422 lines
42 KiB
XML
422 lines
42 KiB
XML
<document id="57222EEA789623523466E735171775F4" ID-DOI="10.1649/1066.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268265367" checkinUser="valdenar" docAuthor="Ratcliffe, Brett C." docDate="1821" docId="041BEE3DB042A77EFEE3FBB6FCB2FA2C" docLanguage="en" docName="1066.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1066.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cyclocephala melanopoda Ratcliffe 2008, new species" docType="treatment" docVersion="3" lastPageNumber="241" masterDocId="F8229645B04CA76EFFDDFFBAFF8DFFAC" masterDocTitle="More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1699205293535" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="20883C24CC74D2CD1BC66A697462739F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4BB753E4377F2DAA13C55AA8C84860C3">
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<mods:title id="BD9FD37BC0B7AB351E3C3F52AE402FBD">More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
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<mods:name id="26EBABFF40B41CCA4BD09C36174E6838" type="personal">
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<mods:role id="FB0478CF3F1EBCA50F2F35536C0273F9">
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<mods:roleTerm id="83AEE4D478DFCB4A9802E217AFFF81A2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B9BF9337989CF36DC0D6E5CD9613C994">Ratcliffe, Brett C.</mods:namePart>
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<mods:typeOfResource id="6701590906D92C281995BFCCF90B66CE">text</mods:typeOfResource>
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<mods:titleInfo id="A4D31B14231758F60429A6553D609E9B">
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<mods:title id="6B1B969007F6A875189EF5F810DFFE57">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="D49BD87381B4EF416E48594D8C62CCF8">
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<mods:date id="06411569D4E2390E52772115795FB28F">1821</mods:date>
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<mods:detail id="E48639E98F2ADC9F95C72D4F642437C0" type="pubDate">
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<mods:number id="8D6507DDA9BDA80EE471108C2B58AC45">2008-06-30</mods:number>
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</mods:detail>
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<mods:detail id="183E09BC94C98A6534F3E8A48A858180" type="volume">
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<mods:number id="AEE4A6E252CFBA9419F90182290DD741">62</mods:number>
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</mods:detail>
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<mods:detail id="8042A8FA19D399F8C5985F28BA4FDB19" type="issue">
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<mods:number id="E264CECBE7BB5706753C1C35E6C084E0">2</mods:number>
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<mods:extent id="36CD9CBF165CA2C9CEDA228209442D52" unit="page">
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<mods:start id="C96477ED4311A245714C23AB471C75E8">221</mods:start>
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<mods:end id="EF5D3C82CCD0A2F59AE2DF912C1CFB42">241</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location id="20EEECD0C318FFF55293B729C01055D1">
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<mods:url id="A5BFCA635F381384F15B422CFD7DFEA3">http://dx.doi.org/10.1649/1066.1</mods:url>
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</mods:location>
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<mods:classification id="B14C26DE08C306F742A21E71AD20C865">journal article</mods:classification>
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<mods:identifier id="79BD7E81752BD151B3CE7818ECE3BF0E" type="DOI">10.1649/1066.1</mods:identifier>
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<mods:identifier id="E8E2FA9BA9E3BB00344DD9BB14256ACB" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="04584E4F87794B9378C2AC4E3A80A510" type="Zenodo-Dep">5369967</mods:identifier>
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</mods:mods>
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<treatment id="041BEE3DB042A77EFEE3FBB6FCB2FA2C" LSID="urn:lsid:plazi:treatment:041BEE3DB042A77EFEE3FBB6FCB2FA2C" httpUri="http://treatment.plazi.org/id/041BEE3DB042A77EFEE3FBB6FCB2FA2C" lastPageId="16" pageId="14">
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<subSubSection id="C4A80CA0B042A760FEE3FBB6FCAFFB8E" box="[318,802,1036,1058]" pageId="14" type="nomenclature">
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<paragraph id="8C0D5F2BB042A760FEE3FBB6FCAFFB8E" blockId="14.[318,802,1036,1086]" box="[318,802,1036,1058]" pageId="14">
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<heading id="D745E847B042A760FEE3FBB6FCAFFB8E" box="[318,802,1036,1058]" centered="true" fontSize="9" level="2" pageId="14" reason="2">
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<emphasis id="BEC68339B042A760FEE3FBB6FCAFFB8E" bold="true" box="[318,802,1036,1058]" pageId="14">
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<taxonomicName id="4BB224A8B042A760FEE3FBB6FD29FB8E" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2008" box="[318,676,1036,1058]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="14" phylum="Arthropoda" rank="species" species="melanopoda" status="sp. nov.">
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<emphasis id="BEC68339B042A760FEE3FBB6FDCEFB8E" bold="true" box="[318,579,1036,1058]" italics="true" pageId="14">Cyclocephala melanopoda</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="A5F53E42B042A760FD6DFBB6FCAFFB8E" box="[688,802,1036,1058]" pageId="14" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B042A760FE33FB92FDF9FB92" box="[494,628,1064,1086]" pageId="14" type="description">
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<paragraph id="8C0D5F2BB042A760FE33FB92FDF9FB92" blockId="14.[318,802,1036,1086]" box="[494,628,1064,1086]" pageId="14">
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(
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<figureCitation id="148943AEB042A760FE28FB92FDE0FB92" box="[501,621,1064,1086]" captionStart="Figs" captionStartId="14.[167,206,915,934]" captionTargetBox="[146,972,194,892]" captionTargetId="figure-203@14.[146,976,194,892]" captionTargetPageId="14" captionText="Figs. 26–30. Cyclocephala melanopoda: 26) dorsal view of paratype; 27–28) parameres (caudal and lateral views); 29–30) right protarsus, median view, of male of 29) C. melanopoda and 30) C. herteli." figureDoi="http://doi.org/10.5281/zenodo.5369979" httpUri="https://zenodo.org/record/5369979/files/figure.png" pageId="14">Figs. 26–29</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B042A761FF77FBE3FCA9FC61" lastPageId="15" pageId="14" type="materials_examined">
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<paragraph id="8C0D5F2BB042A760FF77FBE3FE15FABD" blockId="14.[146,976,1113,1569]" pageId="14">
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<emphasis id="BEC68339B042A760FF77FBE3FECBFBC3" bold="true" box="[170,326,1113,1135]" pageId="14">Type Material.</emphasis>
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<materialsCitation id="3CDA5576B042A760FE8CFBE3FE9CFB09" collectingDate="1997-05-18" collectingDateMax="1997-05-24" collectingDateMin="1997-05-18" collectorName="C. & K. Messenger." country="Ecuador" latitude="-0.21916667" location="Tinalandia Resort" longLatPrecision="21" longitude="-79.152504" pageId="14" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
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<typeStatus id="5309E189B042A760FE8CFBE3FE3CFBC3" box="[337,433,1113,1135]" pageId="14" type="holotype">Holotype</typeStatus>
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male, labeled ‘‘
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<collectingCountry id="F4A51FBBB042A760FD87FBE3FD6DFBC3" box="[602,736,1113,1135]" name="Ecuador" pageId="14">ECUADOR</collectingCountry>
|
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:
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<collectingRegion id="4E7691C9B042A760FD32FBE3FC15FBC3" box="[751,920,1113,1135]" country="Ecuador" name="Pichincha" pageId="14">Pichincha Prov.</collectingRegion>
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||
, Sto. Domingo,
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<location id="896D09F0B042A760FED9FBCEFE49FB26" LSID="urn:lsid:plazi:treatment:041BEE3DB042A77EFEE3FBB6FCB2FA2C:896D09F0B042A760FED9FBCEFE49FB26" box="[260,452,1140,1162]" country="Ecuador" latitude="-0.21916667" longLatPrecision="21" longitude="-79.152504" name="Tinalandia Resort" pageId="14" stateProvince="Pichincha">Tinalandia Resort</location>
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,
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<geoCoordinate id="E98639ECB042A760FE0FFBCEFD9BFB26" box="[466,534,1140,1162]" degrees="0" direction="south" minutes="13" orientation="latitude" pageId="14" precision="15" seconds="9" value="-0.21916667">
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0
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<emphasis id="BEC68339B042A760FE03FBCEFE6AFB25" bold="true" box="[478,487,1140,1161]" italics="true" pageId="14">°</emphasis>
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13
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<emphasis id="BEC68339B042A760FE22FBCEFD8AFB26" bold="true" box="[511,519,1140,1162]" pageId="14">9</emphasis>
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S
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</geoCoordinate>
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,
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<geoCoordinate id="E98639ECB042A760FDF9FBCEFDF2FB26" box="[548,639,1140,1162]" degrees="79" direction="west" minutes="09" orientation="longitude" pageId="14" precision="15" seconds="9" value="-79.152504">
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79
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<emphasis id="BEC68339B042A760FDE1FBCEFDC8FB25" bold="true" box="[572,581,1140,1161]" italics="true" pageId="14">°</emphasis>
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||
09
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<emphasis id="BEC68339B042A760FD80FBCEFDE8FB26" bold="true" box="[605,613,1140,1162]" pageId="14">9</emphasis>
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W
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</geoCoordinate>
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||
,
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<date id="F80C79EBB042A760FD50FBCEFC95FB26" box="[653,792,1140,1162]" pageId="14" value="1997-05-18" valueMax="1997-05-24" valueMin="1997-05-18">
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||
<collectingDate id="E8488003B042A760FD50FBCEFC95FB26" box="[653,792,1140,1162]" pageId="14" value="1997-05-18" valueMax="1997-05-24" valueMin="1997-05-18">
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||
V-18-24-
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<quantity id="4B4AF2CEB042A760FD3BFBCEFC95FB26" box="[742,792,1140,1162]" metricMagnitude="6" metricUnit="m" metricValue="1.99776" pageId="14" unit="m" value="1997760.0">1997</quantity>
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</collectingDate>
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||
</date>
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||
, 760 m,
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<collectorName id="21473AFDB042A760FCAAFBCEFE84FB09" pageId="14">C. & K. Messenger.</collectorName>
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||
’’
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||
</materialsCitation>
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||
<materialsCitation id="3CDA5576B042A760FECAFB35FE15FABD" collectingDate="1976-02-22" collectingDateMax="1999-04-21" collectingDateMin="1976-02-22" collectionCode="T" collectorName="I. P. Smith & D. C. Carlson Collection & Hovore. & One & H. & A. Howden." country="Ecuador" latitude="-0.21916667" location="Rio Palenque" longLatPrecision="21" longitude="-79.152504" municipality="Tinalandia" pageId="14" specimenCount="3" stateProvince="Pichincha" typeStatus="allotype">
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<typeStatus id="5309E189B042A760FECAFB35FEE2FB09" box="[279,367,1167,1189]" pageId="14" type="allotype">Allotype</typeStatus>
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||
female labeled ‘‘
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||
<collectingCountry id="F4A51FBBB042A760FDCAFB35FD10FB09" box="[535,669,1167,1189]" name="Ecuador" pageId="14">ECUADOR</collectingCountry>
|
||
:
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||
<collectingRegion id="4E7691C9B042A760FD7AFB35FC86FB09" box="[679,779,1167,1189]" country="Ecuador" name="Pichincha" pageId="14">Pichincha</collectingRegion>
|
||
Pr.,
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||
<collectingMunicipality id="6C69C551B042A760FCE1FB35FC20FB09" box="[828,941,1167,1189]" pageId="14">Tinalandia</collectingMunicipality>
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||
,
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||
<date id="F80C79EBB042A760FC6AFB35FE8FFB6C" pageId="14" value="1999-04-21">
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<collectingDate id="E8488003B042A760FC6AFB35FE8FFB6C" pageId="14" value="1999-04-21">21 April 1999</collectingDate>
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||
</date>
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||
, F.
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||
<collectionCode id="EAA3C7EEB042A760FEF1FB10FEB2FB6C" box="[300,319,1194,1216]" name="Tavera, Department of Geology and Geophysics" pageId="14">T</collectionCode>
|
||
. Hovore/
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||
<collectorName id="21473AFDB042A760FE7EFB10FD9FFB6C" box="[419,530,1194,1216]" pageId="14">I. P. Smith</collectorName>
|
||
//
|
||
<collectorName id="21473AFDB042A760FDFFFB10FCAEFB6C" box="[546,803,1194,1216]" pageId="14">D. C. Carlson Collection</collectorName>
|
||
, Gift from F.
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||
<collectionCode id="EAA3C7EEB042A760FC64FB10FC41FB6C" box="[953,972,1194,1216]" name="Tavera, Department of Geology and Geophysics" pageId="14">T</collectionCode>
|
||
.
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||
<collectorName id="21473AFDB042A760FF4FFB7FFF67FB77" box="[146,234,1221,1243]" pageId="14">Hovore.</collectorName>
|
||
’’
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||
<collectorName id="21473AFDB042A760FF23FB7FFEA5FB77" box="[254,296,1221,1243]" pageId="14">One</collectorName>
|
||
<typeStatus id="5309E189B042A760FEEEFB7FFE00FB77" box="[307,397,1221,1243]" pageId="14" type="paratype">paratype</typeStatus>
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||
male with same data as allotype.
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||
<collectorName id="21473AFDB042A760FCD8FB7FFCA2FB77" box="[773,815,1221,1243]" pageId="14">One</collectorName>
|
||
<typeStatus id="5309E189B042A760FCE7FB7FFC19FB77" box="[826,916,1221,1243]" pageId="14" type="paratype">paratype</typeStatus>
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||
male labeled ‘‘
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||
<collectingCountry id="F4A51FBBB042A760FF33FB5AFEF9FB5A" box="[238,372,1248,1270]" name="Ecuador" pageId="14">ECUADOR</collectingCountry>
|
||
, 700
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||
<emphasis id="BEC68339B042A760FE7BFB5AFE23FB5A" bold="true" box="[422,430,1248,1270]" pageId="14">9</emphasis>
|
||
,
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||
<location id="896D09F0B042A760FE1DFB5AFDDDFB5A" LSID="urn:lsid:plazi:treatment:041BEE3DB042A77EFEE3FBB6FCB2FA2C:896D09F0B042A760FE1DFB5AFDDDFB5A" box="[448,592,1248,1270]" country="Ecuador" municipality="Tinalandia" name="Rio Palenque" pageId="14" stateProvince="Pichincha">Rio Palenque</location>
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||
,
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||
<locationDeviation id="9DBF4609B042A760FD82FB5AFCC1FB5A" box="[607,844,1248,1270]" location="St. Domingo" pageId="14">
|
||
<quantity id="4B4AF2CEB042A760FD82FB5AFD2DFB5A" box="[607,672,1248,1270]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="14" unit="km" value="47.0">47 km</quantity>
|
||
S St. Domingo
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||
</locationDeviation>
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||
,
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||
<date id="F80C79EBB042A760FC87FB5AFF49FABD" pageId="14" value="1976-02-22" valueMax="1976-02-27" valueMin="1976-02-22">
|
||
<collectingDate id="E8488003B042A760FC87FB5AFF49FABD" pageId="14" value="1976-02-22" valueMax="1976-02-27" valueMin="1976-02-22">Feb. 22–27 1976</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="21473AFDB042A760FF0DFB41FE02FABD" box="[208,399,1275,1297]" pageId="14">H. & A. Howden.</collectorName>
|
||
’’
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB042A760FF77FAADFE37FAD2" blockId="14.[146,976,1113,1569]" pageId="14">
|
||
<materialsCitation id="3CDA5576B042A760FF77FAADFE3AFAD2" collectionCode="CA" collectorName="D. Carlson & Lincoln, NE & One & David Carlson Collection & Fair Oaks" country="Canada" county="Ottawa" location="University of Nebraska State Museum" municipality="Canadian Museum of Nature Collection" pageId="14" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="5309E189B042A760FF77FAADFE87FA81" box="[170,266,1303,1325]" pageId="14" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="5309E189B042A760FEE2FAADFE1CFA81" box="[319,401,1303,1325]" pageId="14" type="allotype">allotype</typeStatus>
|
||
(property of
|
||
<collectorName id="21473AFDB042A760FDC1FAADFD02FA81" box="[540,655,1303,1325]" pageId="14">D. Carlson</collectorName>
|
||
) deposited at the
|
||
<location id="896D09F0B042A760FC95FAADFE13FAE4" LSID="urn:lsid:plazi:treatment:041BEE3DB042A77EFEE3FBB6FCB2FA2C:896D09F0B042A760FC95FAADFE13FAE4" country="Canada" county="Ottawa" municipality="Canadian Museum of Nature Collection" name="University of Nebraska State Museum" pageId="14">University of Nebraska State Museum</location>
|
||
,
|
||
<collectorName id="21473AFDB042A760FE72FA88FDB5FAE4" box="[431,568,1330,1352]" pageId="14">Lincoln, NE</collectorName>
|
||
.
|
||
<collectorName id="21473AFDB042A760FD95FA88FDFFFAE4" box="[584,626,1330,1352]" pageId="14">One</collectorName>
|
||
<typeStatus id="5309E189B042A760FDA2FA88FD54FAE4" box="[639,729,1330,1352]" pageId="14" type="paratype">paratype</typeStatus>
|
||
in the
|
||
<collectorName id="21473AFDB042A760FCEFFA88FF76FACF" pageId="14">David Carlson Collection</collectorName>
|
||
(
|
||
<collectorName id="21473AFDB042A760FED5FAF7FEFFFACF" box="[264,370,1357,1379]" pageId="14">Fair Oaks</collectorName>
|
||
,
|
||
<collectionCode id="EAA3C7EEB042A760FEA1FAF7FE2CFACF" box="[380,417,1357,1379]" country="USA" name="Chicago Academy of Sciences" pageId="14" type="Museum">CA</collectionCode>
|
||
) and
|
||
<specimenCount id="9AB494A2B042A760FE04FAF7FDD3FACF" box="[473,606,1357,1379]" pageId="14" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
in the
|
||
<collectingMunicipality id="6C69C551B042A760FD7EFAF7FF73FAD2" pageId="14">Canadian Museum of Nature Collection</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="656C27A7B042A760FED7FAD2FED5FAD2" box="[266,344,1384,1406]" pageId="14">Ottawa</collectingCounty>
|
||
,
|
||
<collectingCountry id="F4A51FBBB042A760FEB9FAD2FE3AFAD2" box="[356,439,1384,1406]" name="Canada" pageId="14">Canada</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB042A761FF77FA3EFD6FFD58" blockId="14.[146,976,1113,1569]" lastBlockId="15.[175,1005,194,1570]" lastPageId="15" pageId="14">
|
||
<emphasis id="BEC68339B042A760FF77FA3EFE80FA36" bold="true" box="[170,269,1412,1434]" pageId="14">
|
||
<typeStatus id="5309E189B042A760FF77FA3EFE84FA36" box="[170,265,1412,1434]" pageId="14" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. Length
|
||
<quantity id="4B4AF2CEB042A760FE60FA3EFD80FA36" box="[445,525,1412,1434]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="14" unit="mm" value="9.8">9.8 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB042A760FCD0FA3EFCD3FA36" box="[781,862,1412,1434]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="14" unit="mm" value="4.2">4.2 mm</quantity>
|
||
. Color of clypeus, pronotum, elytra, pygidium, femora, and tibiae testaceous; frons, elytral pattern, sides of pygidium, apices of tibiae and femora, meso-and metatarsi, procoxae, venter of meso- and metathorax, and abdominal sternites black; protarsi piceous; protarsal claws black, meso- and metatarsal claws testaceous. Elytra with black, curved line extending from base either side of scutellum to suture at about middle; 3 black, short streaks on disc.
|
||
<emphasis id="BEC68339B043A761FCCFFF78FCC5FF7B" box="[786,840,194,215]" italics="true" pageId="15">Head</emphasis>
|
||
: Surface finely shagreened, opaque. Frons between eyes moderately densely punctate, punctures small. Remainder of head vaguely punctate or roughened. Clypeus with apex parabolic, rimmed in black, weakly reflexed. Interocular width equals 3.0 transverse eye diameters. Antenna 10-segmented, club subequal in length to segments 2–7.
|
||
<emphasis id="BEC68339B043A761FE8CFEF3FE39FEF2" box="[337,436,329,350]" italics="true" pageId="15">Pronotum</emphasis>
|
||
: Surface weakly shining, finely shagreened, punctate; punctures dense (separated by; 1–2 puncture diameters), moderate in size, becoming indistinct near lateral margins. Posterior angles narrowly rounded. Base with complete marginal bead.
|
||
<emphasis id="BEC68339B043A761FE66FE20FE74FE03" box="[443,505,410,431]" italics="true" pageId="15">Elytra</emphasis>
|
||
: Surface weakly shining (a little more so than pronotum), punctate; punctures moderate in density and size (becoming slightly larger on sides), many weakly ocellate, paired punctate striae distinct. Posterior third with sparse, minute, stout setae.
|
||
<taxonomicName id="4BB224A8B043A761FE37FE51FD16FDAD" authority=": Surface" authorityName="Surface" box="[490,667,491,513]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B043A761FE37FE51FDCBFDAC" box="[490,582,491,512]" italics="true" pageId="15">Pygidium</emphasis>
|
||
: Surface
|
||
</taxonomicName>
|
||
weakly shining, finely shagreened, weakly punctate; punctures moderately dense, minute, some setigerous; setae sparse, pale, small. In lateral view, surface weakly convex in basal 3/4, more strongly convex in apical fourth.
|
||
<emphasis id="BEC68339B043A761FE81FD87FE07FDFE" box="[348,394,573,594]" italics="true" pageId="15">Legs</emphasis>
|
||
: Protibia bidentate. Protarsus strongly enlarged; tarsomeres 2–4 each slightly larger than preceding; 5th long, curved, concave ventrally, ventromedial edge deeply emarginate, ventrolateral edge strongly produced onto a lobe (
|
||
<figureCitation id="148943AEB043A761FF30FD37FEBBFD0F" box="[237,310,653,675]" captionStart="Figs" captionStartId="14.[167,206,915,934]" captionTargetBox="[146,972,194,892]" captionTargetId="figure-203@14.[146,976,194,892]" captionTargetPageId="14" captionText="Figs. 26–30. Cyclocephala melanopoda: 26) dorsal view of paratype; 27–28) parameres (caudal and lateral views); 29–30) right protarsus, median view, of male of 29) C. melanopoda and 30) C. herteli." figureDoi="http://doi.org/10.5281/zenodo.5369979" httpUri="https://zenodo.org/record/5369979/files/figure.png" pageId="15">Fig. 29</figureCitation>
|
||
); median claw enlarged, strongly curved, apex obliquely truncate, entire. Metatarsus nearly twice as long as metatibia.
|
||
<emphasis id="BEC68339B043A761FD70FD13FD63FD12" box="[685,750,681,702]" italics="true" pageId="15">Venter</emphasis>
|
||
: Prosternal process short, stout; apex transversely oval with raised, transverse ‘‘button’’ on anterior half. Last sternite very deeply emarginate.
|
||
<emphasis id="BEC68339B043A761FE2FFD65FDD7FD58" box="[498,602,735,756]" italics="true" pageId="15">Parameres</emphasis>
|
||
:
|
||
<figureCitation id="148943AEB043A761FDBAFD64FD50FD58" box="[615,733,734,756]" captionStart="Figs" captionStartId="14.[167,206,915,934]" captionTargetBox="[146,972,194,892]" captionTargetId="figure-203@14.[146,976,194,892]" captionTargetPageId="14" captionText="Figs. 26–30. Cyclocephala melanopoda: 26) dorsal view of paratype; 27–28) parameres (caudal and lateral views); 29–30) right protarsus, median view, of male of 29) C. melanopoda and 30) C. herteli." figureDoi="http://doi.org/10.5281/zenodo.5369979" httpUri="https://zenodo.org/record/5369979/files/figure.png" pageId="15">Figs. 27–28</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB043A761FF1AFD40FCA9FC61" blockId="15.[175,1005,194,1570]" pageId="15">
|
||
<emphasis id="BEC68339B043A761FF1AFD40FEAFFCBC" bold="true" box="[199,290,762,784]" pageId="15">
|
||
<typeStatus id="5309E189B043A761FF1AFD40FE93FCBC" box="[199,286,762,784]" pageId="15" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. Length
|
||
<quantity id="4B4AF2CEB043A761FE0AFD40FDA5FCBC" box="[471,552,762,784]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="15" unit="mm" value="9.3">9.3 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB043A761FCCCFD40FCEFFCBC" box="[785,866,762,784]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="15" unit="mm" value="4.9">4.9 mm</quantity>
|
||
. As
|
||
<typeStatus id="5309E189B043A761FC4EFD40FC60FCBC" box="[915,1005,762,784]" pageId="15" type="holotype">holotype</typeStatus>
|
||
except in the following respects: Color of clypeus dark brownish yellow; elytra with curved, black line interrupted at base either side of scutellum; subapical swelling on lateral margin black.
|
||
<emphasis id="BEC68339B043A761FDC8FCF1FDC6FCCC" box="[533,587,843,864]" italics="true" pageId="15">Head</emphasis>
|
||
: Clypeus weakly shining (not opaque), surface densely, finely rugopunctate.
|
||
<emphasis id="BEC68339B043A761FDF1FCDCFDE6FCD7" box="[556,619,870,891]" italics="true" pageId="15">Elytra</emphasis>
|
||
: In ventral view, epipleuron abruptly narrowed at level of sternite 2; lateral margin just above this slightly swollen. Minute setae sparser.
|
||
<taxonomicName id="4BB224A8B043A761FE41FC27FDCEFC1E" authority=": Setae" authorityName="Setae" box="[412,579,924,946]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B043A761FE41FC27FE76FC1E" box="[412,507,925,946]" italics="true" pageId="15">Pygidium</emphasis>
|
||
: Setae
|
||
</taxonomicName>
|
||
sparse, minute. In lateral view, surface nearly flat.
|
||
<emphasis id="BEC68339B043A761FEFAFC02FEDBFC61" box="[295,342,952,973]" italics="true" pageId="15">Legs</emphasis>
|
||
: Protibia tridentate. Protarsus not enlarged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B043A761FF1AFC69FD99FBF9" pageId="15" type="description">
|
||
<paragraph id="8C0D5F2BB043A761FF1AFC69FD99FBF9" blockId="15.[175,1005,194,1570]" pageId="15">
|
||
<emphasis id="BEC68339B043A761FF1AFC69FEA0FC45" bold="true" box="[199,301,979,1001]" pageId="15">Variation.</emphasis>
|
||
Males (
|
||
<specimenCount id="9AB494A2B043A761FEA2FC69FE77FC45" box="[383,506,979,1001]" pageId="15" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
|
||
). Length 9.8–10.0 mm; width across humeri
|
||
<quantity id="4B4AF2CEB043A761FC1FFC69FE8CFBA8" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.7" metricValueMin="4.6" pageId="15" unit="mm" value="4.65" valueMax="4.7" valueMin="4.6">4.6– 4.7 mm</quantity>
|
||
. As
|
||
<typeStatus id="5309E189B043A761FEF2FC54FE04FBA8" box="[303,393,1006,1028]" pageId="15" type="holotype">holotype</typeStatus>
|
||
except in the following respects: In
|
||
<specimenCount id="9AB494A2B043A761FCDFFC54FC02FBA8" box="[770,911,1006,1028]" pageId="15" type="generic">one specimen</specimenCount>
|
||
, color of clypeus and sides of pygidium dark brownish yellow; pronotum with trace of light brown, longitudinal macula either side of midline on basal half; elytra with posterior 2 discal spots connected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B043A761FF1AFBE1FD4EFB20" pageId="15" type="etymology">
|
||
<paragraph id="8C0D5F2BB043A761FF1AFBE1FD4EFB20" blockId="15.[175,1005,194,1570]" pageId="15">
|
||
<emphasis id="BEC68339B043A761FF1AFBE1FEB7FBDD" bold="true" box="[199,314,1115,1137]" pageId="15">Etymology.</emphasis>
|
||
The name is derived from the Greek words
|
||
<emphasis id="BEC68339B043A761FCCCFBE1FCC4FBDC" box="[785,841,1115,1136]" italics="true" pageId="15">melos</emphasis>
|
||
, for black, and
|
||
<emphasis id="BEC68339B043A761FF72FBCCFF51FB27" box="[175,220,1142,1163]" italics="true" pageId="15">pous</emphasis>
|
||
, for foot, in reference to the black tarsomeres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B043A761FF1AFB2BFE87FABF" pageId="15" type="distribution">
|
||
<paragraph id="8C0D5F2BB043A761FF1AFB2BFE87FABF" blockId="15.[175,1005,194,1570]" pageId="15">
|
||
<emphasis id="BEC68339B043A761FF1AFB2BFECBFB0B" bold="true" box="[199,326,1169,1191]" pageId="15">Distribution.</emphasis>
|
||
<taxonomicName id="4BB224A8B043A761FE88FB28FDECFB0B" authorityName="Ratcliffe" authorityYear="2008" box="[341,609,1170,1191]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B043A761FE88FB28FDECFB0B" box="[341,609,1170,1191]" italics="true" pageId="15">Cyclocephala melanopoda</emphasis>
|
||
</taxonomicName>
|
||
is known from the southeast and southwest of Santo Domingo de los Colorados on the west side of the Andes (Tinalandia Resort on Hwy 30;
|
||
<collectingRegion id="4E7691C9B043A761FE23FB7DFD4EFB71" box="[510,707,1223,1245]" country="Ecuador" name="Pichincha" pageId="15">Pichincha Province</collectingRegion>
|
||
and Rio Palenque Research
|
||
<collectingRegion id="4E7691C9B043A761FF72FB58FF7EFB54" box="[175,243,1250,1272]" country="France" name="Centre" pageId="15">Center</collectingRegion>
|
||
off of Hwy 25;
|
||
<collectingRegion id="4E7691C9B043A761FE4EFB58FDDDFB54" box="[403,592,1250,1272]" country="Ecuador" name="Los Rios" pageId="15">Los Rios Province</collectingRegion>
|
||
, respectively). The specimens were taken at lights.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B043A77EFF1AFAA3FCB2FA2C" lastPageId="16" pageId="15" type="diagnosis">
|
||
<paragraph id="8C0D5F2BB043A77EFF1AFAA3FE09FC66" blockId="15.[175,1005,194,1570]" lastBlockId="16.[146,975,927,970]" lastPageId="16" pageId="15">
|
||
<emphasis id="BEC68339B043A761FF1AFAA3FEA2FA83" bold="true" box="[199,303,1305,1327]" pageId="15">Diagnosis.</emphasis>
|
||
The parameres are similar to those of
|
||
<taxonomicName id="4BB224A8B043A761FD04FAA3FC14FA83" authority="Endrodi" authorityName="Endrodi" authorityYear="1964" box="[729,921,1305,1327]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B043A761FD04FAA3FCB7FA82" box="[729,826,1305,1326]" italics="true" pageId="15">C. herteli</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BB224A8B043A761FC08FAA3FEFFFAE6" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="1992" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="santaritae">
|
||
<emphasis id="BEC68339B043A761FC08FAA3FE9FFAE5" italics="true" pageId="15">C. santaritae</emphasis>
|
||
Ratcliffe
|
||
</taxonomicName>
|
||
(both from
|
||
<collectingCountry id="F4A51FBBB043A761FE29FA8EFDC6FAE6" box="[500,587,1332,1354]" name="Panama" pageId="15">Panama</collectingCountry>
|
||
).
|
||
<taxonomicName id="4BB224A8B043A761FD83FA8EFCEFFAE5" authorityName="Ratcliffe" authorityYear="2008" box="[606,866,1332,1353]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B043A761FD83FA8EFCEFFAE5" box="[606,866,1332,1353]" italics="true" pageId="15">Cyclocephala melanopoda</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4BB224A8B043A761FC08FA8EFF63FAC8" authorityName="Endrodi" authorityYear="1964" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B043A761FC08FA8EFF63FAC8" italics="true" pageId="15">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
in most external characteristics, except that in
|
||
<taxonomicName id="4BB224A8B043A761FD16FAF5FCD2FAC8" authorityName="Ratcliffe" authorityYear="2008" box="[715,863,1359,1380]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B043A761FD16FAF5FCD2FAC8" box="[715,863,1359,1380]" italics="true" pageId="15">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
the frons and most of the venter is black (testaceous or light brown in
|
||
<taxonomicName id="4BB224A8B043A761FCC5FAD0FCF4FAD3" authorityName="Endrodi" authorityYear="1964" box="[792,889,1386,1407]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B043A761FCC5FAD0FCF4FAD3" box="[792,889,1386,1407]" italics="true" pageId="15">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
), the male pygidium is setose (glabrous in
|
||
<taxonomicName id="4BB224A8B043A761FDC3FA3CFD09FA37" authorityName="Endrodi" authorityYear="1964" box="[542,644,1414,1435]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B043A761FDC3FA3CFD09FA37" box="[542,644,1414,1435]" italics="true" pageId="15">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
), and the tarsomeres are black (testaceous in
|
||
<taxonomicName id="4BB224A8B043A761FEE2FA1AFE11FA19" authorityName="Endrodi" authorityYear="1964" box="[319,412,1440,1461]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B043A761FEE2FA1AFE11FA19" box="[319,412,1440,1461]" italics="true" pageId="15">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
). In females,
|
||
<taxonomicName id="4BB224A8B043A761FDF8FA1AFD37FA19" authorityName="Ratcliffe" authorityYear="2008" box="[549,698,1440,1461]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B043A761FDF8FA1AFD37FA19" box="[549,698,1440,1461]" italics="true" pageId="15">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
has the elytral margin slightly swollen just above the abrupt constriction of the epipleuron, but in
|
||
<taxonomicName id="4BB224A8B043A761FCB5FA06FC48FA7D" authorityName="Endrodi" authorityYear="1964" box="[872,965,1468,1489]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B043A761FCB5FA06FC48FA7D" box="[872,965,1468,1489]" italics="true" pageId="15">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
the elytral margin is slightly swollen but also bi-emarginate. The most distinctive difference is the form of the male 5th protarsomere. In
|
||
<taxonomicName id="4BB224A8B043A761FCCBFA48FC3DF9AB" authorityName="Ratcliffe" authorityYear="2008" box="[790,944,1522,1543]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B043A761FCCBFA48FC3DF9AB" box="[790,944,1522,1543]" italics="true" pageId="15">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
, it is concave ventrally with the ventromedial edge deeply emarginate, and the 236 (caudal and lateral views).
|
||
</paragraph>
|
||
<caption id="D8CD0FA3B05CA77EFF7AFC25FC42FC1E" ID-DOI="http://doi.org/10.5281/zenodo.5369981" ID-Zenodo-Dep="5369981" box="[167,975,927,947]" httpUri="https://zenodo.org/record/5369981/files/figure.png" pageId="16" startId="16.[167,206,927,946]" targetBox="[146,976,194,905]" targetPageId="16">
|
||
<paragraph id="8C0D5F2BB05CA77EFF7AFC25FC42FC1E" blockId="16.[146,975,927,970]" box="[167,975,927,947]" pageId="16">
|
||
<emphasis id="BEC68339B05CA77EFF7AFC25FE95FC1E" bold="true" box="[167,280,927,946]" pageId="16">Figs. 31–33.</emphasis>
|
||
<taxonomicName id="4BB224A8B05CA77EFEF0FC1AFE70FC1F" authorityName="Ratcliffe" authorityYear="2008" box="[301,509,928,947]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="monacha">
|
||
<emphasis id="BEC68339B05CA77EFEF0FC1AFE70FC1F" box="[301,509,928,947]" italics="true" pageId="16">Cyclocephala monacha</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="BEC68339B05CA77EFDD3FC25FDA7FC1E" bold="true" box="[526,554,927,946]" pageId="16">31)</emphasis>
|
||
dorsal view of holotype;
|
||
<emphasis id="BEC68339B05CA77EFCF7FC25FCEAFC1E" bold="true" box="[810,871,927,946]" pageId="16">32–33)</emphasis>
|
||
parameres
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C0D5F2BB05CA77EFF4FFC48FC5DFB91" blockId="16.[146,976,1010,1408]" pageId="16">
|
||
ventrolateral edge strongly produced onto a lobe (
|
||
<figureCitation id="148943AEB05CA77EFD09FC48FCADFBA4" box="[724,800,1010,1032]" captionStart="Figs" captionStartId="14.[167,206,915,934]" captionTargetBox="[146,972,194,892]" captionTargetId="figure-203@14.[146,976,194,892]" captionTargetPageId="14" captionText="Figs. 26–30. Cyclocephala melanopoda: 26) dorsal view of paratype; 27–28) parameres (caudal and lateral views); 29–30) right protarsus, median view, of male of 29) C. melanopoda and 30) C. herteli." figureDoi="http://doi.org/10.5281/zenodo.5369979" httpUri="https://zenodo.org/record/5369979/files/figure.png" pageId="16">Fig. 29</figureCitation>
|
||
). In
|
||
<taxonomicName id="4BB224A8B05CA77EFCBFFC48FC45FBAB" authorityName="Endrodi" authorityYear="1964" box="[866,968,1010,1031]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B05CA77EFCBFFC48FC45FBAB" box="[866,968,1010,1031]" italics="true" pageId="16">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
, conversely, the ventral edges either side of the concavity each have a small ‘‘tooth’’ near the base, and the teeth are subequal in size to one another (
|
||
<figureCitation id="148943AEB05CA77EFCAAFB9DFC48FB91" box="[887,965,1063,1085]" captionStart="Figs" captionStartId="14.[167,206,915,934]" captionTargetBox="[146,972,194,892]" captionTargetId="figure-203@14.[146,976,194,892]" captionTargetPageId="14" captionText="Figs. 26–30. Cyclocephala melanopoda: 26) dorsal view of paratype; 27–28) parameres (caudal and lateral views); 29–30) right protarsus, median view, of male of 29) C. melanopoda and 30) C. herteli." figureDoi="http://doi.org/10.5281/zenodo.5369979" httpUri="https://zenodo.org/record/5369979/files/figure.png" pageId="16">Fig. 30</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB05CA77EFF77FBF9FC45FB55" blockId="16.[146,976,1010,1408]" pageId="16">
|
||
Although the parameres of
|
||
<taxonomicName id="4BB224A8B05CA77EFE0DFBF9FDE4FBF4" authorityName="Ratcliffe" authorityYear="2008" box="[464,617,1091,1112]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B05CA77EFE0DFBF9FDE4FBF4" box="[464,617,1091,1112]" italics="true" pageId="16">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BB224A8B05CA77EFD7FFBF9FCAAFBF4" authorityName="Ratcliffe" authorityYear="1992" box="[674,807,1091,1112]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="santaritae">
|
||
<emphasis id="BEC68339B05CA77EFD7FFBF9FCAAFBF4" box="[674,807,1091,1112]" italics="true" pageId="16">C. santaritae</emphasis>
|
||
</taxonomicName>
|
||
are similar, the similarities end there.
|
||
<taxonomicName id="4BB224A8B05CA77EFEA8FBE4FDF5FBDF" authorityName="Ratcliffe" authorityYear="2008" box="[373,632,1118,1139]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B05CA77EFEA8FBE4FDF5FBDF" box="[373,632,1118,1139]" italics="true" pageId="16">Cyclocephala melanopoda</emphasis>
|
||
</taxonomicName>
|
||
is extensively black on the frons, elytral pattern, either side of the pygidium, legs, and venter, but
|
||
<taxonomicName id="4BB224A8B05CA77EFCF9FBC2FC2BFB21" authorityName="Ratcliffe" authorityYear="1992" box="[804,934,1144,1165]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="santaritae">
|
||
<emphasis id="BEC68339B05CA77EFCF9FBC2FC2BFB21" box="[804,934,1144,1165]" italics="true" pageId="16">C. santaritae</emphasis>
|
||
</taxonomicName>
|
||
has only a faint and different pattern on the elytra. In addition, the pygidium in
|
||
<taxonomicName id="4BB224A8B05CA77EFC6AFB29FE84FB6F" authorityName="Ratcliffe" authorityYear="2008" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B05CA77EFC6AFB29FE84FB6F" italics="true" pageId="16">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
has sparse, small setae, whereas the pygidium in
|
||
<taxonomicName id="4BB224A8B05CA77EFCC3FB14FC2EFB6F" authorityName="Ratcliffe" authorityYear="1992" box="[798,931,1198,1219]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="santaritae">
|
||
<emphasis id="BEC68339B05CA77EFCC3FB14FC2EFB6F" box="[798,931,1198,1219]" italics="true" pageId="16">C. santaritae</emphasis>
|
||
</taxonomicName>
|
||
has dense, long setae. In the females, the swelling on the lateral margin of the elytra is small in
|
||
<taxonomicName id="4BB224A8B05CA77EFF36FB59FE0FFB54" authorityName="Ratcliffe" authorityYear="2008" box="[235,386,1251,1272]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B05CA77EFF36FB59FE0FFB54" box="[235,386,1251,1272]" italics="true" pageId="16">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
, but in
|
||
<taxonomicName id="4BB224A8B05CA77EFE0AFB59FDD4FB54" authorityName="Ratcliffe" authorityYear="1992" box="[471,601,1251,1272]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="santaritae">
|
||
<emphasis id="BEC68339B05CA77EFE0AFB59FDD4FB54" box="[471,601,1251,1272]" italics="true" pageId="16">C. santaritae</emphasis>
|
||
</taxonomicName>
|
||
it is developed into a distinct lobe.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB05CA77EFF77FB45FCB2FA2C" blockId="16.[146,976,1010,1408]" pageId="16">
|
||
<taxonomicName id="4BB224A8B05CA77EFF77FB45FE20FAB8" authorityName="Ratcliffe" authorityYear="2008" box="[170,429,1279,1300]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B05CA77EFF77FB45FE20FAB8" box="[170,429,1279,1300]" italics="true" pageId="16">Cyclocephala melanopoda</emphasis>
|
||
</taxonomicName>
|
||
superficially resembles
|
||
<taxonomicName id="4BB224A8B05CA77EFD7DFB45FC2FFAB9" authority="Hohne" authorityName="Hohne" authorityYear="1923" box="[672,930,1279,1301]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="quadripunctata">
|
||
<emphasis id="BEC68339B05CA77EFD7DFB45FCD8FAB8" box="[672,853,1279,1300]" italics="true" pageId="16">C. quadripunctata</emphasis>
|
||
Höhne
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BB224A8B05CA77EFF4FFAA0FEEFFA83" authority="Endrodi" authorityName="Endrodi" authorityYear="1966" box="[146,354,1305,1327]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="peruana">
|
||
<emphasis id="BEC68339B05CA77EFF4FFAA0FE8FFA83" box="[146,258,1306,1327]" italics="true" pageId="16">C. peruana</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
, but in these two latter species there are short setae on the clypeus and/or pronotum, and the parameres are of different form and more robust than those of
|
||
<taxonomicName id="4BB224A8B05CA77EFEBBFAF5FE76FAC8" authorityName="Ratcliffe" authorityYear="2008" box="[358,507,1359,1380]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="melanopoda">
|
||
<emphasis id="BEC68339B05CA77EFEBBFAF5FE76FAC8" box="[358,507,1359,1380]" italics="true" pageId="16">C. melanopoda</emphasis>
|
||
</taxonomicName>
|
||
. This species will key as far as couplet 190 (
|
||
<taxonomicName id="4BB224A8B05CA77EFC6AFAF5FF5CFAD3" authorityName="Endrodi" authorityYear="1964" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="herteli">
|
||
<emphasis id="BEC68339B05CA77EFC6AFAF5FF5CFAD3" italics="true" pageId="16">C. herteli</emphasis>
|
||
</taxonomicName>
|
||
) in the key to male
|
||
<taxonomicName id="4BB224A8B05CA77EFE78FAD0FDF5FA2C" box="[421,632,1386,1408]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="BEC68339B05CA77EFE78FAD0FDA7FAD3" box="[421,554,1386,1407]" italics="true" pageId="16">Cyclocephala</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="E82322DAB05CA77EFD41FAD0FCB6FA2C" author="Endrodi" box="[668,827,1386,1408]" pageId="16" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |