F70BEF83E29FEDCFA8F" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
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<subSubSection id="CB8150A17B119F74BEF83E29FDC2FE18" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8324032A7B119F74BEF83E29FDDFFE39" blockId="8.[151,569,364,423]" box="[151,569,364,390]" pageId="8" pageNumber="9">
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<heading id="D86CB4467B119F74BEF83E29FDDFFE39" bold="true" box="[151,569,364,390]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="B1EFDF387B119F74BEF83E29FDDFFE39" bold="true" box="[151,569,364,390]" pageId="8" pageNumber="9">
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Genus
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<taxonomicName id="449B78A97B119F74BE813E29FDDFFE39" ID-CoL="7SXT" authority="Roewer, 1959" authorityName="Roewer" authorityYear="1959" box="[238,569,364,390]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BE813E29FE6DFE39" bold="true" box="[238,395,364,390]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
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<bibRefCitation id="E70A7EDB7B119F74BFFD3E29FDDFFE39" author="Roewer" box="[402,569,364,390]" pageId="8" pageNumber="9" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II. (Lycosidae). In: Institute des Parcs Nationaux du Congo et du Rwanda (Ed.), Exploration du Parc National de l'Upemba - Mission G. F. de Witte, Vol. 55, pp. 1 - 518." type="journal article" year="1959">Roewer, 1959</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8324032A7B119F74BEF83ECBFDC2FE18" blockId="8.[151,569,364,423]" box="[151,548,398,423]" pageId="8" pageNumber="9">Australian Union-Jack wolf spiders</paragraph>
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</subSubSection>
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<subSubSection id="CB8150A17B119F74BEF83E9DFD9BFDB4" pageId="8" pageNumber="9" type="reference_group">
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<paragraph id="8324032A7B119F74BEF83E9DFE0EFE53" blockId="8.[151,637,470,523]" box="[151,488,470,493]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="A38B24047B119F74BEF83E9DFE0EFE53" box="[151,488,470,493]" pageId="8" pageNumber="9">
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<taxonomicName id="449B78A97B119F74BEF83E9DFE05FE53" ID-CoL="7SXT" authority="Roewer, 1959: 351" authorityName="Roewer" authorityPageNumber="351" authorityYear="1959" box="[151,483,470,493]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BEF83E9DFEFCFE52" box="[151,282,472,493]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
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<treatmentCitation id="023A253B7B119F74BF4F3E92FE05FE53" author="Roewer" box="[288,483,470,492]" page="351" pageId="8" pageNumber="9" year="1959">
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<bibRefCitation id="E70A7EDB7B119F74BF4F3E92FE57FE53" author="Roewer" box="[288,433,470,492]" pageId="8" pageNumber="9" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II. (Lycosidae). In: Institute des Parcs Nationaux du Congo et du Rwanda (Ed.), Exploration du Parc National de l'Upemba - Mission G. F. de Witte, Vol. 55, pp. 1 - 518." type="journal article" year="1959">Roewer, 1959</bibRefCitation>
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: 351
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8324032A7B119F74BEF83EB0FD9BFDB4" blockId="8.[151,637,470,523]" box="[151,637,501,523]" pageId="8" pageNumber="9">
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<treatmentCitationGroup id="A38B24047B119F74BEF83EB0FD9BFDB4" box="[151,637,501,523]" pageId="8" pageNumber="9">
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<taxonomicName id="449B78A97B119F74BEF83EB0FE2DFDB4" authority="Roewer, 1960: 774" authorityName="Roewer" authorityPageNumber="774" authorityYear="1960" box="[151,459,501,523]" class="Arachnida" family="Lycosidae" genus="Orthocosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BEF83EB0FEE4FDB4" box="[151,258,501,523]" italics="true" pageId="8" pageNumber="9">Orthocosa</emphasis>
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<treatmentCitation id="023A253B7B119F74BF673EB3FE2DFDB4" author="Roewer" box="[264,459,501,523]" page="774" pageId="8" pageNumber="9" year="1960">
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<bibRefCitation id="E70A7EDB7B119F74BF673EB3FE7EFDB4" author="Roewer" box="[264,408,501,523]" pageId="8" pageNumber="9" refString="Roewer, C. F. (1960) Araneae Lycosaeformia II. (Lycosidae) (Fortsetzung und Schluss). In: Institute des Parcs Nationaux du Congo et du Rwanda (Ed), Exploration du Parc National de l'Upemba - Mission G. F. de Witte, Vol. 55, pp. 519 - 1040." type="journal article" year="1960">Roewer, 1960</bibRefCitation>
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: 774
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</treatmentCitation>
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||
</taxonomicName>
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||
.
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<emphasis id="B1EFDF387B119F74BFBA3EB0FD9BFDB4" bold="true" box="[469,637,501,523]" pageId="8" pageNumber="9">New synonymy.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="CB8150A17B119F74BEF83D72FC2BFDCA" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8324032A7B119F74BEF83D72FC2BFDCA" blockId="8.[151,1437,567,2033]" pageId="8" pageNumber="9">
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<emphasis id="B1EFDF387B119F74BEF83D72FED2FDEF" bold="true" box="[151,308,567,592]" pageId="8" pageNumber="9">Type species.</emphasis>
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<taxonomicName id="449B78A97B119F74BF513D7CFD7AFDEF" authority="Hogg, 1905" authorityName="Hogg" authorityYear="1905" box="[318,668,567,592]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tasmanica">
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<emphasis id="B1EFDF387B119F74BF513D7CFDEEFDEF" box="[318,520,569,592]" italics="true" pageId="8" pageNumber="9">Lycosa tasmanica</emphasis>
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<bibRefCitation id="E70A7EDB7B119F74BC7D3D7DFD7AFDEF" author="Hogg" box="[530,668,567,592]" pageId="8" pageNumber="9" refString="Hogg, H. R. (1905) On some South Australian spiders of the family Lycosidae. Proceedings of the Zoological Society London, 1905, 569 - 590." type="proceedings paper" year="1905">Hogg, 1905</bibRefCitation>
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</taxonomicName>
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, by original designation (
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<bibRefCitation id="E70A7EDB7B119F74BDAE3D7DFB80FDEF" author="Roewer" box="[961,1126,567,592]" pageId="8" pageNumber="9" refString="Roewer, C. F. (1959) Araneae Lycosaeformia II. (Lycosidae). In: Institute des Parcs Nationaux du Congo et du Rwanda (Ed.), Exploration du Parc National de l'Upemba - Mission G. F. de Witte, Vol. 55, pp. 1 - 518." type="journal article" year="1959">Roewer, 1959</bibRefCitation>
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||
).
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<taxonomicName id="449B78A97B119F74BA143D7CFAA0FDEF" box="[1147,1350,569,592]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tasmanica">
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<emphasis id="B1EFDF387B119F74BA143D7CFAA0FDEF" box="[1147,1350,569,592]" italics="true" pageId="8" pageNumber="9">Lycosa tasmanica</emphasis>
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</taxonomicName>
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is here considered a junior synonym of
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<taxonomicName id="449B78A97B119F74BC6F3D18FC2FFDCA" authority="L. Koch, 1965" authorityName="L. Koch" authorityYear="1965" box="[512,969,604,629]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="godeffroyi">
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<emphasis id="B1EFDF387B119F74BC6F3D18FCEDFDCA" box="[512,779,604,629]" italics="true" pageId="8" pageNumber="9">Tasmanicosa godeffroyi</emphasis>
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(L. Koch, 1965)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CB8150A17B119F74BEA83D3AFE09FA0F" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8324032A7B119F74BEA83D3AFE09FA0F" blockId="8.[151,1437,567,2033]" pageId="8" pageNumber="9">
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<emphasis id="B1EFDF387B119F74BEA83D3AFEA7FD27" bold="true" box="[199,321,639,664]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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The most conspicuous character of
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<taxonomicName id="449B78A97B119F74BC833DC4FC9DFD27" box="[748,891,641,664]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BC833DC4FC9DFD27" box="[748,891,641,664]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
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</taxonomicName>
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is the Union-Jack pattern of the carapace that consists of dark and light radiating lines, including an undulating line under the lateral eyes to the front of the carapace (e.g.
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<figureCitation id="1BA01FAF7B119F74BF503D82FDCAFD5F" box="[319,556,711,736]" captionStart="FIGURE 1" captionStartId="4.[151,250,1789,1811]" captionTargetBox="[216,1374,195,1755]" captionTargetId="figure@4.[212,1374,195,1765]" captionTargetPageId="4" captionText="FIGURE 1 A – H. Life images of Tasmanicosa. A, T. godeffroyi (L. Koch, 1865) comb. nov., male from Canberra (Australian Capital Territory); B – D, T. hughjackmani sp. nov. from Epsom (Victoria); B, female; C, female in burrow, D, male; E – F, T. leuckartii (Thorell, 1870) comb. nov. from Perth (Western Australia); E, female; F, penultimate female; G, T. musgravei (McKay, 1974) comb. nov., female in burrow from Mt Kosciuszko (New South Wales); H, T. semicincta (L. Koch, 1877) comb. nov., female from Brisbane (Queensland)." httpUri="https://zenodo.org/record/253034/files/figure.png" pageId="8" pageNumber="9">Figs 1A–B, D–E, H</figureCitation>
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). Median and marginal light bands are present, but often poorly developed.
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<taxonomicName id="449B78A97B119F74BEF83DA8FF10FCBB" box="[151,246,749,772]" class="Arachnida" family="Lycosidae" genus="Dingosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BEF83DA8FF10FCBB" box="[151,246,749,772]" italics="true" pageId="8" pageNumber="9">Dingosa</emphasis>
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</taxonomicName>
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is the only Australian wolf spider genus in which spiders also have a Union-Jack pattern on the carapace, but males of this genus differ from those of
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<taxonomicName id="449B78A97B119F74BCCA3C54FCD0FC97" box="[677,822,785,808]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BCCA3C54FCD0FC97" box="[677,822,785,808]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
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</taxonomicName>
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by the heavily enlarged palea and elongated tegular apophysis of the male pedipalp (
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<bibRefCitation id="E70A7EDB7B119F74BC6D3C71FCF9FCF3" author="Framenau" box="[514,799,820,845]" pageId="8" pageNumber="9" refString="Framenau, V. W. & Baehr, B. C. (2007) Revision of the Australian wolf spider genus Dingosa Roewer, 1955 (Araneae, Lycosidae, Lycosinae). Journal of Natural History, 41, 1603 - 1629." type="journal article" year="2007">Framenau & Baehr 2007</bibRefCitation>
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). Amongst other morphological characters,
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<taxonomicName id="449B78A97B119F74BB663C70FA8EFCF3" box="[1289,1384,821,844]" class="Arachnida" family="Lycosidae" genus="Dingosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BB663C70FA8EFCF3" box="[1289,1384,821,844]" italics="true" pageId="8" pageNumber="9">Dingosa</emphasis>
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</taxonomicName>
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also differs by the behaviour of constructing turrets around the entrance to the burrow, not observed in
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<taxonomicName id="449B78A97B119F74BB6B3C1CFA73FCCF" box="[1284,1429,857,880]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B1EFDF387B119F74BB6B3C1CFA73FCCF" box="[1284,1429,857,880]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
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||
</taxonomicName>
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||
.
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||
<taxonomicName id="449B78A97B119F74BEF83C38FEC0FC2B" box="[151,294,893,916]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B1EFDF387B119F74BEF83C38FEC0FC2B" box="[151,294,893,916]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
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||
</taxonomicName>
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differs from
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<taxonomicName id="449B78A97B119F74BFD53C38FDE8FC2B" box="[442,526,893,916]" class="Arachnida" family="Lycosidae" genus="Venator" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B1EFDF387B119F74BFD53C38FDE8FC2B" box="[442,526,893,916]" italics="true" pageId="8" pageNumber="9">Venator</emphasis>
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</taxonomicName>
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by the lack of a retrolateral incision in the tegular apophysis of the male pedipalp (absent in
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<taxonomicName id="449B78A97B119F74BF7D3CE4FD9DFC07" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" box="[274,635,927,952]" class="Arachnida" family="Lycosidae" genus="Venator" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="immansuetus">
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<emphasis id="B1EFDF387B119F74BF7D3CE4FE25FC07" box="[274,451,929,952]" italics="true" pageId="8" pageNumber="9">V. immansuetus</emphasis>
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<bibRefCitation id="E70A7EDB7B119F74BFA23CDAFD9DFC07" author="Simon" box="[461,635,927,952]" pageId="8" pageNumber="9" refString="Simon, E. (1909) Araneae, 2 me partie. In: Michaelsen, W & Hartmeyer, R (Eds), Die Fauna Sudwest-Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise 1905. Gustav Fischer Verlag, Jena, pp. 155 - 212." type="journal article" year="1909">(Simon, 1909))</bibRefCitation>
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</taxonomicName>
|
||
and the lack of sharp lateral edges of the epigyne atrium in females (
|
||
<bibRefCitation id="E70A7EDB7B119F74BEF13C81FEB2FC62" author="Framenau" box="[158,340,964,989]" pageId="8" pageNumber="9" refString="Framenau, V. W. (2015) Review of the Australian wolf spider genus Venator (Araneae, Lycosidae). Zootaxa, 4013, 541 - 555. https: // doi. org / 10.11646 / zootaxa. 4013.4.5" type="journal article" year="2015">Framenau 2015</bibRefCitation>
|
||
). It differs from
|
||
<taxonomicName id="449B78A97B119F74BC653C81FD87FC62" box="[522,609,964,989]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BC653C81FD87FC62" box="[522,609,964,989]" italics="true" pageId="8" pageNumber="9">Knoelle</emphasis>
|
||
</taxonomicName>
|
||
by the lack of a large patch of apical setae on the male pedipalp cymbium (
|
||
<bibRefCitation id="E70A7EDB7B119F74BEF13CADFE84FBBF" author="Framenau" box="[158,354,999,1024]" pageId="8" pageNumber="9" refString="Framenau, V. W. (2006 a) Knoelle, a new monotypic wolf spider genus from Australia (Araneae: Lycosidae). Zootaxa, 1281, 55 - 67." type="journal article" year="2006" yearSuffix="a">Framenau 2006a</bibRefCitation>
|
||
).
|
||
<taxonomicName id="449B78A97B119F74BF1A3CACFDE0FBBF" box="[373,518,1001,1024]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BF1A3CACFDE0FBBF" box="[373,518,1001,1024]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
(with the exception of
|
||
<taxonomicName id="449B78A97B119F74BD7F3CACFC47FBBF" box="[784,929,1001,1024]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="musgravei">
|
||
<emphasis id="B1EFDF387B119F74BD7F3CACFC47FBBF" box="[784,929,1001,1024]" italics="true" pageId="8" pageNumber="9">T. musgravei</emphasis>
|
||
</taxonomicName>
|
||
) differs from
|
||
<taxonomicName id="449B78A97B119F74BA2D3CACFB5DFBBF" box="[1090,1211,1001,1024]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BA2D3CACFB5DFBBF" box="[1090,1211,1001,1024]" italics="true" pageId="8" pageNumber="9">Hoggicosa</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the epigyne, as the distance between the anterior pockets is smaller than the width of the posterior transverse part (
|
||
<bibRefCitation id="E70A7EDB7B119F74BEF13B6AFE0DFBF7" author="Langlands" box="[158,491,1071,1096]" pageId="8" pageNumber="9" refString="Langlands, P. & Framenau, V. W. (2010) Systematic revision of Hoggicosa Roewer, 1960, the Australian ' bicolor ' group of wolf spiders (Araneae: Lycosidae). Zoological Journal of the Linnean Society, 158, 83 - 123. https: // doi. org / 10.1111 / j. 1096 - 3642.2009.00545. x" type="journal article" year="2010">Langlands & Framenau 2010</bibRefCitation>
|
||
). In addition, males of
|
||
<taxonomicName id="449B78A97B119F74BC863B74FC85FBF7" box="[745,867,1073,1096]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BC863B74FC85FBF7" box="[745,867,1073,1096]" italics="true" pageId="8" pageNumber="9">Hoggicosa</emphasis>
|
||
</taxonomicName>
|
||
have a patch of dorsally bent setae on the tip of the cymbium, which is lacking in
|
||
<taxonomicName id="449B78A97B119F74BF983B10FD6EFBD3" box="[503,648,1109,1132]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BF983B10FD6EFBD3" box="[503,648,1109,1132]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="449B78A97B119F74BCF73B10FCCFFBD3" box="[664,809,1109,1132]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BCF73B10FCCFFBD3" box="[664,809,1109,1132]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
distinctly differs from the monotypic
|
||
<taxonomicName id="449B78A97B119F74BA8B3B10FAA0FBD3" box="[1252,1350,1109,1132]" class="Arachnida" family="Lycosidae" genus="Tapetosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BA8B3B10FAA0FBD3" box="[1252,1350,1109,1132]" italics="true" pageId="8" pageNumber="9">Tapetosa</emphasis>
|
||
</taxonomicName>
|
||
, as the spiders are not dorso-ventrally flattened and the legs are not laterigrade (
|
||
<bibRefCitation id="E70A7EDB7B119F74BD9C3B3DFB12FB2F" author="Framenau" box="[1011,1268,1143,1168]" pageId="8" pageNumber="9" refString="Framenau, V. W., Main, B. Y., Waldock, J. M. & Harvey, M. S. (2009) Tapetosa, a new monotypic wolf spider genus from Western Australia (Araneae: Lycosidae: Lycosinae). Records of the Western Australian Museum, 25, 309 - 314. https: // doi. org / 10.18195 / issn. 0312 - 3162.25 (3). 2009.309 - 314" type="journal article" year="2009">
|
||
Framenau
|
||
<emphasis id="B1EFDF387B119F74BA003B3CFB4AFB2F" box="[1135,1196,1143,1168]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="449B78A97B119F74BB643B3CFA7CFB2F" box="[1291,1434,1145,1168]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BB643B3CFA7CFB2F" box="[1291,1434,1145,1168]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="449B78A97B119F74BF483BD8FE6FFB0B" box="[295,393,1181,1204]" class="Arachnida" family="Lycosidae" genus="Mainosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BF483BD8FE6FFB0B" box="[295,393,1181,1204]" italics="true" pageId="8" pageNumber="9">Mainosa</emphasis>
|
||
</taxonomicName>
|
||
by a very different body colouration, i.e. the opisthosoma is not black with transverse light bars on the opisthosoma, and by the lack of turret-building behaviour around its burrow entrance (
|
||
<bibRefCitation id="E70A7EDB7B119F74BB453B85FF04FB42" author="Framenau" pageId="8" pageNumber="9" refString="Framenau, V. W. (2006 b) Mainosa, a new genus for the Australian ' shuttlecock wolf spider' (Araneae, Lycosidae). Journal of Arachnology, 34, 206 - 213." type="journal article" year="2006" yearSuffix="b">Framenau 2006b</bibRefCitation>
|
||
).
|
||
<taxonomicName id="449B78A97B119F74BE9A3BA0FE62FB43" box="[245,388,1253,1276]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BE9A3BA0FE62FB43" box="[245,388,1253,1276]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="449B78A97B119F74BC713BA0FD8DFB43" box="[542,619,1253,1276]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BC713BA0FD8DFB43" box="[542,619,1253,1276]" italics="true" pageId="8" pageNumber="9">Hogna</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the male and female genitalia as
|
||
<emphasis id="B1EFDF387B119F74BAF13BA0FB15FB43" box="[1182,1267,1253,1276]" italics="true" pageId="8" pageNumber="9">
|
||
<taxonomicName id="449B78A97B119F74BAF13BA0FB0BFB43" box="[1182,1261,1253,1276]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Hogna</taxonomicName>
|
||
¸
|
||
</emphasis>
|
||
as represented by its
|
||
<typeStatus id="5C20BD887B119F74BE8A3A4CFEF3FA9E" box="[229,277,1289,1313]" pageId="8" pageNumber="9">type</typeStatus>
|
||
species
|
||
<taxonomicName id="449B78A97B119F74BF113A4CFD24FA9F" authority="Latreille, 1817" authorityName="Latreille" authorityYear="1817" box="[382,706,1287,1312]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="radiata">
|
||
<emphasis id="B1EFDF387B119F74BF113A4CFE1EFA9F" box="[382,504,1287,1312]" italics="true" pageId="8" pageNumber="9">H. radiata</emphasis>
|
||
(Latreille, 1817)
|
||
</taxonomicName>
|
||
, have a very simple, triangular tegular apophysis in the male pedipalp and a simple, inverted T-shaped median septum of the female epigyne (Framenau
|
||
<emphasis id="B1EFDF387B119F74BACC3A68FB3BFAFA" box="[1187,1245,1324,1349]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
|
||
2006), and from
|
||
<taxonomicName id="449B78A97B119F74BEF83A15FEEAFAD8" box="[151,268,1360,1383]" class="Arachnida" family="Lycosidae" genus="Costacosa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BEF83A15FEEAFAD8" box="[151,268,1360,1383]" italics="true" pageId="8" pageNumber="9">Costacosa</emphasis>
|
||
</taxonomicName>
|
||
in the very different shape of the tegular apophysis (
|
||
<bibRefCitation id="E70A7EDB7B119F74BD163A15FB45FAD7" author="Framenau" box="[889,1187,1360,1385]" pageId="8" pageNumber="9" refString="Framenau, V. W. & Leung, A. E. (2013) Costacosa, a new genus of wolf spider (Araneae, Lycosidae) from coastal north-west Western Australia. Records of the Western Australian Museum Supplement, 83, 173 - 184. https: // doi. org / 10.18195 / issn. 0313 - 122 x. 83.2013.173 - 184" type="journal article" year="2013">Framenau & Leung 2013</bibRefCitation>
|
||
).
|
||
<taxonomicName id="449B78A97B119F74BAD73A14FAA1FAD7" box="[1208,1351,1361,1384]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BAD73A14FAA1FAD7" box="[1208,1351,1361,1384]" italics="true" pageId="8" pageNumber="9">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="449B78A97B119F74BEBA3A30FED4FA33" box="[213,306,1397,1420]" class="Arachnida" family="Lycosidae" genus="Venatrix" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BEBA3A30FED4FA33" box="[213,306,1397,1420]" italics="true" pageId="8" pageNumber="9">Venatrix</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="449B78A97B119F74BF043A31FE12FA32" box="[363,500,1396,1421]" class="Arachnida" family="Lycosidae" genus="Tuberculosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B119F74BF043A31FE12FA32" box="[363,500,1396,1421]" italics="true" pageId="8" pageNumber="9">Tuberculosa</emphasis>
|
||
</taxonomicName>
|
||
by the lack of an apical cymbial hook-like macroseta in males (
|
||
<bibRefCitation id="E70A7EDB7B119F74BAA13A31FF35FA0F" author="Framenau" pageId="8" pageNumber="9" refString="Framenau, V. W. & Vink, C. J. (2001) Revision of the wolf spider genus Venatrix Roewer (Araneae: Lycosidae). Invertebrate Taxonomy, 15, 927 - 970." type="journal article" year="2001">Framenau & Vink 2001</bibRefCitation>
|
||
; Framenau & Yoo 2006).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CB8150A17B119F70BEA83AF9FA9DFD4F" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="8324032A7B119F74BEA83AF9FDA1F8A7" blockId="8.[151,1437,567,2033]" pageId="8" pageNumber="9">
|
||
<emphasis id="B1EFDF387B119F74BEA83AF9FEBFFA6A" bold="true" box="[199,345,1468,1493]" pageId="8" pageNumber="9">Description.</emphasis>
|
||
Large and robust wolf spiders, total length ca. 12.0–25.0 in males and ca. 14.0–30.0 in females. Prosoma longer than wide with rounded postero-lateral corners. Prominent Union-Jack pattern dorsally, consisting of black and light radiating lines; the most anterior line reaches under the eyes and approaches the front of the carapace. The row of the anterior eyes is slightly procurved, the PME are larger than PLE; the row of the AE slightly narrower than the row of PME; the row of PME is narrower than that of the PLE (
|
||
<figureCitation id="1BA01FAF7B119F74BAFD3909FB0BF9DB" box="[1170,1261,1612,1637]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
|
||
). The labium is subquadrate, i.e. about as long as wide, and the endites about twice as long as the labium; both generally dark brown to black, sometimes with lighter anterior rim. The chelicerae are dark brown to black with an elongated patch of white, silvery, yellow or golden setae (
|
||
<figureCitation id="1BA01FAF7B119F74BCD539F2FCA7F96F" box="[698,833,1719,1744]" captionStart="FIGURE 1" captionStartId="4.[151,250,1789,1811]" captionTargetBox="[216,1374,195,1755]" captionTargetId="figure@4.[212,1374,195,1765]" captionTargetPageId="4" captionText="FIGURE 1 A – H. Life images of Tasmanicosa. A, T. godeffroyi (L. Koch, 1865) comb. nov., male from Canberra (Australian Capital Territory); B – D, T. hughjackmani sp. nov. from Epsom (Victoria); B, female; C, female in burrow, D, male; E – F, T. leuckartii (Thorell, 1870) comb. nov. from Perth (Western Australia); E, female; F, penultimate female; G, T. musgravei (McKay, 1974) comb. nov., female in burrow from Mt Kosciuszko (New South Wales); H, T. semicincta (L. Koch, 1877) comb. nov., female from Brisbane (Queensland)." httpUri="https://zenodo.org/record/253034/files/figure.png" pageId="8" pageNumber="9">Figs. 1D, F</figureCitation>
|
||
; 2A). They have three promarginal teeth, the middle largest and its base is often fused with a smaller apical one, the proximal one is more distant. They also have three large retromarginal teeth of equal size.
|
||
</paragraph>
|
||
<paragraph id="8324032A7B119F74BEA83861FE01F84E" blockId="8.[151,1437,567,2033]" pageId="8" pageNumber="9">
|
||
The leg formula of males and females is IV> I> II> III. Legs are uniform greyish-brown with the coxal venter of the same colour as the sternum, generally dark brown to black, with the exception of
|
||
<taxonomicName id="449B78A97B119F74BAE0380CFAC5F8DF" box="[1167,1315,1865,1888]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="semicincta">
|
||
<emphasis id="B1EFDF387B119F74BAE0380CFAC5F8DF" box="[1167,1315,1865,1888]" italics="true" pageId="8" pageNumber="9">T. semicincta</emphasis>
|
||
</taxonomicName>
|
||
with light brown sternum and ventral coxae. The femora are generally the lightest, ventral patellae and apical tibiae are often darker than the remainder of the legs. Legs I and II have entire ventral scopulae on tarsi and metatarsi, which also cover parts of the tibiae in females. Legs III and IV have scopulae on tarsi and half of metatarsi in males and all of tarsi and metatarsi in females.
|
||
</paragraph>
|
||
<caption id="D7E453A27B109F75BEF83833FB5CF857" httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="9" pageNumber="10" targetBox="[295,1315,209,1885]" targetPageId="9">
|
||
<paragraph id="8324032A7B109F75BEF83833FB5CF857" blockId="9.[151,1437,1910,2024]" pageId="9" pageNumber="10">
|
||
<emphasis id="B1EFDF387B109F75BEF83833FED9F833" bold="true" box="[151,319,1910,1932]" pageId="9" pageNumber="10">FIGURE 2A–F</emphasis>
|
||
. Key morphological terminology for the revision of
|
||
<taxonomicName id="449B78A97B109F75BDE8383DFBECF832" box="[903,1034,1912,1933]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B109F75BDE8383DFBECF832" box="[903,1034,1912,1933]" italics="true" pageId="9" pageNumber="10">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
, exemplified by the type species,
|
||
<taxonomicName id="449B78A97B109F75BBE6383DFE48F814" authority="L. Koch, 1865" authorityName="L. Koch" authorityYear="1865" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="godeffroyi" status="comb. nov.">
|
||
<emphasis id="B1EFDF387B109F75BBE6383DFF18F814" italics="true" pageId="9" pageNumber="10">T. godeffroyi</emphasis>
|
||
(L. Koch, 1865)
|
||
</taxonomicName>
|
||
<emphasis id="B1EFDF387B109F75BFD838D0FDC8F814" bold="true" box="[439,558,1941,1963]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="AADC62437B109F75BFD838D0FDC8F814" box="[439,558,1941,1963]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; A, Eye arrangement in frontal view;
|
||
<emphasis id="B1EFDF387B109F75BDAE38D0FC34F814" bold="true" box="[961,978,1941,1963]" pageId="9" pageNumber="10">B</emphasis>
|
||
, male right pedipalp, ventral view;
|
||
<emphasis id="B1EFDF387B109F75BB2438D0FABBF814" bold="true" box="[1355,1373,1941,1963]" pageId="9" pageNumber="10">C</emphasis>
|
||
, right tegular apophysis, ventral view,
|
||
<emphasis id="B1EFDF387B109F75BF8638F1FE1DF875" bold="true" box="[489,507,1972,1994]" pageId="9" pageNumber="10">D</emphasis>
|
||
, right palea, ventral view;
|
||
<emphasis id="B1EFDF387B109F75BD7638F1FCCCF875" bold="true" box="[793,810,1972,1994]" pageId="9" pageNumber="10">E</emphasis>
|
||
, female epigyne, ventral view;
|
||
<emphasis id="B1EFDF387B109F75BA1738F1FB63F875" bold="true" box="[1144,1157,1972,1994]" pageId="9" pageNumber="10">F</emphasis>
|
||
, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D7E453A27B139F76BEF83884FF22F849" httpUri="https://zenodo.org/record/253036/files/figure.png" pageId="10" pageNumber="11" targetBox="[210,1379,203,1953]" targetPageId="10">
|
||
<paragraph id="8324032A7B139F76BEF83884FF22F849" blockId="10.[151,1435,1985,2038]" pageId="10" pageNumber="11">
|
||
<emphasis id="B1EFDF387B139F76BEF83884FEA7F868" bold="true" box="[151,321,1985,2007]" pageId="10" pageNumber="11">FIGURE 3A–T.</emphasis>
|
||
Illustrations for the key to the
|
||
<taxonomicName id="449B78A97B139F76BC133887FCE6F868" box="[636,768,1986,2007]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B139F76BC133887FCE6F868" box="[636,768,1986,2007]" italics="true" pageId="10" pageNumber="11">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
species of Australia. Roman letters refer to the key choices in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D7E453A27B129F77BEF838E2FF22F863" httpUri="https://zenodo.org/record/253037/files/figure.png" pageId="11" pageNumber="12" targetBox="[192,1397,195,1921]" targetPageId="11">
|
||
<paragraph id="8324032A7B129F77BEF838E2FF22F863" blockId="11.[151,1435,1959,2012]" pageId="11" pageNumber="12">
|
||
<emphasis id="B1EFDF387B129F77BEF838E2FEA3F802" bold="true" box="[151,325,1959,1981]" pageId="11" pageNumber="12">FIGURE 4A–K.</emphasis>
|
||
Illustrations for the key to the
|
||
<taxonomicName id="449B78A97B129F77BC1138ECFCE4F801" box="[638,770,1961,1982]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B129F77BC1138ECFCE4F801" box="[638,770,1961,1982]" italics="true" pageId="11" pageNumber="12">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
species of Australia. Roman letters refer to the key choices in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8324032A7B159F70BEA83FD2FCF3FEA3" blockId="12.[151,1437,151,1328]" pageId="12" pageNumber="13">
|
||
The sternum is generally dark brown to black, except in
|
||
<emphasis id="B1EFDF387B159F70BD163FDCFBFEFF0F" box="[889,1048,153,176]" italics="true" pageId="12" pageNumber="13">
|
||
<taxonomicName id="449B78A97B159F70BD163FDCFBF3FF0F" box="[889,1045,153,176]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="semicincta">T. semicincta</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
in which it is light brown. The opisthosoma has a dorsal, dark folium pattern that is laterally bordered by light lines or patches (
|
||
<figureCitation id="1BA01FAF7B159F70BAB83FF9FF49FF47" captionStart="FIGURE 1" captionStartId="4.[151,250,1789,1811]" captionTargetBox="[216,1374,195,1755]" captionTargetId="figure@4.[212,1374,195,1765]" captionTargetPageId="4" captionText="FIGURE 1 A – H. Life images of Tasmanicosa. A, T. godeffroyi (L. Koch, 1865) comb. nov., male from Canberra (Australian Capital Territory); B – D, T. hughjackmani sp. nov. from Epsom (Victoria); B, female; C, female in burrow, D, male; E – F, T. leuckartii (Thorell, 1870) comb. nov. from Perth (Western Australia); E, female; F, penultimate female; G, T. musgravei (McKay, 1974) comb. nov., female in burrow from Mt Kosciuszko (New South Wales); H, T. semicincta (L. Koch, 1877) comb. nov., female from Brisbane (Queensland)." httpUri="https://zenodo.org/record/253034/files/figure.png" pageId="12" pageNumber="13">Figs 1A–B, D–E, H</figureCitation>
|
||
). The venter is either uniformly dark brown to black, but this patch is often reduced in some species displaying a species-specific pattern (e.g.
|
||
<taxonomicName id="449B78A97B159F70BFB33E40FD85FEA2" box="[476,611,260,285]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="leuckartii">
|
||
<emphasis id="B1EFDF387B159F70BFB33E40FD85FEA2" box="[476,611,260,285]" italics="true" pageId="12" pageNumber="13">T. leuckartii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="449B78A97B159F70BC1E3E40FCE3FEA3" box="[625,773,261,284]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="semicincta">
|
||
<emphasis id="B1EFDF387B159F70BC1E3E40FCE3FEA3" box="[625,773,261,284]" italics="true" pageId="12" pageNumber="13">T. semicincta</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8324032A7B159F70BEA83E62FC5EFDDF" blockId="12.[151,1437,151,1328]" pageId="12" pageNumber="13">
|
||
The male pedipalp cymbium is dorsally covered with a dense layer of silvery setae. The tip of the cymbium carries ca. 2–10 macrosetae that are slightly bent ventrally. The tegular apophysis is directed retrolaterally (
|
||
<figureCitation id="1BA01FAF7B159F70BB503E09FA72FEDB" box="[1343,1428,332,357]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2B</figureCitation>
|
||
) and has a ridge of species-specific shape between its apical point and its ventral process (e.g.
|
||
<figureCitation id="1BA01FAF7B159F70BAA23E2AFACFFE37" box="[1229,1321,367,392]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation id="1BA01FAF7B159F70BB583E2AFABCFE37" box="[1335,1370,367,392]" captionStart="FIGURE 6" captionStartId="16.[151,250,1920,1942]" captionTargetBox="[220,1366,813,1892]" captionTargetId="figure@16.[213,1374,805,1899]" captionTargetPageId="16" captionText="FIGURE 6 A – H. Scanning Electron Micrographs (SEM) of right male Tasmanicosa tegular apophyses, ventral view. A, T. godeffroyi (L. Koch) (NMV K 11547); B, T. kochorum sp. nov. (QM S 68828); C, T. gilberta (Hogg) (QM S 70800); D, T. leuckartii (Thorell) (NMV K 11541); E, T. phyllis (Hogg) (NMV 11539); F, T. musgravei (McKay) (QM S 70787); G, T. semicincta (L. Koch) (QM S 71191); H, T. ramosa (L. Koch) (QM S 71125). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/253039/files/figure.png" pageId="12" pageNumber="13">6B</figureCitation>
|
||
). The ventral process is well-developed. The length of this ridge corresponds to the length of the median septum in females (see
|
||
<bibRefCitation id="E70A7EDB7B159F70BF5E3EF2FE2DFE6F" author="Zyuzin" box="[305,459,439,464]" pageId="12" pageNumber="13" refString="Zyuzin, A. A. (1993) Studies on the wolf spiders (Araneae: Lycosidae). I. A new genus and species from Kazakhstan, with comments on the Lycosinae. Memoirs of the Queensland Museum, 33, 693 - 700." type="journal article" year="1993">Zyuzin, 1993</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="E70A7EDB7B159F70BC653EF2FD43FE6F" author="Sadana" box="[522,677,439,464]" pageId="12" pageNumber="13" refString="Sadana, G. L. (1972) Mechanics of copulation in Lycosa chaperi Simon (Araneida: Lycosidae). Bulletin of the British Arachnological Society, 2, 87 - 89" type="journal article" year="1972">Sadana, 1972</bibRefCitation>
|
||
for a functional explanation of male and female genitalia in the Lycosinae). The embolus is long and generally sickle-shaped as characteristic for most Lycosinae (
|
||
<figureCitation id="1BA01FAF7B159F70BB6F3E99FABAFE4B" box="[1280,1372,476,501]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2D</figureCitation>
|
||
). The terminal apophysis is generally broad and sickle-shaped (
|
||
<figureCitation id="1BA01FAF7B159F70BD443EBAFC63FDA7" box="[811,901,511,536]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2D</figureCitation>
|
||
) but apically modified in a number of species aiding in their identification (e.g.
|
||
<taxonomicName id="449B78A97B159F70BC5C3D60FD25FD82" box="[563,707,548,573]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="leuckartii">
|
||
<emphasis id="B1EFDF387B159F70BC5C3D60FD25FD82" box="[563,707,548,573]" italics="true" pageId="12" pageNumber="13">L. leuckartii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="449B78A97B159F70BCB73D60FC97FD83" box="[728,881,549,572]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="musgravei">
|
||
<emphasis id="B1EFDF387B159F70BCB73D60FC97FD83" box="[728,881,549,572]" italics="true" pageId="12" pageNumber="13">L. musgravei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="449B78A97B159F70BDE93D60FBE2FD82" box="[902,1028,548,573]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gilberta">
|
||
<emphasis id="B1EFDF387B159F70BDE93D60FBE2FD82" box="[902,1028,548,573]" italics="true" pageId="12" pageNumber="13">L. gilberta</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="1BA01FAF7B159F70BA4C3D61FB43FD83" box="[1059,1189,548,573]" captionStart="FIGURE 7" captionStartId="17.[151,250,1293,1315]" captionTargetBox="[214,1370,198,1263]" captionTargetId="figure@17.[206,1381,193,1271]" captionTargetPageId="17" captionText="FIGURE 7 A – H. Scanning Electron Micrographs (SEM) of right Tasmanicosa palea, ventral view. A, T. ramosa (L. Koch) (QM S 71125) B, T. kochorum sp. nov. (QM S 68828); C, T. semicincta (L. Koch) (QM S 71191); D, T. phyllis (Hogg) (NMV 11539); E, T. godeffroyi (L. Koch) (NMV K 11547); F, T. musgravei (McKay) (QM S 70787); G, T. leuckartii (Thorell) (NMV K 11541); H, T. gilberta (Hogg) (QM S 70800); Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/253040/files/figure.png" pageId="12" pageNumber="13">Figs 7F–H</figureCitation>
|
||
,
|
||
<figureCitation id="1BA01FAF7B159F70BAD83D61FB11FD83" box="[1207,1271,548,572]" captionStart="FIGURE 11" captionStartId="23.[151,250,1617,1639]" captionTargetBox="[254,1350,186,1586]" captionTargetId="figure@23.[248,1357,179,1594]" captionTargetPageId="23" captionText="FIGURE 11 A – L. Tasmanicosa gilberta (Hogg) comb. nov., male and female (QM S 70800); A, C, male, dorsal (A) and ventral (C) view; B, D, female, dorsal (B) and ventral (D) view; E, F, right male pedipalp, retrolateral (E) and ventral (F) view; G, H, female epigyne, ventral (G) and dorsal (H) view; I, J, palea region of right male pedipalp, ventral (I) and retrolateral (J) view (SEM); K, L, tegular apophysis of right male pedipalp in (K) apical and (L) ventral view;. Scale bars: A – D, 5 mm; E, F, 1 mm; G – I, K, L 0.1 mm." httpUri="https://zenodo.org/record/253044/files/figure.png" pageId="12" pageNumber="13">
|
||
<date id="F72525EA7B159F70BAD83D61FB3DFD83" box="[1207,1243,548,572]" pageId="12" pageNumber="13" value="2011-01">11I</date>
|
||
–J
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="1BA01FAF7B159F70BB653D61FAD7FD82" box="[1290,1329,548,573]" captionStart="FIGURE 15" captionStartId="29.[151,250,1565,1587]" captionTargetBox="[269,1316,200,1530]" captionTargetId="figure@29.[263,1324,193,1544]" captionTargetPageId="29" captionText="FIGURE 15 A – K. Tasmanicosa kochorum sp. nov., male (QM S 68828) and female (QM S 68829); A, C, male, dorsal (A) and ventral (C) view; B, D, female, dorsal (B) and ventral (D) view; E, F, right male pedipalp, retrolateral (E) and ventral (F) view; G, H, female epigyne, ventral (G) and dorsal (H) view; I, palea of right pedipalp, ventral view; J, K, tegular apophysis of right pedipalp in (J) apical and (K) ventral view. Scale bars: A – D, 5 mm; E, F, 1 mm; G – K, 0.1 mm." httpUri="https://zenodo.org/record/253049/files/figure.png" pageId="12" pageNumber="13">
|
||
<date id="F72525EA7B159F70BB653D61FAD7FD82" box="[1290,1329,548,573]" pageId="12" pageNumber="13" value="2015-01">15I</date>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="1BA01FAF7B159F70BB2B3D61FA8BFD83" box="[1348,1389,548,572]" captionStart="FIGURE 18" captionStartId="34.[151,250,1549,1571]" captionTargetBox="[283,1308,201,1500]" captionTargetId="figure@34.[275,1313,195,1525]" captionTargetPageId="34" captionText="FIGURE 18 A – K. Tasmanicosa musgravei (McKay) comb. nov., male (QM S 70787) and female (QM S 70785); A, C, male, dorsal (A) and ventral (C) view; B, D, female, dorsal (B) and ventral (D) view; E, F, right male pedipalp, retrolateral (E) and ventral (F) view; G, H, female epigyne, ventral (G) and dorsal (H) view; I, palea of right pedipalp, ventral view; J, K, tegular apophysis of right pedipalp in (J) ventral and (K) apical view. Scale bars: A – D, 5 mm; E, F, 1 mm; G – K, 0.1 mm." httpUri="https://zenodo.org/record/253052/files/figure.png" pageId="12" pageNumber="13">
|
||
<date id="F72525EA7B159F70BB2B3D61FA8BFD83" box="[1348,1389,548,572]" pageId="12" pageNumber="13" value="1918-01">18I</date>
|
||
</figureCitation>
|
||
). A subterminal apophysis is present and generally sickle-shaped (
|
||
<figureCitation id="1BA01FAF7B159F70BD3E3D02FC4AFDDF" box="[849,940,583,608]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8324032A7B159F70BEA83D29FA9DFD4F" blockId="12.[151,1437,151,1328]" pageId="12" pageNumber="13">
|
||
The median septum of the female epigyne is inverted T-shaped, i.e. with a wide posterior transverse part (
|
||
<figureCitation id="1BA01FAF7B159F70BB003D29FF51FD17" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2E</figureCitation>
|
||
) and within this general configuration is variable between species, in particular in reference to it widening anteriorly. The spermathecal heads (
|
||
<figureCitation id="1BA01FAF7B159F70BC5F3DF1FD63FD73" box="[560,645,692,717]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2F</figureCitation>
|
||
) are of varying shape (i.e. spherical, elongated or kidney-shaped) and generally wider than the spermathecal stalks (
|
||
<figureCitation id="1BA01FAF7B159F70BCF83D92FD0BFD4F" box="[663,749,727,752]" captionStart="FIGURE 2" captionStartId="9.[151,250,1910,1932]" captionTargetBox="[295,1315,209,1885]" captionTargetId="figure@9.[268,1318,193,1889]" captionTargetPageId="9" captionText="FIGURE 2 A – F. Key morphological terminology for the revision of Tasmanicosa, exemplified by the type species, T. godeffroyi (L. Koch, 1865) comb. nov.; A, Eye arrangement in frontal view; B, male right pedipalp, ventral view; C, right tegular apophysis, ventral view, D, right palea, ventral view; E, female epigyne, ventral view; F, female epigyne (cleared), dorsal view. AME, ALE, anterior median and lateral eyes; PME, PLE, posterior median and lateral eyes." httpUri="https://zenodo.org/record/253035/files/figure.png" pageId="12" pageNumber="13">Fig. 2F</figureCitation>
|
||
). The latter are generally twisted, S-shaped or convoluted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CB8150A17B159F70BEA83DB9FEDCFA8F" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8324032A7B159F70BEA83DB9FD0BFB1F" blockId="12.[151,1437,151,1328]" pageId="12" pageNumber="13">
|
||
<emphasis id="B1EFDF387B159F70BEA83DB9FED2FCAA" bold="true" box="[199,308,764,789]" pageId="12" pageNumber="13">Remarks</emphasis>
|
||
. Roewer’s (1959: 351) justification of the genus
|
||
<taxonomicName id="449B78A97B159F70BD383DB8FC0EFCAB" box="[855,1000,765,788]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BD383DB8FC0EFCAB" box="[855,1000,765,788]" italics="true" pageId="12" pageNumber="13">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
was insufficient by modern taxonomic standards and did not include an accurate description of the genus. The diagnosis was limited to a single character referring to the distance between the PME (from German): “
|
||
<taxonomicName id="449B78A97B159F70BD243C00FB42FCE2" authority="Hogg, 1905" authorityName="Hogg" authorityYear="1905" box="[843,1188,836,861]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tasmanica">
|
||
<emphasis id="B1EFDF387B159F70BD243C00FBF3FCE3" box="[843,1045,837,860]" italics="true" pageId="12" pageNumber="13">Lycosa tasmanica</emphasis>
|
||
<bibRefCitation id="E70A7EDB7B159F70BA713C00FB42FCE2" author="Hogg" box="[1054,1188,836,861]" pageId="12" pageNumber="13" refString="Hogg, H. R. (1905) On some South Australian spiders of the family Lycosidae. Proceedings of the Zoological Society London, 1905, 569 - 590." type="proceedings paper" year="1905">Hogg, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
(in the catalogue, 2a, 240, sub
|
||
<taxonomicName id="449B78A97B159F70BE913C2CFEBBFC3F" box="[254,349,873,896]" class="Arachnida" family="Lycosidae" genus="Dingosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BE913C2CFEBBFC3F" box="[254,349,873,896]" italics="true" pageId="12" pageNumber="13">Dingosa</emphasis>
|
||
</taxonomicName>
|
||
) is better assigned to a new genus (for which we here propose
|
||
<taxonomicName id="449B78A97B159F70BA4A3C2CFB52FC3F" box="[1061,1204,873,896]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BA4A3C2CFB52FC3F" box="[1061,1204,873,896]" italics="true" pageId="12" pageNumber="13">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
n. g.), which differs from
|
||
<taxonomicName id="449B78A97B159F70BEBA3CC9FEA6FC1A" box="[213,320,908,933]" class="Arachnida" family="Lycosidae" genus="Lycosella" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BEBA3CC9FEA6FC1A" box="[213,320,908,933]" italics="true" pageId="12" pageNumber="13">Lycosella</emphasis>
|
||
</taxonomicName>
|
||
(see the following genus) in the distance between the PME which is smaller than one diameter of the PME.” The problems associated with this poor diagnosis and much of Roewer’s taxonomic work on the
|
||
<taxonomicName id="449B78A97B159F70BEF83C91FEEFFC52" box="[151,265,980,1005]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
|
||
are reflected in the fact that
|
||
<taxonomicName id="449B78A97B159F70BC0E3C91FC94FC53" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" box="[609,882,980,1005]" class="Arachnida" family="Lycosidae" genus="Lycosella" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BC0E3C91FD2DFC52" box="[609,715,980,1005]" italics="true" pageId="12" pageNumber="13">Lycosella</emphasis>
|
||
Thorell, 1890
|
||
</taxonomicName>
|
||
belongs to a different subfamily altogether, the
|
||
<taxonomicName id="449B78A97B159F70BEF83CB2FE21FBAF" authority="Framenau, 2007" authorityName="Framenau" authorityYear="2007" box="[151,455,1015,1040]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="subFamily" subFamily="Artoriinae">Artoriinae Framenau, 2007</taxonomicName>
|
||
(Framenau 2007). The genus
|
||
<taxonomicName id="449B78A97B159F70BD783CBCFC40FBAF" box="[791,934,1017,1040]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BD783CBCFC40FBAF" box="[791,934,1017,1040]" italics="true" pageId="12" pageNumber="13">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
is here included in the subfamily Lycosinae, as it conforms to the subfamily diagnosis given by
|
||
<bibRefCitation id="E70A7EDB7B159F70BCB93B59FC6FFB8A" author="Dondale" box="[726,905,1052,1077]" pageId="12" pageNumber="13" refString="Dondale, C. D. (1986) The subfamilies of wolf spiders (Araneae: Lycosidae). Actas X Congreso Internacional de Aracnologia, Jaca, Espana, pp. 327 - 332." type="book chapter" year="1986">Dondale (1986)</bibRefCitation>
|
||
, i.e. the tegular apophysis of the male pedipalp is transverse with a ventrally directed spur and has a sinuous channel on the dorsal surface. Within the Lycosinae, molecular sequence data places the genus in a Gondwanan clade, together with
|
||
<taxonomicName id="449B78A97B159F70BA2F3B20FB5FFBC3" box="[1088,1209,1125,1148]" class="Arachnida" family="Lycosidae" genus="Hoggicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BA2F3B20FB5FFBC3" box="[1088,1209,1125,1148]" italics="true" pageId="12" pageNumber="13">Hoggicosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="449B78A97B159F70BAA33B21FAC2FBC2" box="[1228,1316,1124,1149]" class="Arachnida" family="Lycosidae" genus="Knoelle" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BAA33B21FAC2FBC2" box="[1228,1316,1124,1149]" italics="true" pageId="12" pageNumber="13">Knoelle</emphasis>
|
||
</taxonomicName>
|
||
and some
|
||
<taxonomicName id="449B78A97B159F70BEF83BCCFF05FB1F" box="[151,227,1161,1184]" class="Arachnida" family="Lycosidae" genus="Hogna" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BEF83BCCFF05FB1F" box="[151,227,1161,1184]" italics="true" pageId="12" pageNumber="13">Hogna</emphasis>
|
||
</taxonomicName>
|
||
, albeit with low support (
|
||
<bibRefCitation id="E70A7EDB7B159F70BC6D3BC2FD07FB1F" author="Murphy" box="[514,737,1159,1184]" pageId="12" pageNumber="13" refString="Murphy, N. P., Framenau, V. W., Donnellan, S. C., Harvey, M. S., Park, Y. - C. & Austin, A. D. (2006) Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12 S rRNA, 28 S rRNA, and NADH 1 genes: implications for classification, biogeography, and the evolution of web building behaviour. Molecular Phylogenetics and Evolution, 38, 583 - 602." type="journal article" year="2006">
|
||
Murphy
|
||
<emphasis id="B1EFDF387B159F70BC0A3BCCFD78FB1F" box="[613,670,1159,1184]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8324032A7B159F70BEA83BE9FEDCFA8F" blockId="12.[151,1437,151,1328]" pageId="12" pageNumber="13">
|
||
The transfer of
|
||
<taxonomicName id="449B78A97B159F70BF163BE9FCD4FB7B" authority="L. Koch, 1877" authorityName="L. Koch" authorityYear="1877" box="[377,818,1196,1221]" class="Arachnida" family="Lycosidae" genus="Orthocosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="semicincta">
|
||
<emphasis id="B1EFDF387B159F70BF163BE9FD96FB7B" box="[377,624,1196,1221]" italics="true" pageId="12" pageNumber="13">Orthocosa semicincta</emphasis>
|
||
(
|
||
<bibRefCitation id="E70A7EDB7B159F70BCEE3BE9FCCDFB7B" author="Koch" box="[641,811,1196,1221]" pageId="12" pageNumber="13" refString="Koch, L. (1877) Die Arachniden Australiens nach der Natur beschrieben und abgebildet. 1. Theil. 7. Lieferung. Verlag von Bauer und Raspe, Nurnberg, pp. 889 - 968." type="journal article" year="1877">L. Koch, 1877</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, the type species of
|
||
<taxonomicName id="449B78A97B159F70BA723BE9FB75FB7A" box="[1053,1171,1196,1221]" class="Arachnida" family="Lycosidae" genus="Orthocosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BA723BE9FB75FB7A" box="[1053,1171,1196,1221]" italics="true" pageId="12" pageNumber="13">Orthocosa</emphasis>
|
||
</taxonomicName>
|
||
, places
|
||
<taxonomicName id="449B78A97B159F70BA9E3BE9FA81FB7A" box="[1265,1383,1196,1221]" class="Arachnida" family="Lycosidae" genus="Orthocosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BA9E3BE9FA81FB7A" box="[1265,1383,1196,1221]" italics="true" pageId="12" pageNumber="13">Orthocosa</emphasis>
|
||
</taxonomicName>
|
||
into junior synonymy of
|
||
<taxonomicName id="449B78A97B159F70BFF03B94FDD6FB57" box="[415,560,1233,1256]" class="Arachnida" family="Lycosidae" genus="Tasmanicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BFF03B94FDD6FB57" box="[415,560,1233,1256]" italics="true" pageId="12" pageNumber="13">Tasmanicosa</emphasis>
|
||
</taxonomicName>
|
||
. Other non-Australian species currently listed in
|
||
<taxonomicName id="449B78A97B159F70BAC23B8AFAC5FB57" box="[1197,1315,1231,1256]" class="Arachnida" family="Lycosidae" genus="Orthocosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1EFDF387B159F70BAC23B8AFAC5FB57" box="[1197,1315,1231,1256]" italics="true" pageId="12" pageNumber="13">Orthocosa</emphasis>
|
||
</taxonomicName>
|
||
are here morphologically assessed based on their original descriptions and are here transferred to more appropriate genera (Appendix A).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |