535 lines
114 KiB
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535 lines
114 KiB
XML
<document id="AE37981063A2047C5DF8594153EDFA49" ID-CLB-Dataset="274285" ID-DOI="10.5852/ejt.2023.893.2285" ID-GBIF-Dataset="e441bc9d-bb44-4ee6-b29f-a6724b96592b" ID-ISSN="2118-9773" ID-Zenodo-Dep="8385630" ID-ZooBank="D9DA4879-BC1D-4947-96E8-7D574DE334BA" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1695891225180" checkinUser="plazi" docAuthor="Leng, Zhaoqi, Tran, Anh Duc & Ye, Zhen" docDate="2023" docId="045E878DFFC529263A79FD0972C90CE2" docLanguage="en" docName="EJT.2023.893.1-73.pdf" docOrigin="European Journal of Taxonomy 893 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2285/9863" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhyacobates turgidus Leng & Tran 2023, sp. nov." docType="treatment" docUuid="A61745D8-0A50-4E9C-89CA-B557C637ED2E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="30" masterDocId="F867FFF5FFDC293B381FFFB270530F1A" masterDocTitle="Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="26" updateTime="1699690649759" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E8FFA294610DCEF32EE8C92671B7038D">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="0E93426739B7FF6B31E8CEBF9F5A43A2">Leng, Zhaoqi</mods:namePart>
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<mods:affiliation id="2BC323216E79CBD6DBA3852227E41D1C">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:nameIdentifier id="B04FF2B328B9ED8EE6C1284247E70E82" type="email">zhaoqileng0712@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E3A4CA0C805D2DB62F1A4E1F83C12DA7">Tran, Anh Duc</mods:namePart>
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<mods:affiliation id="61A8C906C00550D7639687FE252C2D02">Faculty of Biology, VNU University of Science, Vietnam National University, Hanoi, Nguyen Trai, Thanh Xuan, Hanoi, Vietnam.</mods:affiliation>
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<mods:nameIdentifier id="A63C1537E566038BC6A004B3CFA9F820" type="email">tran.anhduc@hus.edu.vn</mods:nameIdentifier>
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<mods:namePart id="DB44C295EC6B5B7C320A0367932C6759">Ye, Zhen</mods:namePart>
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<mods:affiliation id="2ABF0EF1E6FAA8E5070CAAB0EC5C44D0">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:nameIdentifier id="255B3266B164E991EE1FE89EEB7253C8" type="email">yezhen1987331@nankai.edu.cn</mods:nameIdentifier>
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<mods:title id="9DA64BB7AA5073E624C95C0909A937E3">European Journal of Taxonomy</mods:title>
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<mods:date id="8222249C76177546588F74983C34A36C">2023</mods:date>
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<mods:number id="CFE4666D7D47537D93FD8D4CF47C88B3">2023-09-28</mods:number>
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<treatment id="045E878DFFC529263A79FD0972C90CE2" ID-DOI="http://doi.org/10.5281/zenodo.10064063" ID-GBIF-Taxon="214246599" ID-Zenodo-Dep="10064063" LSID="urn:lsid:zoobank.org:act:A61745D8-0A50-4E9C-89CA-B557C637ED2E" httpUri="http://treatment.plazi.org/id/045E878DFFC529263A79FD0972C90CE2" lastPageId="29" lastPageNumber="30" pageId="25" pageNumber="26">
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<subSubSection id="C4ED6510FFC529223A79FD09739D0DCC" box="[614,974,699,726]" pageId="25" pageNumber="26" type="nomenclature">
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<heading id="D70081F7FFC529223A79FD09739D0DCC" box="[614,974,699,726]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
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<taxonomicName id="4BF74D18FFC529223A79FD09733F0DCC" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[614,876,699,726]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
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<emphasis id="BE83EA89FFC529223A79FD09733F0DCC" bold="true" box="[614,876,699,726]" italics="true" pageId="25" pageNumber="26">Rhyacobates turgidus</emphasis>
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<taxonomicNameLabel id="A5B057F2FFC529223B6CFD0E739D0DCC" box="[883,974,700,726]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C4ED6510FFC529223A53FCB673B40C04" box="[588,999,772,799]" pageId="25" pageNumber="26" type="description">
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<paragraph id="8C48369BFFC529223A53FCB673B40C04" blockId="25.[362,1226,699,799]" box="[588,999,772,799]" pageId="25" pageNumber="26">
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<figureCitation id="14CC2A1EFFC529223A53FCB6729D0C04" box="[588,718,772,798]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="25" pageNumber="26">Figs 3C–D</figureCitation>
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,
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<figureCitation id="14CC2A1EFFC529223AC4FCB672AC0C04" box="[731,767,772,798]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="25" pageNumber="26">4C</figureCitation>
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,
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<figureCitation id="14CC2A1EFFC529223B14FCB673630C05" box="[779,816,772,799]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="25" pageNumber="26">5C</figureCitation>
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,
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<figureCitation id="14CC2A1EFFC529223B23FCB673320C04" box="[828,865,772,798]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="25" pageNumber="26">6C</figureCitation>
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<figureCitation id="14CC2A1EFFC529223B72FCB673C10C04" box="[877,914,772,798]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="25" pageNumber="26">8C</figureCitation>
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,
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<figureCitation id="14CC2A1EFFC529223B81FCB673B40C04" box="[926,999,772,798]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="27.[189,232,1963,1989]" captionStartId-1="28.[189,232,1787,1813]" captionTargetBox-0="[260,1340,536,1853]" captionTargetBox-1="[264,1323,291,1729]" captionTargetId-0="figure-53@27.[189,1398,511,1911]" captionTargetId-1="figure-15@28.[228,1359,265,1747]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." captionText-1="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385676" figureDoi-1="http://doi.org/10.5281/zenodo.8385678" httpUri-0="https://zenodo.org/record/8385676/files/figure.png" httpUri-1="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">19–20</figureCitation>
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</paragraph>
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<subSubSection id="C4ED6510FFC5292238A2FCF471EB0A5A" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph id="8C48369BFFC5292238A2FCF471650C7A" blockId="25.[189,310,838,864]" box="[189,310,838,864]" pageId="25" pageNumber="26">
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<heading id="D70081F7FFC5292238A2FCF471650C7A" bold="true" box="[189,310,838,864]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
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<emphasis id="BE83EA89FFC5292238A2FCF471650C7A" bold="true" box="[189,310,838,864]" pageId="25" pageNumber="26">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8C48369BFFC5292238A2FCC771EB0A5A" blockId="25.[189,1401,885,1344]" pageId="25" pageNumber="26">
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Body length of apterous females 7.82–8.50, of apterous males 5.90–6.31. In both sexes, abdominal mediotergite I distinctly swollen and large, about as long as the three subsequent abdominal terga together (
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<figureCitation id="14CC2A1EFFC52922392BFC0F71EC0CC2" box="[308,447,957,984]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="25" pageNumber="26">Figs 3C–D</figureCitation>
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,
|
||
<figureCitation id="14CC2A1EFFC5292239CFFC0C71A50CC2" box="[464,502,958,984]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="25" pageNumber="26">4C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC529223A18FC0C72D90CC2" box="[519,650,958,984]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">20A–B, H</figureCitation>
|
||
). Female: pronotum mainly black with a median brownish-yellow spot; mesonotum and metanotum mainly black with a median brownish-yellow stripe (
|
||
<figureCitation id="14CC2A1EFFC5292238DAFBB771780B3A" box="[197,299,1029,1056]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="25" pageNumber="26">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC529223924FBB471230B3A" box="[315,368,1030,1056]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[260,1340,536,1853]" captionTargetId="figure-53@27.[189,1398,511,1911]" captionTargetPageId="27" captionText="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="25" pageNumber="26">19A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC52922399EFBB471EA0B3A" box="[385,441,1030,1056]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">20A</figureCitation>
|
||
); abdominal mediotergite I with a median brownish-yellow stripe; abdominal terga II–VI distinctly short (
|
||
<figureCitation id="14CC2A1EFFC529223A12FB9B72C70B5E" box="[525,660,1065,1092]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="25" pageNumber="26">Figs 3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC529223ABFFB9872A80B5E" box="[672,763,1066,1092]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">20A–B</figureCitation>
|
||
); abdominal mediotergite II medially hidden beneath the abdominal mediotergite I but not laterally (
|
||
<figureCitation id="14CC2A1EFFC529223B13FBFF73210B72" box="[780,882,1101,1128]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="25" pageNumber="26">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC529223B9CFBFC73E80B72" box="[899,955,1102,1128]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">20A</figureCitation>
|
||
); laterosternites of connexivum VII expanded dorsad, folded mesad, but not meeting in middle of abdominal dorsum; posterior margin of sternum VII with a short and pointed median process (
|
||
<figureCitation id="14CC2A1EFFC529223B6EFB2774570BAA" box="[881,1028,1173,1200]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">Fig. 20E–G</figureCitation>
|
||
). Male: middle trochanter with one spine; middle femur with scattered small spines, not arranged in distinct row (
|
||
<figureCitation id="14CC2A1EFFC529223C8EFB0B74AB0BCE" box="[1169,1272,1209,1236]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">Fig. 20J</figureCitation>
|
||
); length of middle tibia ca 1.4 times length of hind tibia; proctiger with angular lobes laterally (
|
||
<figureCitation id="14CC2A1EFFC529223CA0FB6F75700BED" box="[1215,1315,1245,1271]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="25" pageNumber="26">Figs 6C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC529223D2EFB6C753A0BE2" box="[1329,1385,1246,1272]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">20K</figureCitation>
|
||
); paramere relatively slender, strongly curved at basal third, distal part tapering towards hook-shaped apex (
|
||
<figureCitation id="14CC2A1EFFC529223918FA97713A0A5A" box="[263,361,1317,1344]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="25" pageNumber="26">Figs 8C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC529223969FA9471F80A5A" box="[374,427,1318,1344]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="25" pageNumber="26">20L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFC5292238A2FADA72F80ACC" pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph id="8C48369BFFC5292238A2FADA71170A98" blockId="25.[189,324,1384,1410]" box="[189,324,1384,1410]" pageId="25" pageNumber="26">
|
||
<heading id="D70081F7FFC5292238A2FADA71170A98" bold="true" box="[189,324,1384,1410]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="BE83EA89FFC5292238A2FADA71170A98" bold="true" box="[189,324,1384,1410]" pageId="25" pageNumber="26">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC5292238A2FA2572F80ACC" blockId="25.[189,1399,1431,1494]" pageId="25" pageNumber="26">The species epithet is derived from the Latin adjective ʻturgidusʼ, meaning ʻswollenʼ and refers to the extremely large abdominal mediotergite I.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFC5292238A2FA4F73FE089F" pageId="25" pageNumber="26" type="materials_examined">
|
||
<paragraph id="8C48369BFFC5292238A2FA4F71F9090D" blockId="25.[189,426,1533,1559]" box="[189,426,1533,1559]" pageId="25" pageNumber="26">
|
||
<heading id="D70081F7FFC5292238A2FA4F71F9090D" bold="true" box="[189,426,1533,1559]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
|
||
<emphasis id="BE83EA89FFC5292238A2FA4F71F9090D" bold="true" box="[189,426,1533,1559]" pageId="25" pageNumber="26">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC5292238F5F99F71B3095D" blockId="25.[189,1399,1581,1679]" box="[234,480,1581,1607]" pageId="25" pageNumber="26">
|
||
<emphasis id="BE83EA89FFC5292238F5F99F710F095D" bold="true" box="[234,348,1581,1607]" pageId="25" pageNumber="26">
|
||
<typeStatus id="534C8839FFC5292238F5F99F710F095D" box="[234,348,1581,1607]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="14CC2A1EFFC529223973F99F7184095D" box="[364,471,1581,1607]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[260,1340,536,1853]" captionTargetId="figure-53@27.[189,1398,511,1911]" captionTargetPageId="27" captionText="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="25" pageNumber="26">Fig. 19A</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC5292238A2F9E374730995" blockId="25.[189,1399,1581,1679]" pageId="25" pageNumber="26">
|
||
<materialsCitation id="3C9F3CC6FFC5292238A2F9E374730995" ID-GBIF-Occurrence="4429869336" collectingDate="2016-08-09" collectionCode="NKUM" collectorName="Chen-guang Zheng" country="China" county="He-jiang County" elevation="741" latitude="28.584333" location="Lu-zhou City" longLatPrecision="1" longitude="106.25264" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan" typeStatus="holotype">
|
||
<collectingCountry id="F4E0760BFFC5292238A2F9E3714B0971" box="[189,280,1617,1643]" name="China" pageId="25" pageNumber="26">CHINA</collectingCountry>
|
||
•
|
||
<specimenCount id="9AF1FD12FFC529223932F9E371100971" box="[301,323,1617,1643]" pageId="25" pageNumber="26" type="female">♀</specimenCount>
|
||
(apterous);
|
||
<collectingRegion id="4E33F879FFC5292239D3F9E372C80971" box="[460,667,1617,1643]" country="China" name="Sichuan" pageId="25" pageNumber="26">Sichuan Province</collectingRegion>
|
||
,
|
||
<location id="89286040FFC529223ABBF9E373120971" LSID="urn:lsid:plazi:treatment:045E878DFFC529263A79FD0972C90CE2:89286040FFC529223ABBF9E373120971" box="[676,833,1617,1643]" country="China" county="He-jiang County" latitude="106.25264" longLatPrecision="1" longitude="28.584333" name="Lu-zhou City" pageId="25" pageNumber="26" stateProvince="Sichuan">Lu-zhou City</location>
|
||
,
|
||
<collectingCounty id="65294E17FFC529223B55F9E3745D0971" box="[842,1038,1617,1643]" pageId="25" pageNumber="26">He-jiang County</collectingCounty>
|
||
, Tian-tang-ba;
|
||
<geoCoordinate id="E9C3505CFFC529223CA5F9FD75210971" box="[1210,1394,1615,1643]" degrees="106" direction="east" minutes="15" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="09.5" value="106.25264">106°15′09.5″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFC5292238A2F9C171040995" box="[189,343,1651,1679]" degrees="28" direction="north" minutes="35" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="3.6" value="28.584333">28°35′3.6″ E</geoCoordinate>
|
||
;
|
||
<elevation id="07DAD1A8FFC52922397CF9C771BD0995" box="[355,494,1653,1679]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="25" pageNumber="26" unit="m" value="741.0">
|
||
<quantity id="4B0F9B7EFFC52922397CF9C771FD0995" box="[355,430,1653,1679]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="25" pageNumber="26" unit="m" value="741.0">741 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="F849105BFFC5292239E4F9C772DC0995" box="[507,655,1653,1679]" pageId="25" pageNumber="26" value="2016-08-09">
|
||
<collectingDate id="E80DE9B3FFC5292239E4F9C772DC0995" box="[507,655,1653,1679]" pageId="25" pageNumber="26" value="2016-08-09">9 Aug. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFC529223A84F9C7732F0995" box="[667,892,1653,1679]" pageId="25" pageNumber="26">Chen-guang Zheng</collectorName>
|
||
leg.;
|
||
<collectionCode id="EAE6AE5EFFC529223BA1F9C7744F0995" box="[958,1052,1653,1679]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="25" pageNumber="26">NKUM</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC5292238F5F90571BA09CB" blockId="25.[189,1398,1719,1926]" box="[234,489,1719,1745]" pageId="25" pageNumber="26">
|
||
<emphasis id="BE83EA89FFC5292238F5F905713409CB" bold="true" box="[234,359,1719,1745]" pageId="25" pageNumber="26">
|
||
<typeStatus id="534C8839FFC5292238F5F905713409CB" box="[234,359,1719,1745]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="14CC2A1EFFC529223967F90571B309CB" box="[376,480,1719,1745]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[260,1340,536,1853]" captionTargetId="figure-53@27.[189,1398,511,1911]" captionTargetPageId="27" captionText="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="25" pageNumber="26">Fig. 19B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC5292238A2F96973FE089F" blockId="25.[189,1398,1719,1926]" pageId="25" pageNumber="26">
|
||
<materialsCitation id="3C9F3CC6FFC5292238A2F96971440803" ID-GBIF-Occurrence="4429869388" collectingDate="2016-08-09" collectionCode="NKUM" collectorName="Chen-guang Zheng" country="China" county="He-jiang County" elevation="741" latitude="28.584333" location="Lu-zhou City" longLatPrecision="1" longitude="106.25264" pageId="25" pageNumber="26" specimenCount="3" specimenCount-female="3" stateProvince="Sichuan" typeStatus="paratype">
|
||
<collectingCountry id="F4E0760BFFC5292238A2F969714B09EF" box="[189,280,1755,1781]" name="China" pageId="25" pageNumber="26">CHINA</collectingCountry>
|
||
• 3 ♁♁,
|
||
<specimenCount id="9AF1FD12FFC529223997F969719809EF" box="[392,459,1755,1781]" pageId="25" pageNumber="26" type="female">3 ♀♀</specimenCount>
|
||
(apterous),
|
||
<specimenCount id="9AF1FD12FFC529223A44F96972DA09EF" box="[603,649,1755,1781]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
|
||
(dealated macropterous); same collection data as for holotype;
|
||
<collectionCode id="EAE6AE5EFFC5292238A2F94D71440803" box="[189,279,1791,1817]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="25" pageNumber="26">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFC529223921F94D73620827" ID-GBIF-Occurrence="4429869358" collectingDate="2016-08-10" collectionCode="NKUM" collectorName="Yan-fei Li" country="China" elevation="895" latitude="28.608444" location="Si-mian Mountain" longLatPrecision="1" longitude="106.372" municipality="City" pageId="25" pageNumber="26" specimenCount="3" specimenCount-female="3" stateProvince="Chongqing" typeStatus="paratype">
|
||
<specimenCount id="9AF1FD12FFC529223921F94D71D40803" box="[318,391,1791,1817]" pageId="25" pageNumber="26" type="female">3 ♀♀</specimenCount>
|
||
(apterous);
|
||
<collectingRegion id="4E33F879FFC529223A3DF94D72F70803" box="[546,676,1791,1817]" country="China" name="Chongqing" pageId="25" pageNumber="26">Chongqing</collectingRegion>
|
||
<collectingMunicipality id="6C2CACE1FFC529223AACF94D72B40803" box="[691,743,1791,1817]" pageId="25" pageNumber="26">City</collectingMunicipality>
|
||
,
|
||
<location id="89286040FFC529223AE5F94D73880803" LSID="urn:lsid:plazi:treatment:045E878DFFC529263A79FD0972C90CE2:89286040FFC529223AE5F94D73880803" box="[762,987,1791,1817]" country="China" latitude="28.608444" longLatPrecision="1" longitude="106.372" municipality="City" name="Si-mian Mountain" pageId="25" pageNumber="26" stateProvince="Chongqing">Si-mian Mountain</location>
|
||
;
|
||
<geoCoordinate id="E9C3505CFFC529223BF2F94F74F30803" box="[1005,1184,1789,1817]" degrees="28" direction="north" minutes="36" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="30.4" value="28.608444">28°36′30.4″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFC529223CACF94F75220803" box="[1203,1393,1789,1817]" degrees="106" direction="east" minutes="22" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="19.2" value="106.372">106°22′19.2″ E</geoCoordinate>
|
||
;
|
||
<elevation id="07DAD1A8FFC5292238A2F891711E0827" box="[189,333,1827,1854]" metricMagnitude="2" metricUnit="m" metricValue="8.95" pageId="25" pageNumber="26" unit="m" value="895.0">
|
||
<quantity id="4B0F9B7EFFC5292238A2F89171590827" box="[189,266,1827,1854]" metricMagnitude="2" metricUnit="m" metricValue="8.95" pageId="25" pageNumber="26" unit="m" value="895.0">895 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="F849105BFFC529223941F89172550827" box="[350,518,1827,1853]" pageId="25" pageNumber="26" value="2016-08-10">
|
||
<collectingDate id="E80DE9B3FFC529223941F89172550827" box="[350,518,1827,1853]" pageId="25" pageNumber="26" value="2016-08-10">10 Aug. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFC529223A0BF89172DC0827" box="[532,655,1827,1853]" pageId="25" pageNumber="26">Yan-fei Li</collectorName>
|
||
leg.;
|
||
<collectionCode id="EAE6AE5EFFC529223AC8F89173620827" box="[727,817,1827,1853]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="25" pageNumber="26">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFC529223B4FF89173FA089F" ID-GBIF-Occurrence="4429869361" collectingDate="2013-08-15" collectionCode="NKUM" collectorName="Yang Liu & Zhen Ye" country="China" county="Xu-yong County" elevation="741" latitude="28.272055" location="National Nature Reserve" longLatPrecision="1" longitude="105.541" municipality="Lu-zhou City" pageId="25" pageNumber="26" specimenCount="4" specimenCount-female="4" stateProvince="Sichuan" typeStatus="paratype">
|
||
1 ♁,
|
||
<specimenCount id="9AF1FD12FFC529223B8FF89173860827" box="[912,981,1827,1853]" pageId="25" pageNumber="26" type="female">4 ♀♀</specimenCount>
|
||
(apterous);
|
||
<collectingRegion id="4E33F879FFC529223C76F891756E0827" box="[1129,1341,1827,1853]" country="China" name="Sichuan" pageId="25" pageNumber="26">Sichuan Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6C2CACE1FFC529223D53F8917165087B" pageId="25" pageNumber="26">Lu-zhou City</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="65294E17FFC529223959F8F57240087B" box="[326,531,1863,1889]" pageId="25" pageNumber="26">Xu-yong County</collectingCounty>
|
||
, Hua-gao-xi
|
||
<location id="89286040FFC529223AABF8F573B0087B" LSID="urn:lsid:plazi:treatment:045E878DFFC529263A79FD0972C90CE2:89286040FFC529223AABF8F573B0087B" box="[692,995,1863,1889]" country="China" county="Xu-yong County" latitude="28.272055" longLatPrecision="1" longitude="105.541" municipality="Lu-zhou City" name="National Nature Reserve" pageId="25" pageNumber="26" stateProvince="Sichuan">National Nature Reserve</location>
|
||
;
|
||
<geoCoordinate id="E9C3505CFFC529223BEBF8F774F6087B" box="[1012,1189,1861,1889]" degrees="28" direction="north" minutes="16" orientation="latitude" pageId="25" pageNumber="26" precision="1" seconds="19.4" value="28.272055">28°16′19.4″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFC529223CAAF8F77522087B" box="[1205,1393,1861,1889]" degrees="105" direction="east" minutes="32" orientation="longitude" pageId="25" pageNumber="26" precision="1" seconds="27.6" value="105.541">105°32′27.6″ E</geoCoordinate>
|
||
;
|
||
<elevation id="07DAD1A8FFC5292238A2F8D97114089F" box="[189,327,1899,1925]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="25" pageNumber="26" unit="m" value="741.0">
|
||
<quantity id="4B0F9B7EFFC5292238A2F8D97154089F" box="[189,263,1899,1925]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="25" pageNumber="26" unit="m" value="741.0">741 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="F849105BFFC52922394AF8D971A4089F" box="[341,503,1899,1926]" pageId="25" pageNumber="26" value="2013-08-15">
|
||
<collectingDate id="E80DE9B3FFC52922394AF8D971A4089F" box="[341,503,1899,1926]" pageId="25" pageNumber="26" value="2013-08-15">15 Aug. 2013</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFC529223A1DF8D9723C089F" box="[514,623,1899,1925]" pageId="25" pageNumber="26">Yang Liu</collectorName>
|
||
and
|
||
<collectorName id="2102534DFFC529223AB7F8D9735A089F" box="[680,777,1899,1925]" pageId="25" pageNumber="26">Zhen Ye</collectorName>
|
||
leg.;
|
||
<collectionCode id="EAE6AE5EFFC529223B54F8D973FA089F" box="[843,937,1899,1925]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="25" pageNumber="26">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFC6292638A2FEBA731D0ECB" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27" type="description">
|
||
<paragraph id="8C48369BFFC6292138A2FEBA711D0E38" blockId="26.[189,334,264,290]" box="[189,334,264,290]" pageId="26" pageNumber="27">
|
||
<heading id="D70081F7FFC6292138A2FEBA711D0E38" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
|
||
<emphasis id="BE83EA89FFC6292138A2FEBA711D0E38" bold="true" box="[189,334,264,290]" pageId="26" pageNumber="27">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138F5FE8A71EB0E48" blockId="26.[185,1400,312,585]" box="[234,440,312,338]" pageId="26" pageNumber="27">
|
||
<emphasis id="BE83EA89FFC6292138F5FE8A71EB0E48" bold="true" box="[234,440,312,338]" pageId="26" pageNumber="27">Apterous female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138A2FEEE73020D52" blockId="26.[185,1400,312,585]" pageId="26" pageNumber="27">
|
||
<smallCapsWord id="8AAEA047FFC6292138A2FEEE713D0E6E" baselines="367,367" box="[189,366,348,374]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="26" pageNumber="27">MEASUREMENTS</smallCapsWord>
|
||
. Body length 7.82–8.50 (
|
||
<typeStatus id="534C8839FFC629213ABEFEE973590E6F" box="[673,778,347,373]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
8.20), width 3.01–3.60 (
|
||
<typeStatus id="534C8839FFC629213C2CFEE974CF0E6F" box="[1075,1180,347,373]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
3.48), head width 1.48, interocular width 0.71, eye length (dorsal view) 0.43; relative lengths of antennal segments I– IV: 3.89:1.02: 1.18: 0.92; pronotum: length 0.81, width 1.65; mesonotum: length 3.78, width 3.25; metanotum: length 0.84, width 2.84; abdomen length (ventral view) 2.94; abdominal sternum VII: length 1.56, width 1.05; abdominal mediotergite I: length 0.86, width 1.71; relative lengths of leg segments (femur: tibia: tarsal segment I:tarsal segment II): fore leg: 3.94:3.38: 1.93: 0.88, middle leg: 11.12: 6.88: 4.25:0.45, hind leg: 11.35: 5.41:0.16: 0.24.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138A6FDC2727F0C0C" blockId="26.[185,1399,624,791]" pageId="26" pageNumber="27">
|
||
C OLORATION. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum and metanotum mainly black with a median brownish-yellow stripe (
|
||
<figureCitation id="14CC2A1EFFC6292138DAFD0471760DCA" box="[197,293,694,720]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="26" pageNumber="27">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC62921392DFD0471340DCA" box="[306,359,694,720]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[260,1340,536,1853]" captionTargetId="figure-53@27.[189,1398,511,1911]" captionTargetPageId="27" captionText="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="26" pageNumber="27">19A</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC62921396BFD0471F80DCA" box="[372,427,694,720]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20A</figureCitation>
|
||
). Connexivum dorsally brownish-yellow. Abdominal mediotergite I with a median yellow stripe. Mesosternum chiefly blackish with a median subtriangular yellowish spot (
|
||
<figureCitation id="14CC2A1EFFC629213CE4FD6B75390DE9" box="[1275,1386,729,755]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">Fig. 20C</figureCitation>
|
||
). Abdominal venter light yellow.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138A2FC8C74340B8F" blockId="26.[187,1401,830,1173]" pageId="26" pageNumber="27">
|
||
<smallCapsWord id="8AAEA047FFC6292138A2FC8C717F0C4C" baselines="849,849" box="[189,300,830,856]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="26" pageNumber="27">ABDOMEN</smallCapsWord>
|
||
. Abdomen relatively short, almost straight (
|
||
<figureCitation id="14CC2A1EFFC629213B58FC8C73F50C43" box="[839,934,830,857]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="26" pageNumber="27">Fig. 5C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213BA9FC8C73BE0C42" box="[950,1005,830,856]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20D</figureCitation>
|
||
). Connexiva erect on abdominal segments I–VI, parallel to each other above mediotergites, reflexed over terminal mediotergites (
|
||
<figureCitation id="14CC2A1EFFC6292138D9FC3671070C84" box="[198,340,900,926]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">Fig. 20A–B</figureCitation>
|
||
).Abdominal mediotergite I distinctly swollen, nearly as long as three subsequent abdominal terga together, sparsely covered with silvery pubescence on each side; abdominal terga II–VI distinctly short in length (
|
||
<figureCitation id="14CC2A1EFFC629213961FC7972560CFF" box="[382,517,971,997]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="26" pageNumber="27">Figs 3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213A0BFC79723E0CFF" box="[532,621,971,997]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20A–B</figureCitation>
|
||
). Abdominal mediotergite II medially hidden beneath abdominal mediotergite I, but laterally exposed (
|
||
<figureCitation id="14CC2A1EFFC629213A9BFC5C73580B12" box="[644,779,1006,1032]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="26" pageNumber="27">Figs 3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213B04FC5C73010B12" box="[795,850,1006,1032]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20A</figureCitation>
|
||
). Abdominal segment VII elongate, nearly as long as two preceding abdominal segments together, length of ventral margin distinctly longer than that of dorsal margin in lateral view (
|
||
<figureCitation id="14CC2A1EFFC629213A54FB8672FE0B55" box="[587,685,1076,1103]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="26" pageNumber="27">Figs 5C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213AA3FB8672A00B54" box="[700,755,1076,1102]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20D</figureCitation>
|
||
); laterosternites of connexivum VII expanded dorsad, folded mesad, not meeting in middle of abdominal dorsum (
|
||
<figureCitation id="14CC2A1EFFC629213B62FBE573B40B68" box="[893,999,1111,1138]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">Fig. 20E</figureCitation>
|
||
). Abdominal sternum VII tapering caudad, posterior margin with a short and pointed median process (
|
||
<figureCitation id="14CC2A1EFFC629213BCFFBC974090B8F" box="[976,1114,1147,1173]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">Fig. 20E–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138F5FB0E723B0BCC" blockId="26.[188,1398,1212,1344]" box="[234,616,1212,1238]" pageId="26" pageNumber="27">
|
||
<emphasis id="BE83EA89FFC6292138F5FB0E723B0BCC" bold="true" box="[234,616,1212,1238]" pageId="26" pageNumber="27">Dealated macropterous female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138A2FB6D74160A5A" blockId="26.[188,1398,1212,1344]" pageId="26" pageNumber="27">
|
||
Similar to apterous female in general structure and coloration with following exceptions: thorax with a pronotal lobe, anterior part with a subrhombic yellow marking, posterior part elongate, covering most of mesonotum; posterior margin broadly rounded and brownish (
|
||
<figureCitation id="14CC2A1EFFC629213B8AFA9473A50A5A" box="[917,1014,1318,1344]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="26" pageNumber="27">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213C1CFA94746B0A5A" box="[1027,1080,1318,1344]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138A2FADA71110AF1" blockId="26.[188,1399,1383,1515]" pageId="26" pageNumber="27">
|
||
<smallCapsWord id="8AAEA047FFC6292138A2FADA713D0A9A" baselines="1403,1403" box="[189,366,1384,1410]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="26" pageNumber="27">MEASUREMENTS</smallCapsWord>
|
||
. Body length 7.81, width 2.90, head width 1.45, interocular width 0.66, eye length (dorsal view) 0.53; pronotum: length 2.97, width 2.41; mesonotum width 2.90; metanotum: length 0.67, width 2.51; abdominal sternum VII: length 1.05, width 1.49; abdominal mediotergite I: length 0.67, width 1.19.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138F5F9A071F20936" blockId="26.[185,1399,1554,1827]" box="[234,417,1554,1580]" pageId="26" pageNumber="27">
|
||
<emphasis id="BE83EA89FFC6292138F5F9A071F20936" bold="true" box="[234,417,1554,1580]" pageId="26" pageNumber="27">Apterous male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292138A2F98471A30839" blockId="26.[185,1399,1554,1827]" pageId="26" pageNumber="27">
|
||
<smallCapsWord id="8AAEA047FFC6292138A2F984713D0954" baselines="1609,1609" box="[189,366,1590,1616]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="26" pageNumber="27">MEASUREMENTS</smallCapsWord>
|
||
. Body length 5.90–6.31, width 2.14–2.20, head width 1.32, interocular width 0.60, eye length (dorsal view) 0.42; relative lengths of antennal segments I–IV: 3.28: 0.92: 1.03: 0.87; pronotum: length 0.71, width 1.41; mesonotum: length 1.30, width 2.14; metanotum: length 0.71, width 1.82; abdomen length (ventral view) 1.98; abdominal sternum VII: length 0.46, width 0.83; abdominal mediotergite I: length 0.46, width 0.85; relative lengths of leg segments (femur: tibia: tarsal segment I: tarsal segment II): fore leg: 3.33: 2.88: 1.02: 0.63, middle leg: 8.72: 5.41: 2.66: 0.43, hind leg: 8.95:3.78: 0.16: 0.23.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC6292038A2F8F871F20E5C" blockId="26.[189,1399,1866,1963]" lastBlockId="27.[189,1398,265,327]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
<smallCapsWord id="8AAEA047FFC6292138A2F8F8711B0879" baselines="1885,1886" box="[189,328,1866,1892]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="26" pageNumber="27">COLORATION</smallCapsWord>
|
||
. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum mainly black with a median brownish-yellow stripe. Metanotum chiefly black, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFC629213A83F82272AD08B1" box="[668,766,1936,1963]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="26" pageNumber="27">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213B14F823736D08B1" box="[779,830,1937,1963]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[260,1340,536,1853]" captionTargetId="figure-53@27.[189,1398,511,1911]" captionTargetPageId="27" captionText="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="26" pageNumber="27">19B</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC629213B54F82373D108B1" box="[843,898,1937,1963]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="26" pageNumber="27">20H</figureCitation>
|
||
). Connexivum dorsally blackish (
|
||
<figureCitation id="14CC2A1EFFC629213D0FF822752108B1" box="[1296,1394,1936,1963]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="26" pageNumber="27">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC7292038A2FEBB70A20E39" box="[189,241,265,291]" captionStart="Fig" captionStartId="27.[189,232,1963,1989]" captionTargetBox="[260,1340,536,1853]" captionTargetId="figure-53@27.[189,1398,511,1911]" captionTargetPageId="27" captionText="Fig. 19. Habitus of Rhyacobates turgidus sp. nov., dorsal view. A. Holotype, apterous female (NKUM). B. Paratype, apterous male (NKUM). Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="27" pageNumber="28">19B</figureCitation>
|
||
). Mesosternum chiefly blackish with a median subtriangular yellowish spot (
|
||
<figureCitation id="14CC2A1EFFC729203C62FEBB748C0E39" box="[1149,1247,265,291]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="27" pageNumber="28">Fig. 20I</figureCitation>
|
||
). Abdominal venter dark-yellow.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC7292038A2FEDC732C0EB1" blockId="27.[189,1399,366,427]" pageId="27" pageNumber="28">
|
||
<smallCapsWord id="8AAEA047FFC7292038A2FEDC717F0E9C" baselines="385,385" box="[189,300,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="27" pageNumber="28">ABDOMEN</smallCapsWord>
|
||
. Abdomen relatively short. Abdominal mediotergite I extremely swollen (
|
||
<figureCitation id="14CC2A1EFFC729203CA2FEDC75710E92" box="[1213,1314,366,392]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="27" pageNumber="28">Figs 4C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC729203D2CFEDC753A0E92" box="[1331,1385,366,392]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="27" pageNumber="28">20H</figureCitation>
|
||
), nearly as long as three subsequent abdominal terga together.
|
||
</paragraph>
|
||
<caption id="D8886613FFC7292038A2F819730108F3" ID-DOI="http://doi.org/10.5281/zenodo.8385676" ID-Zenodo-Dep="8385676" httpUri="https://zenodo.org/record/8385676/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1963,1989]" targetBox="[260,1340,536,1853]" targetPageId="27">
|
||
<paragraph id="8C48369BFFC7292038A2F819730108F3" blockId="27.[189,1399,1963,2025]" pageId="27" pageNumber="28">
|
||
<emphasis id="BE83EA89FFC7292038A2F819714908DF" bold="true" box="[189,282,1963,1989]" pageId="27" pageNumber="28">Fig. 19.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4BF74D18FFC7292039BDF81972F308DF" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[418,672,1963,1989]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
|
||
<emphasis id="BE83EA89FFC7292039BDF81972F308DF" box="[418,672,1963,1989]" italics="true" pageId="27" pageNumber="28">Rhyacobates turgidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B057F2FFC729203AB9F81E735008DC" box="[678,771,1964,1990]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, dorsal view.
|
||
<emphasis id="BE83EA89FFC729203BBDF81973E408DF" bold="true" box="[930,951,1963,1989]" pageId="27" pageNumber="28">A</emphasis>
|
||
. Holotype, apterous female (NKUM).
|
||
<emphasis id="BE83EA89FFC7292038A2F87C708208F2" bold="true" box="[189,209,1998,2024]" pageId="27" pageNumber="28">B</emphasis>
|
||
. Paratype, apterous male (NKUM). Scale bar = 3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D8886613FFC0292738A2F94973B108F3" ID-DOI="http://doi.org/10.5281/zenodo.8385678" ID-Zenodo-Dep="8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1787,1813]" targetBox="[264,1323,291,1729]" targetPageId="28">
|
||
<paragraph id="8C48369BFFC0292738A2F94973B108F3" blockId="28.[188,1399,1787,2025]" pageId="28" pageNumber="29">
|
||
<emphasis id="BE83EA89FFC0292738A2F9497148080F" bold="true" box="[189,283,1787,1813]" pageId="28" pageNumber="29">Fig. 20.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName id="4BF74D18FFC029273A41F949730F080F" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[606,860,1787,1813]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
|
||
<emphasis id="BE83EA89FFC029273A41F949730F080F" box="[606,860,1787,1813]" italics="true" pageId="28" pageNumber="29">Rhyacobates turgidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B057F2FFC029273B7CF94E73ED080C" box="[867,958,1788,1814]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="BE83EA89FFC029273BD9F94973AE080F" bold="true" box="[966,1021,1787,1813]" pageId="28" pageNumber="29">A–B</emphasis>
|
||
. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view.
|
||
<emphasis id="BE83EA89FFC029273B30F8AC73170822" bold="true" box="[815,836,1822,1848]" pageId="28" pageNumber="29">C</emphasis>
|
||
. Body of female, excluding head and prothorax, ventral view.
|
||
<emphasis id="BE83EA89FFC029273947F8F0713E0846" bold="true" box="[344,365,1858,1884]" pageId="28" pageNumber="29">D</emphasis>
|
||
. Body of female, dealated macropterous form, lateral view.
|
||
<emphasis id="BE83EA89FFC029273C3AF8F0746A0846" bold="true" box="[1061,1081,1858,1884]" pageId="28" pageNumber="29">E</emphasis>
|
||
. Abdominal end of female, dorsal view.
|
||
<emphasis id="BE83EA89FFC029273945F8D7713F0865" bold="true" box="[346,364,1893,1919]" pageId="28" pageNumber="29">F</emphasis>
|
||
. Abdominal end of female, ventral view.
|
||
<emphasis id="BE83EA89FFC029273B6FF8D673D40864" bold="true" box="[880,903,1892,1918]" pageId="28" pageNumber="29">G</emphasis>
|
||
. Abdominal end of female, lateral view.
|
||
<emphasis id="BE83EA89FFC0292738A2F83A708708B8" bold="true" box="[189,212,1928,1954]" pageId="28" pageNumber="29">H</emphasis>
|
||
. Body of male, dorsal view.
|
||
<emphasis id="BE83EA89FFC029273A2DF83A726F08B8" bold="true" box="[562,572,1928,1954]" pageId="28" pageNumber="29">I</emphasis>
|
||
. Body of male, excluding head and prothorax, ventral view.
|
||
<emphasis id="BE83EA89FFC029273D07F83A757608B8" bold="true" box="[1304,1317,1928,1954]" pageId="28" pageNumber="29">J</emphasis>
|
||
. Basal part of right middle leg of male, ventral view.
|
||
<emphasis id="BE83EA89FFC029273AD2F81972B708DF" bold="true" box="[717,740,1963,1989]" pageId="28" pageNumber="29">K</emphasis>
|
||
. Proctiger of male, dorsal view.
|
||
<emphasis id="BE83EA89FFC029273C45F819743D08DF" bold="true" box="[1114,1134,1963,1989]" pageId="28" pageNumber="29">L</emphasis>
|
||
. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C48369BFFC1292638A2FEBB71390E5D" blockId="29.[187,1398,265,327]" pageId="29" pageNumber="30">
|
||
<smallCapsWord id="8AAEA047FFC1292638A2FEBB70B80E3B" baselines="284,284" box="[189,235,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="29" pageNumber="30">LEG</smallCapsWord>
|
||
. Middle trochanter with one spine; middle femur with scattered small spines, not arranged in distinct row (
|
||
<figureCitation id="14CC2A1EFFC1292638E6FE9F710E0E5D" box="[249,349,301,327]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="29" pageNumber="30">Fig. 20J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC1292638A2FEDD731D0ECB" blockId="29.[189,1399,367,466]" pageId="29" pageNumber="30">
|
||
<smallCapsWord id="8AAEA047FFC1292638A2FEDD71620E92" baselines="386,387" box="[189,305,367,393]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="29" pageNumber="30">GENITALIA</smallCapsWord>
|
||
. Abdominal segment VIII ventro-laterally impressed. Pygophore large, ovate. Proctiger with angular lobes laterally (
|
||
<figureCitation id="14CC2A1EFFC1292639CEFE2172630EB7" box="[465,560,403,429]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="29" pageNumber="30">Figs 6C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263A23FE2172200EB7" box="[572,627,403,429]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="29" pageNumber="30">20K</figureCitation>
|
||
). Paramere relatively slender, strongly curved at basal third, distal part tapering towards hook-shaped apex (
|
||
<figureCitation id="14CC2A1EFFC129263ABEFE0573520ECB" box="[673,769,439,465]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="29" pageNumber="30">Figs 8C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263B11FE0573110ECB" box="[782,834,439,465]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="29" pageNumber="30">20L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFC1292638A2FE4B72590D59" box="[189,522,505,579]" pageId="29" pageNumber="30" type="distribution">
|
||
<paragraph id="8C48369BFFC1292638A2FE4B71050D09" blockId="29.[189,342,505,531]" box="[189,342,505,531]" pageId="29" pageNumber="30">
|
||
<heading id="D70081F7FFC1292638A2FE4B71050D09" bold="true" box="[189,342,505,531]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis id="BE83EA89FFC1292638A2FE4B71050D09" bold="true" box="[189,342,505,531]" pageId="29" pageNumber="30">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC1292638A2FD9B72590D59" blockId="29.[189,522,553,579]" box="[189,522,553,579]" pageId="29" pageNumber="30">
|
||
<collectingCountry id="F4E0760BFFC1292638A2FD9B715B0D59" box="[189,264,553,579]" name="China" pageId="29" pageNumber="30">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="4E33F879FFC12926390CFD9B71CB0D59" box="[275,408,553,579]" country="China" name="Chongqing" pageId="29" pageNumber="30">Chongqing</collectingRegion>
|
||
,
|
||
<collectingRegion id="4E33F879FFC1292639BBFD9B72550D59" box="[420,518,553,579]" country="China" name="Sichuan" pageId="29" pageNumber="30">Sichuan</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFC1292638A2FDD972C90CE2" pageId="29" pageNumber="30" type="discussion">
|
||
<paragraph id="8C48369BFFC1292638A2FDD971F90D9F" blockId="29.[189,426,619,645]" box="[189,426,619,645]" pageId="29" pageNumber="30">
|
||
<heading id="D70081F7FFC1292638A2FDD971F90D9F" bold="true" box="[189,426,619,645]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis id="BE83EA89FFC1292638A2FDD971F90D9F" bold="true" box="[189,426,619,645]" pageId="29" pageNumber="30">Comparative notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFC1292638A2FD2872C90CE2" blockId="29.[189,1399,666,1016]" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4BF74D18FFC1292638A2FD2871930DAE" authority="Leng & Tran, 2023" authorityName="Leng & Tran" authorityYear="2023" box="[189,448,666,692]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
|
||
<emphasis id="BE83EA89FFC1292638A2FD2871930DAE" box="[189,448,666,692]" italics="true" pageId="29" pageNumber="30">Rhyacobates turgidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A5B057F2FFC1292639D8FD2972700DAF" box="[455,547,667,693]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is closely related to
|
||
<taxonomicName id="4BF74D18FFC129263B06FD2973F10DAE" authorityName="Andersen & Chen" authorityYear="1995" box="[793,930,667,692]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="BE83EA89FFC129263B06FD2973F10DAE" box="[793,930,667,692]" italics="true" pageId="29" pageNumber="30">R. recurvus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFC129263BC4FD2974D30DAE" authorityName="Tran & Nguyen" authorityYear="2016" box="[987,1152,667,692]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC129263BC4FD2974D30DAE" box="[987,1152,667,692]" italics="true" pageId="29" pageNumber="30">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
. These three species possess a robust body and similar outline of abdominal segment VII (
|
||
<figureCitation id="14CC2A1EFFC129263C00FD0C74960DC2" box="[1055,1221,702,729]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="29" pageNumber="30">Fig. 3C, I, N</figureCitation>
|
||
).
|
||
<taxonomicName id="4BF74D18FFC129263CC1FD0C71730DE6" authorityName="Leng & Tran" authorityYear="2023" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turgidus">
|
||
<emphasis id="BE83EA89FFC129263CC1FD0C71730DE6" italics="true" pageId="29" pageNumber="30">Rhyacobates turgidus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4BF74D18FFC129263A6AFD51734D0DE6" authorityName="Tran & Nguyen" authorityYear="2016" box="[629,798,739,764]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC129263A6AFD51734D0DE6" box="[629,798,739,764]" italics="true" pageId="29" pageNumber="30">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
by median yellowish stripes on the mesonotum, metanotum and abdominal mediotergite I (
|
||
<figureCitation id="14CC2A1EFFC129263AFDFCB4731F0C3A" box="[738,844,774,800]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="29" pageNumber="30">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263B7FFCB473C40C3A" box="[864,919,774,800]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="29" pageNumber="30">20A</figureCitation>
|
||
), which are absent in
|
||
<taxonomicName id="4BF74D18FFC129263CDDFCB5753C0C3A" authorityName="Tran & Nguyen" authorityYear="2016" box="[1218,1391,775,800]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="BE83EA89FFC129263CDDFCB5753C0C3A" box="[1218,1391,775,800]" italics="true" pageId="29" pageNumber="30">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4BF74D18FFC1292638A2FC9871960C5E" authorityName="Leng & Tran" authorityYear="2023" box="[189,453,810,836]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turgidus">
|
||
<emphasis id="BE83EA89FFC1292638A2FC9871960C5E" box="[189,453,810,836]" italics="true" pageId="29" pageNumber="30">Rhyacobates turgidus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4BF74D18FFC129263B3FFC9973FE0C5E" authorityName="Andersen & Chen" authorityYear="1995" box="[800,941,811,836]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="BE83EA89FFC129263B3FFC9973FE0C5E" box="[800,941,811,836]" italics="true" pageId="29" pageNumber="30">R. recurvus</emphasis>
|
||
</taxonomicName>
|
||
by the following characteristics: (1) in lateral view,
|
||
<taxonomicName id="4BF74D18FFC129263962FCFD72570C72" authorityName="Leng & Tran" authorityYear="2023" box="[381,516,846,872]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turgidus">
|
||
<emphasis id="BE83EA89FFC129263962FCFD72570C72" box="[381,516,846,872]" italics="true" pageId="29" pageNumber="30">R. turgidus</emphasis>
|
||
</taxonomicName>
|
||
has a nearly straight abdomen (
|
||
<figureCitation id="14CC2A1EFFC129263B8FFCFC73BC0C73" box="[912,1007,846,873]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="29" pageNumber="30">Fig. 5C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263BE0FCFC74650C72" box="[1023,1078,846,872]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="29" pageNumber="30">20D</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4BF74D18FFC129263CA6FCFD75160C72" authorityName="Andersen & Chen" authorityYear="1995" box="[1209,1349,847,872]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="BE83EA89FFC129263CA6FCFD75160C72" box="[1209,1349,847,872]" italics="true" pageId="29" pageNumber="30">R. recurvus</emphasis>
|
||
</taxonomicName>
|
||
has its abdomen curved dorsad (
|
||
<figureCitation id="14CC2A1EFFC129263A3FFCC072DE0C97" box="[544,653,882,909]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="29" pageNumber="30">Figs 5M</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263A82FCC072800C96" box="[669,723,882,908]" captionStart="Fig" captionStartId="54.[189,232,1787,1813]" captionTargetBox="[268,1308,289,1709]" captionTargetId="figure-14@54.[239,1349,267,1738]" captionTargetPageId="54" captionText="Fig. 31. Morphological features of Rhyacobates recurvus Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385702" httpUri="https://zenodo.org/record/8385702/files/figure.png" pageId="29" pageNumber="30">31C</figureCitation>
|
||
); (2)
|
||
<taxonomicName id="4BF74D18FFC129263B03FCC173F70C96" authorityName="Leng & Tran" authorityYear="2023" box="[796,932,882,908]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turgidus">
|
||
<emphasis id="BE83EA89FFC129263B03FCC173F70C96" box="[796,932,882,908]" italics="true" pageId="29" pageNumber="30">R. turgidus</emphasis>
|
||
</taxonomicName>
|
||
has an extremely swollen abdominal mediotergite I, which is nearly as long as the three subsequent abdominal tergites, and distinctly short abdominal terga II–VI (
|
||
<figureCitation id="14CC2A1EFFC129263A3EFC0872FA0CCE" box="[545,681,954,980]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="29" pageNumber="30">Figs 3C–D</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263AA9FC0873430CCE" box="[694,784,954,980]" captionStart="Fig" captionStartId="28.[189,232,1787,1813]" captionTargetBox="[264,1323,291,1729]" captionTargetId="figure-15@28.[228,1359,265,1747]" captionTargetPageId="28" captionText="Fig. 20. Morphological features of Rhyacobates turgidus sp. nov. A–B. Body of female, apterous form (A) and dealated macropterous form (B), dorsal view. C. Body of female, excluding head and prothorax, ventral view. D. Body of female, dealated macropterous form, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view.K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385678" httpUri="https://zenodo.org/record/8385678/files/figure.png" pageId="29" pageNumber="30">20A–B</figureCitation>
|
||
), whereas abdominal terga I–VI of
|
||
<taxonomicName id="4BF74D18FFC129263CDFFC0975180CCE" authorityName="Andersen & Chen" authorityYear="1995" box="[1216,1355,955,980]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="BE83EA89FFC129263CDFFC0975180CCE" box="[1216,1355,955,980]" italics="true" pageId="29" pageNumber="30">R. recurvus</emphasis>
|
||
</taxonomicName>
|
||
are relatively normal-sized (
|
||
<figureCitation id="14CC2A1EFFC1292639FBFC6C721B0CE2" box="[484,584,990,1016]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="29" pageNumber="30">Figs 3N</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFC129263A4AFC6C72DE0CE2" box="[597,653,990,1016]" captionStart="Fig" captionStartId="54.[189,232,1787,1813]" captionTargetBox="[268,1308,289,1709]" captionTargetId="figure-14@54.[239,1349,267,1738]" captionTargetPageId="54" captionText="Fig. 31. Morphological features of Rhyacobates recurvus Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D. Abdominal end of female, lateral view. E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Body of male, dorsal view. H. Body of male, excluding head and prothorax, ventral view. I. Basal part of right middle leg of male, ventral view. J. Proctiger of male, dorsal view. K. Left paramere, lateral aspect, two different views. Scale bars: A–I = 1 mm; J–K = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385702" httpUri="https://zenodo.org/record/8385702/files/figure.png" pageId="29" pageNumber="30">31A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |