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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.877.36320" ID-GBIF-Dataset="f9a141b7-89bb-4142-806b-29b01f066e54" ID-PMC="PMC6785569" ID-Pensoft-Pub="1313-2970-877-81" ID-Pensoft-UUID="05447C1FDBD45B16ADA93F21D6FB90E1" ID-PubMed="31616203" ID-ZooBank="F3C1B49B67FE4569AB8A31228CDB6E0C" ModsDocID="1313-2970-877-81" checkinTime="1570026169361" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Borowiec, Lech" docDate="2019" docId="B8DBB77AFD375F93AFAE0F4064C150BD" docLanguage="en" docName="ZooKeys 877: 81-131" docOrigin="ZooKeys 877" docSource="http://dx.doi.org/10.3897/zookeys.877.36320" docTitle="Temnothorax triangularis Salata &amp; Borowiec, 2019, sp. nov." docType="treatment" docVersion="4" id="05447C1FDBD45B16ADA93F21D6FB90E1" lastPageNumber="81" masterDocId="05447C1FDBD45B16ADA93F21D6FB90E1" masterDocTitle="Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species" masterLastPageNumber="131" masterPageNumber="81" pageNumber="81" updateTime="1668167840577" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Preliminary division of not socially parasitic Greek Temnothorax Mayr, 1861 (Hymenoptera, Formicidae) with a description of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salata, Sebastian</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Borowiec, Lech</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>877</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>131</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.877.36320</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.877.36320</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-877-81</mods:identifier>
<mods:identifier type="ZooBank">F3C1B49B67FE4569AB8A31228CDB6E0C</mods:identifier>
<mods:identifier type="Pensoft-UUID">05447C1FDBD45B16ADA93F21D6FB90E1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159619252" LSID="urn:lsid:plazi:treatment:B8DBB77AFD375F93AFAE0F4064C150BD" httpUri="http://treatment.plazi.org/id/B8DBB77AFD375F93AFAE0F4064C150BD" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="B8DBB77A-FD37-5F93-AFAE-0F4064C150BD" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">Temnothorax triangularis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">Figs 16-17</figureCitation>
,
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">18-20</figureCitation>
,
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">21, 22</figureCitation>
,
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">23-24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax triangularis</emphasis>
</taxonomicName>
belongs to the
<taxonomicName lsidName="T. nylanderi" pageId="0" pageNumber="81" rank="species" species="nylanderi">
<emphasis italics="true" pageId="0" pageNumber="81">T. nylanderi</emphasis>
</taxonomicName>
species group. It differs from most of members of this group in uniformly brown body with darker frons and posterior band of first gastral tergite, and absent or very shallow, inconspicuous metanotal groove. There are four other known species with dark body colouration:
<taxonomicName lsidName="T. laconicus" pageId="0" pageNumber="81" rank="species" species="laconicus">
<emphasis italics="true" pageId="0" pageNumber="81">T. laconicus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="T. artvinensis" pageId="0" pageNumber="81" rank="species" species="artvinensis">
<emphasis italics="true" pageId="0" pageNumber="81">T. artvinensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="T. sordidulus" pageId="0" pageNumber="81" rank="species" species="sordidulus">
<emphasis italics="true" pageId="0" pageNumber="81">T. sordidulus</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="T. tergestinus" pageId="0" pageNumber="81" rank="species" species="tergestinus">
<emphasis italics="true" pageId="0" pageNumber="81">T. tergestinus</emphasis>
</taxonomicName>
.
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax triangularis</emphasis>
</taxonomicName>
differs from all of them in extremely shallow metanotal groove and frons lacking reticulation and covered with dense, thick, longitudinal striations, additionally frons centre has sculpture weaker or reduced, it differs also from
<taxonomicName lsidName="T. laconicus" pageId="0" pageNumber="81" rank="species" species="laconicus">
<emphasis italics="true" pageId="0" pageNumber="81">T. laconicus</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="T. artvinensis" pageId="0" pageNumber="81" rank="species" species="artvinensis">
<emphasis italics="true" pageId="0" pageNumber="81">T. artvinensis</emphasis>
</taxonomicName>
in shorter, triangular propodeal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="etymology">
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="81">Named after short, triangular propodeal spines.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Holotype</emphasis>
, worker (pin) (CASENT0846901): GREECE, Sterea Ellas, Eubea | 2.4 km SW of Stropones |
<geoCoordinate degrees="38,60327" direction="north" orientation="latitude" precision="1" value="38.60327">38,60327N</geoCoordinate>
/
<geoCoordinate degrees="23,87" direction="east" orientation="longitude" precision="555" value="23.87">23,87E</geoCoordinate>
, 1025 m |10 VI 2018, L. Borowiec || Collection L. Borowiec |
<taxonomicName family="Formicidae" lsidName="" pageId="0" pageNumber="81" rank="family">Formicidae</taxonomicName>
| LBC-GR02682 (MNHW).
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Paratypes</emphasis>
, 25w., 1Q.(pin) (CASENT0846661-CASENT0846691): the same nest sample as holotype (DBET, BMNH, CASC, MHNG); 5w. (EtOH): the same locality as holotype, collected in litter (DBET); 11w, 1Q (pin) (CASENT0846692-CASENT0846704): GREECE, Sterea Ellas | Eubea |3.7 km SW of Metochi, 830 m |
<geoCoordinate degrees="38.60402" direction="north" orientation="latitude" precision="1" value="38.60402">38.60402N</geoCoordinate>
/
<geoCoordinate degrees="23.91683" direction="east" orientation="longitude" precision="1" value="23.91683">23.91683E</geoCoordinate>
, | 13 VI 2018, L. Borowiec (DBET).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="terra typica">
<paragraph pageId="0" pageNumber="81">Terra typica.</paragraph>
<paragraph pageId="0" pageNumber="81">Euboea, Greece.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="materials_examined">
<paragraph pageId="0" pageNumber="81">Other material.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">GREECE, Sterea Ellas, Euboea</emphasis>
: 2w. (EtOH): 1.5 km SW of Koutourla,
<geoCoordinate degrees="38.62838" direction="north" orientation="latitude" precision="1" value="38.62838">38.62838N</geoCoordinate>
/
<geoCoordinate degrees="23.92772" direction="east" orientation="longitude" precision="1" value="23.92772">23.92772E</geoCoordinate>
, 695 m, 2018-06-13, leg. L. Borowiec; 1w.(pin) (CASENT0846902), 2w. (EtOH): 2.3 km S of Stropones,
<geoCoordinate degrees="38.9933" direction="north" orientation="latitude" precision="5" value="38.9933">38.9933N</geoCoordinate>
/
<geoCoordinate degrees="23.87807" direction="east" orientation="longitude" precision="1" value="23.87807">23.87807E</geoCoordinate>
, 860 m, 2018-06-10, leg. L. Borowiec; 2w. (EtOH): 2.7 km SE of Stropones,
<geoCoordinate degrees="38.59851" direction="north" orientation="latitude" precision="1" value="38.59851">38.59851N</geoCoordinate>
/
<geoCoordinate degrees="23.9085" direction="east" orientation="longitude" precision="5" value="23.9085">23.9085E</geoCoordinate>
, 855 m, 2018-06-13, leg. L. Borowiec; 3w. (EtOH): 2.9 km S of Stropones,
<geoCoordinate degrees="38.59133" direction="north" orientation="latitude" precision="1" value="38.59133">38.59133N</geoCoordinate>
/
<geoCoordinate degrees="23.88562" direction="east" orientation="longitude" precision="1" value="23.88562">23.88562E</geoCoordinate>
, 880 m, 2018-06-13, leg. L. Borowiec; 43w. (EtOH): 3.7 km SW of Metochi,
<geoCoordinate degrees="38.60402" direction="north" orientation="latitude" precision="1" value="38.60402">38.60402N</geoCoordinate>
/
<geoCoordinate degrees="23.91683" direction="east" orientation="longitude" precision="1" value="23.91683">23.91683E</geoCoordinate>
, 830 m, 2018-06-13, leg. L. Borowiec.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Description of worker</emphasis>
(n = 10): HL: 0.684
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 (0.671-0.708); HW: 0.603
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02 (0.578-0.650); SL: 0.504
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02 (0.484-0.534); EL: 0.140
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 (0.124-0.149); EW: 0.105
<normalizedToken originalValue="±">+/-</normalizedToken>
0.006 (0.093-0.112); WL: 0.769
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02 (0.742-0.820); PSL: 0.161
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02 (0.143-0.183); SDL: 0.119
<normalizedToken originalValue="±">+/-</normalizedToken>
0.006 (0.112-0.130); PEL: 0.294
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02 (0.273-0.323); PPL: 0.182
<normalizedToken originalValue="±">+/-</normalizedToken>
0.008 (0.174-0.199); PEH: 0.245
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 (0.236-0.270); PPH: 0.231
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 (0.217-0.248); PNW: 0.403
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 (0.388-0.435); PLW: 0.177
<normalizedToken originalValue="±">+/-</normalizedToken>
0.009 (0.168-0.199); PPW: 0.241
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01 (0.230-0.267); CI: 88.2
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2 (85.3-92.7); SI1: 73.7
<normalizedToken originalValue="±">+/-</normalizedToken>
1.5 (71.2-76.4); SI2: 83.6
<normalizedToken originalValue="±">+/-</normalizedToken>
2.4 (78.9-87.5); MI: 52.4
<normalizedToken originalValue="±">+/-</normalizedToken>
1.2 (50.8-54.7); EI1: 75.0
<normalizedToken originalValue="±">+/-</normalizedToken>
4.2 (68.8-85.0); EI2: 15.3
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0 (13.6-16.7); PI: 120.1
<normalizedToken originalValue="±">+/-</normalizedToken>
5.4 (108.0-128.8); PPI: 79.0
<normalizedToken originalValue="±">+/-</normalizedToken>
2.8 (75.0-85.7); PSI: 135.8
<normalizedToken originalValue="±">+/-</normalizedToken>
12.4 (120.0-155.6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="colour">
<paragraph pageId="0" pageNumber="81">Colour.</paragraph>
<paragraph pageId="0" pageNumber="81">
Whole body uniformly brown to bright brown, sometimes mesosoma and genae brighter. Legs and antennae bright brown to dark yellow, femora in central part darkened (
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">Figs 16</figureCitation>
,
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">17</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="81">Head.</emphasis>
Oval, 1.14 times as long as wide, lateral surfaces below and above eyes gently convex, posterior edges convex, occipital margin of head straight or slightly concave (
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">Figs 18</figureCitation>
,
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">19</figureCitation>
). Anterior margin of the clypeus slightly convex, medial notch absent. Eyes small, oval, 1.33 times as long as wide. Antennal scape short, in lateral view slightly curved, 0.73 times as long as length of the head, in apex gradually widened, its base with small, triangular tooth, funiculus long, club 3-segmented (
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">Fig. 18</figureCitation>
). Surface of scape with very fine microreticulation, shiny, covered with thin, moderate dense, decumbent setae. Mandibles rounded with thick sparse, longitudinal striae, shiny. Clypeus shiny with thick, longitudinal striae, area between striae shiny with few longitudinal wrinkles. Frontal carinae short, not extending beyond frontal lobes. Antennal fossa deep, with sparse, thin roundly curved striae, area between striae with sparse and fine reticulation, shiny. Frontal lobes narrow, smooth with slight, dense longitudinal striation (
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">Figs 18</figureCitation>
,
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">19</figureCitation>
). Space between frontal carinae and vertex with dense, thick, longitudinal striation, sparser or reduced in the central part, striae sometimes interrupted, surface between striae smooth and shiny; space between frontal carinae ad eyes, temples and malar area with longitudinal, thick reticulation, space between reticulation smooth or with very sparse microreticulation, shiny; genae with very sparse thick reticulation, partly smooth, always shiny. Frons and vertex with erect, pale, short and thick setae.
<emphasis bold="true" pageId="0" pageNumber="81">Mesosoma.</emphasis>
Elongate, 1.9 times as long as wide, slightly arched in profile. Metanotal groove absent or very shallow, inconspicuous. Pronotum convex on sides. Propodeal spines short, triangular, with wide base, directed upward, with angulate tips (
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">Fig. 21</figureCitation>
), only in fewer than 20% of specimens are propodeal spines moderately long, with wide bases and sharp tips (
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">Fig. 22</figureCitation>
). Lateral surfaces of pronotum with thick and sparse longitudinal striation or reticulation, its dorsal surface with thick, sparse, irregular reticulation; mesonotum and propodeum on the whole surface with thick, denser than on pronotum, irregular reticulation. Area between thick sculpture shiny, smooth or sometimes with sparse, fine microreticulation (
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">Fig. 17</figureCitation>
). Entire mesosoma bearing erect, pale, short and thick setae (
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">Fig. 17</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="81">Petiole.</emphasis>
In lateral view, with short peduncle, node high, with anterior face flat to slightly convex, and posterior face and dorsum convex. Peduncle and petiolar node shiny, with thick, dense reticulation, area between rugae smooth, dorsum with sparser reticulation. Dorsal surface bearing sparse, short, erect setae (
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">Figs 21</figureCitation>
,
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">22</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="81">Postpetiole.</emphasis>
In lateral view, regularly convex, apical half with gently convex sides (
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">Figs 21</figureCitation>
,
<figureCitation captionStart="Figures 21, 22" captionStartId="F9" captionText="Figures 21, 22. Variation of propodeal spines of Temnothorax triangularis sp. nov." figureDoi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures21-22">22</figureCitation>
), on the whole surface shiny, with thick, dense reticulation, dorsum with sparser reticulation; area between rugae smooth. Dorsal surface bearing sparse, short, semierect to erect setae.
<emphasis bold="true" pageId="0" pageNumber="81">Gaster.</emphasis>
Smooth and shiny, bearing erect, thin, pale setae (
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">Figs 16</figureCitation>
,
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. Worker of Temnothorax triangularis sp. nov. 16 Dorsal 17 Lateral." figureDoi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures16-17">17</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.877.36320.figures16-17" httpUri="https://binary.pensoft.net/fig/344060" pageId="0" pageNumber="81" start="Figures 16, 17" startId="F7">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 16, 17.</emphasis>
Worker of
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax triangularis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="81">16</emphasis>
Dorsal
<emphasis bold="true" pageId="0" pageNumber="81">17</emphasis>
Lateral.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Description of gyne</emphasis>
(n = 2).
<emphasis bold="true" pageId="0" pageNumber="81">Colour.</emphasis>
Head brown, temples slightly brighter coloured than frontal parts. Antennae uniformly yellow. Mesosoma, petiole and postpetiole bright brown, legs yellow. First gastral tergite mostly dark brown with yellowish brown spot basally, remaining tergites brown with dark brown posterior margins (
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">Figs 23</figureCitation>
,
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">24</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="81">Head.</emphasis>
Eyes big, almost round [EL / HW: 0.26]. Antennal scape short [SL / HW: 0.76], not reaching occipital margin of head. Clypeus smooth and shiny laterally with diffused, longitudinal carinulae. Antennal fossa deep, rugulose with concentric carinae. Frontal lobes wide 0.43 times as wide as head width, rugulose with thick longitudinal costae, interstices microreticulate (
<figureCitation captionStart="Figures 18–20" captionStartId="F8" captionText="Figures 18 - 20. Temnothorax triangularis sp. nov. 18 Worker head and antennae 19 Worker head sculpture 20 Gyne head and antennae." figureDoi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures18-20">Fig. 20</figureCitation>
). Frons shiny, entire surface longitudinally costate and rugose, interstices distinctly microreticulate. Area above eyes and sides of head rugulose and partly longitudinally costate, only small area behind eyes smooth and shiny. Entire head bearing erect, pale and thin setae.
<emphasis bold="true" pageId="0" pageNumber="81">Mesosoma.</emphasis>
Pronotum with thick rugosities in anterior part, sides with thick rugosity and dense longitudinal costae. Surface between rugosities microreticulate. Scutum with dense, thick longitudinal costae and microreticulation between costae but appears shiny. Scutellum laterally with thick longitudinal costae, to the centre costae partly diffused, along middle smooth and shiny area (
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">Figs 23</figureCitation>
,
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">24</figureCitation>
). Metanotum with fine longitudinal costae and microreticulated background. Propodeum with area above propodeal spines with transverse and on sides with longitudinal costae and microreticulated background. Propodeal spines short [PSL / HW: 0.28], triangular, with wide base, straight and angulate apex. Area between and below propodeal spines with distinct microreticulation tends to form transverse ridges. Anepisternum and katepisternum with gentle, dense longitudinal costae. Metaepisternum and metakatepisternum, with dense, longitudinal costae and shiny area close to ventral margin. Surface between costae microreticulate. Dorsal surface of mesosoma with sparse, erect, long, thick and pale setae (
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">Figs 23</figureCitation>
,
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">24</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="81">Petiole and postpetiole.</emphasis>
Microreticulate, the entire surface punctate to rugulose, dorsal surface longitudinally costulate.
<emphasis bold="true" pageId="0" pageNumber="81">Gaster.</emphasis>
Smooth and shiny, bearing sparse, long, erect setae (
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">Figs 23</figureCitation>
,
<figureCitation captionStart="Figures 23, 24" captionStartId="F10" captionText="Figures 23, 24. Gyne of Temnothorax triangularis sp. nov. 23 Dorsal 24 Lateral." figureDoi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.877.36320.figures23-24">24</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.877.36320.figures18-20" httpUri="https://binary.pensoft.net/fig/344061" pageId="0" pageNumber="81" start="Figures 18–20" startId="F8">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 18-20.</emphasis>
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax triangularis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="81">18</emphasis>
Worker head and antennae
<emphasis bold="true" pageId="0" pageNumber="81">19</emphasis>
Worker head sculpture
<emphasis bold="true" pageId="0" pageNumber="81">20</emphasis>
Gyne head and antennae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">General distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">Greece, Sterea Ellas, Euboea.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="biology_ecology">
<paragraph pageId="0" pageNumber="81">Biology.</paragraph>
<paragraph pageId="0" pageNumber="81">Alpine species. Ants were observed on stones, dry branches, and herbs in coniferous forest, or coniferous forest with an admixture of chestnut. Nests were found in the dry branches of conifers lying on the ground.</paragraph>
<paragraph pageId="0" pageNumber="81">
The following ant species were recorded in the same areas as
<taxonomicName lsidName="T. triangularis" pageId="0" pageNumber="81" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">T. triangularis</emphasis>
</taxonomicName>
:
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Sterea Ellas, Euboea, 1.5 km SW of Koutourla</emphasis>
:
<taxonomicName class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenogaster subterranea" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis italics="true" pageId="0" pageNumber="81">Aphaenogaster subterranea</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Formica fusca" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="81">Formica fusca</emphasis>
</taxonomicName>
Linnaeus,
<taxonomicName class="Insecta" family="Formicidae" genus="Lasius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasius alienus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="alienus">
<emphasis italics="true" pageId="0" pageNumber="81">Lasius alienus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
,
<taxonomicName lsidName="L. brunneus" pageId="0" pageNumber="81" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="81">L. brunneus</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName lsidName="L. illyricus" pageId="0" pageNumber="81" rank="species" species="illyricus">
<emphasis italics="true" pageId="0" pageNumber="81">L. illyricus</emphasis>
</taxonomicName>
Zimmermann,
<taxonomicName class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole pallidula" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="pallidula">
<emphasis italics="true" pageId="0" pageNumber="81">Pheidole pallidula</emphasis>
</taxonomicName>
(Nylander),
<taxonomicName class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis pygmaea" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis italics="true" pageId="0" pageNumber="81">Plagiolepis pygmaea</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName lsidName="P. pallescens" pageId="0" pageNumber="81" rank="species" species="pallescens">
<emphasis italics="true" pageId="0" pageNumber="81">P. pallescens</emphasis>
</taxonomicName>
Forel,
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax helenae" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="helenae">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax helenae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
et al.,
<taxonomicName lsidName="T. lichtensteini" pageId="0" pageNumber="81" rank="species" species="lichtensteini">
<emphasis italics="true" pageId="0" pageNumber="81">T. lichtensteini</emphasis>
</taxonomicName>
(Bondroit),
<taxonomicName lsidName="T. unifasciatus" pageId="0" pageNumber="81" rank="species" species="unifasciatus">
<emphasis italics="true" pageId="0" pageNumber="81">T. unifasciatus</emphasis>
</taxonomicName>
(Latreille);
<emphasis bold="true" pageId="0" pageNumber="81">2.3 km S of Stropones</emphasis>
: see
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brackoi" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="brackoi">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax brackoi</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="81">2.4 km SW of Stropones</emphasis>
:
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aethiops" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis italics="true" pageId="0" pageNumber="81">Camponotus aethiops</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Formica fusca" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="81">Formica fusca</emphasis>
</taxonomicName>
Linnaeus,
<taxonomicName lsidName="F. sanguinea" pageId="0" pageNumber="81" rank="species" species="sanguinea">
<emphasis italics="true" pageId="0" pageNumber="81">F. sanguinea</emphasis>
</taxonomicName>
Latreille,
<taxonomicName class="Insecta" family="Formicidae" genus="Lasius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasius alienus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="alienus">
<emphasis italics="true" pageId="0" pageNumber="81">Lasius alienus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
,
<taxonomicName lsidName="L. illyricus" pageId="0" pageNumber="81" rank="species" species="illyricus">
<emphasis italics="true" pageId="0" pageNumber="81">L. illyricus</emphasis>
</taxonomicName>
Zimmermann,
<taxonomicName class="Insecta" family="Formicidae" genus="Messor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Messor structor" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="structor">
<emphasis italics="true" pageId="0" pageNumber="81">Messor structor</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax crasecundus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crasecundus">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax crasecundus</emphasis>
</taxonomicName>
Seifert &amp;
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
,
<taxonomicName lsidName="T. unifasciatus" pageId="0" pageNumber="81" rank="species" species="unifasciatus">
<emphasis italics="true" pageId="0" pageNumber="81">T. unifasciatus</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetramorium impurum" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="impurum">
<emphasis italics="true" pageId="0" pageNumber="81">Tetramorium impurum</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
);
<emphasis bold="true" pageId="0" pageNumber="81">2.7 km SE of Stropones</emphasis>
:
<taxonomicName class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenogaster subterranea" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis italics="true" pageId="0" pageNumber="81">Aphaenogaster subterranea</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName lsidName="A. subterranea" pageId="0" pageNumber="81" rank="species" species="subterranea">A. cf. subterranea</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus aethiops" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis italics="true" pageId="0" pageNumber="81">Camponotus aethiops</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName lsidName="C. oertzeni" pageId="0" pageNumber="81" rank="species" species="oertzeni">
<emphasis italics="true" pageId="0" pageNumber="81">C. oertzeni</emphasis>
</taxonomicName>
Forel,
<taxonomicName class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Formica fusca" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="fusca">
<emphasis italics="true" pageId="0" pageNumber="81">Formica fusca</emphasis>
</taxonomicName>
Linnaeus,
<taxonomicName class="Insecta" family="Formicidae" genus="Lasius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasius alienus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="alienus">
<emphasis italics="true" pageId="0" pageNumber="81">Lasius alienus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
,
<taxonomicName lsidName="L. distinguendus" pageId="0" pageNumber="81" rank="species" species="distinguendus">
<emphasis italics="true" pageId="0" pageNumber="81">L. distinguendus</emphasis>
</taxonomicName>
(Emery),
<taxonomicName class="Insecta" family="Formicidae" genus="Tapinoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tapinoma erraticum" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="erraticum">
<emphasis italics="true" pageId="0" pageNumber="81">Tapinoma erraticum</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax helenae" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="helenae">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax helenae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
et al.,
<taxonomicName lsidName="T. unifasciatus" pageId="0" pageNumber="81" rank="species" species="unifasciatus">
<emphasis italics="true" pageId="0" pageNumber="81">T. unifasciatus</emphasis>
</taxonomicName>
(Latreille);
</paragraph>
<caption doi="10.3897/zookeys.877.36320.figures21-22" httpUri="https://binary.pensoft.net/fig/344062" pageId="0" pageNumber="81" start="Figures 21, 22" startId="F9">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 21, 22.</emphasis>
Variation of propodeal spines of
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax triangularis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">2.9 km S of Stropones</emphasis>
:
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus piceus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="piceus">
<emphasis italics="true" pageId="0" pageNumber="81">Camponotus piceus</emphasis>
</taxonomicName>
(Leach),
<taxonomicName lsidName="C. vagus" pageId="0" pageNumber="81" rank="species" species="vagus">
<emphasis italics="true" pageId="0" pageNumber="81">C. vagus</emphasis>
</taxonomicName>
(Scopoli),
<taxonomicName class="Insecta" family="Formicidae" genus="Lasius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasius brunneus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="81">Lasius brunneus</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName lsidName="L. flavus" pageId="0" pageNumber="81" rank="species" species="flavus">
<emphasis italics="true" pageId="0" pageNumber="81">L. flavus</emphasis>
</taxonomicName>
(Fabricius),
<taxonomicName class="Insecta" family="Formicidae" genus="Myrmica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmica scabrinodis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="scabrinodis">
<emphasis italics="true" pageId="0" pageNumber="81">Myrmica scabrinodis</emphasis>
</taxonomicName>
Nylander,
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax helenae" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="helenae">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax helenae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
et al.,
<taxonomicName lsidName="T. unifasciatus" pageId="0" pageNumber="81" rank="species" species="unifasciatus">
<emphasis italics="true" pageId="0" pageNumber="81">T. unifasciatus</emphasis>
</taxonomicName>
(Latreille);
<emphasis bold="true" pageId="0" pageNumber="81">3.7 km SW of Metochi</emphasis>
:
<taxonomicName class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenogaster subterranea" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis italics="true" pageId="0" pageNumber="81">Aphaenogaster subterranea</emphasis>
</taxonomicName>
(Latreille),
<taxonomicName lsidName="A. subterranea" pageId="0" pageNumber="81" rank="species" species="subterranea">A. cf. subterranea</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus fallax" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="fallax">
<emphasis italics="true" pageId="0" pageNumber="81">Camponotus fallax</emphasis>
</taxonomicName>
(Nylander),
<taxonomicName lsidName="C. vagus" pageId="0" pageNumber="81" rank="species" species="vagus">
<emphasis italics="true" pageId="0" pageNumber="81">C. vagus</emphasis>
</taxonomicName>
(Scopoli),
<taxonomicName class="Insecta" family="Formicidae" genus="Lasius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lasius illyricus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="illyricus">
<emphasis italics="true" pageId="0" pageNumber="81">Lasius illyricus</emphasis>
</taxonomicName>
Zimmermann,
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax crasecundus" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="crasecundus">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax crasecundus</emphasis>
</taxonomicName>
Seifert &amp;
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
,
<taxonomicName lsidName="T. helenae" pageId="0" pageNumber="81" rank="species" species="helenae">
<emphasis italics="true" pageId="0" pageNumber="81">T. helenae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
et al.,
<taxonomicName lsidName="T. unifasciatus" pageId="0" pageNumber="81" rank="species" species="unifasciatus">
<emphasis italics="true" pageId="0" pageNumber="81">T. unifasciatus</emphasis>
</taxonomicName>
(Latreille).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="notes">
<paragraph pageId="0" pageNumber="81">Note.</paragraph>
<paragraph pageId="0" pageNumber="81">
Although this species has inconspicuous metanotal groove, which tends to disappear in some specimens, we decided to place it in the
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax nylanderi" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="nylanderi">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax nylanderi</emphasis>
</taxonomicName>
group. General body shape, structure of petiole, unicolourous antennae, head and mesosoma sculpture presented by
<taxonomicName lsidName="T. triangularis" pageId="0" pageNumber="81" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">T. triangularis</emphasis>
</taxonomicName>
are very similar to those observed in large species of the group i.e.
<taxonomicName lsidName="T. nylanderi" pageId="0" pageNumber="81" rank="species" species="nylanderi">
<emphasis italics="true" pageId="0" pageNumber="81">T. nylanderi</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
),
<taxonomicName lsidName="T. crassispinus" pageId="0" pageNumber="81" rank="species" species="crassispinus">
<emphasis italics="true" pageId="0" pageNumber="81">T. crassispinus</emphasis>
</taxonomicName>
(Karavaiev) or
<taxonomicName lsidName="T. crasecundus" pageId="0" pageNumber="81" rank="species" species="crasecundus">
<emphasis italics="true" pageId="0" pageNumber="81">T. crasecundus</emphasis>
</taxonomicName>
Seifert &amp;
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
. Moreover, an inconspicuous metanotal groove was observed also in some samples of small species of the
<taxonomicName genus="Komnina" lsidName="Komnina nylanderi" pageId="0" pageNumber="81" rank="species" species="nylanderi">
<emphasis italics="true" pageId="0" pageNumber="81">nylanderi</emphasis>
</taxonomicName>
group, such as
<taxonomicName lsidName="T. helenae" pageId="0" pageNumber="81" rank="species" species="helenae">
<emphasis italics="true" pageId="0" pageNumber="81">T. helenae</emphasis>
</taxonomicName>
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
, Heinze &amp;
<normalizedToken originalValue="Mikó">Miko</normalizedToken>
. Usually the depth of metanotal groove is more or less constant within nest samples.
</paragraph>
<caption doi="10.3897/zookeys.877.36320.figures23-24" httpUri="https://binary.pensoft.net/fig/344063" pageId="0" pageNumber="81" start="Figures 23, 24" startId="F10">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 23, 24.</emphasis>
Gyne of
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax triangularis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax triangularis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="81">23</emphasis>
Dorsal
<emphasis bold="true" pageId="0" pageNumber="81">24</emphasis>
Lateral.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.877.36320.figures25-27" httpUri="https://binary.pensoft.net/fig/344064" pageId="0" pageNumber="81" start="Figures 25–27" startId="F11">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 25-27.</emphasis>
Distribution in Greece
<emphasis bold="true" pageId="0" pageNumber="81">25</emphasis>
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brackoi" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="brackoi">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax brackoi</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="81">26</emphasis>
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax messiniaensis" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="messiniaensis">
<emphasis italics="true" pageId="0" pageNumber="81">Temnothorax messiniaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="81">27</emphasis>
<taxonomicName lsidName="T. turcicus" pageId="0" pageNumber="81" rank="species" species="turcicus">
<emphasis italics="true" pageId="0" pageNumber="81">T. turcicus</emphasis>
</taxonomicName>
(Santschi) denoted by red circles and
<taxonomicName lsidName="T. triangularis" pageId="0" pageNumber="81" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="81">T. triangularis</emphasis>
</taxonomicName>
sp. nov. by black circles.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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