402 lines
55 KiB
XML
402 lines
55 KiB
XML
<document id="0C33B42F89E90CAAF69407D2A8919091" ID-DOI="10.11646/zootaxa.4019.1.21" ID-GBIF-Dataset="845d040b-01f0-4bd4-a545-e7972311d1d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="233296" ID-ZooBank="4638C91A-F573-4DF1-9E8F-C25C9BBABDDF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461266503283" checkinUser="plazi" docAuthor="Paxton, Hannelore & Budaeva, Nataliya" docDate="2015" docId="B31F87A5FF99BA26C3A2FA7FFB8F9D17" docLanguage="en" docName="zt04019p634.pdf" docOrigin="Zootaxa 4019 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhamphobrachium (Minibrachium) fractum Paxton & Budaeva, 2015, n. sp." docType="treatment" docVersion="8" lastPageNumber="629" masterDocId="4F26FFDDFF9CBA2EC335FFE7FFC89827" masterDocTitle="Minibrachium, a new subgenus of Rhamphobrachium (Annelida: Onuphidae) from Australia with the description of three new species" masterLastPageNumber="634" masterPageNumber="621" pageNumber="626" updateTime="1698610514063" updateUser="plazi">
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<mods:mods id="2EAA9C12C8B07057FEA5CB0A121D7F9E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2E5758185BDD1D19558B9B7C360D684D">
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<mods:title id="594280D1147C6F058DEBD5ED13E3EC36">Minibrachium, a new subgenus of Rhamphobrachium (Annelida: Onuphidae) from Australia with the description of three new species</mods:title>
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<mods:name id="F56BA6D813EEC70220E95960D78F0116" type="personal">
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<mods:roleTerm id="2BDC6A69FEEE00EDE0B4B0AB785A41FD">Author</mods:roleTerm>
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<mods:namePart id="D9E1F415BEB2365BF6170BA6DB728BEF">Paxton, Hannelore</mods:namePart>
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<mods:name id="65F46BCC0DD462E6E8F50FDDB3C9446B" type="personal">
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<mods:roleTerm id="8BC822EEAD327C8753328E4F9C525734">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4C209FDB2ADACFB7CCF0016A79798464">Budaeva, Nataliya</mods:namePart>
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<mods:typeOfResource id="F918777EA5C30085790F50EA88DC2B6C">text</mods:typeOfResource>
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<mods:title id="1D71A5DCD3054EA6301C701F9EA5D7B9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9C7A0972BCE525E2EF2DBC0F441F0126">
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<mods:date id="3F763D3FFA307301BA7EEF055FF597E7">2015</mods:date>
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<mods:detail id="C2AACCACE19705637809CC4E2B856059" type="volume">
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<mods:number id="2052C803BC8EE643878C204A0658AA5D">4019</mods:number>
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<mods:detail id="39FFC0DABC5D09407B71677B5F40CA6A" type="issue">
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<mods:number id="6A4A8C1EFCA4C3EE8E45006D7363D161">1</mods:number>
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<mods:identifier id="4EDD0FD6081A8431BFAD303827D55402" type="DOI">10.11646/zootaxa.4019.1.21</mods:identifier>
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<mods:identifier id="411176AD72C615F61D48AB414B866D99" type="GBIF-Dataset">845d040b-01f0-4bd4-a545-e7972311d1d0</mods:identifier>
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<mods:identifier id="1EC4C61D61A67A1E83EF54B8AC06401D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="B31F87A5FF99BA26C3A2FA7FFB8F9D17" ID-DOI="http://doi.org/10.5281/zenodo.5662758" ID-GBIF-Taxon="119638082" ID-Zenodo-Dep="5662758" LSID="urn:lsid:plazi:treatment:B31F87A5FF99BA26C3A2FA7FFB8F9D17" httpUri="http://treatment.plazi.org/id/B31F87A5FF99BA26C3A2FA7FFB8F9D17" lastPageId="8" lastPageNumber="629" pageId="5" pageNumber="626">
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<subSubSection id="73AC6538FF99BA2BC3A2FA7FFC519E3E" pageId="5" pageNumber="626" type="nomenclature">
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<paragraph id="3B0936B3FF99BA2BC3A2FA7FFD379D94" blockId="5.[151,767,1432,1492]" box="[151,767,1432,1459]" pageId="5" pageNumber="626">
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<heading id="604181DFFF99BA2BC3A2FA7FFD379D94" bold="true" box="[151,767,1432,1459]" fontSize="11" level="1" pageId="5" pageNumber="626" reason="1">
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<emphasis id="09C2EAA1FF99BA2BC3A2FA7FFD379D94" bold="true" box="[151,767,1432,1459]" pageId="5" pageNumber="626">
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<taxonomicName id="FCB64D30FF99BA2BC3A2FA7FFD7D9D95" ID-CoL="4S2LJ" box="[151,693,1432,1459]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="626" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
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<emphasis id="09C2EAA1FF99BA2BC3A2FA7FFE4B9D95" bold="true" box="[151,387,1432,1458]" italics="true" pageId="5" pageNumber="626">Rhamphobrachium</emphasis>
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(
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<emphasis id="09C2EAA1FF99BA2BC2A1FA7FFD8C9D95" bold="true" box="[404,580,1432,1458]" italics="true" pageId="5" pageNumber="626">Minibrachium</emphasis>
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)
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<emphasis id="09C2EAA1FF99BA2BC160FA7FFD7D9D95" bold="true" box="[597,693,1432,1458]" italics="true" pageId="5" pageNumber="626">fractum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="12F157DAFF99BA2BC189FA7EFD379D94" box="[700,767,1433,1459]" pageId="5" pageNumber="626" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="3B0936B3FF99BA2BC3A2FA5CFEC39DF3" blockId="5.[151,767,1432,1492]" box="[151,267,1467,1492]" pageId="5" pageNumber="626">
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(
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<figureCitation id="A38D2A36FF99BA2BC3AAFA5CFF2F9DF3" box="[159,231,1467,1492]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="5" pageNumber="626">Figs 2</figureCitation>
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,
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<figureCitation id="A38D2A36FF99BA2BC3C6FA5CFEC99DF3" box="[243,257,1467,1492]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="5" pageNumber="626">5</figureCitation>
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)
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</paragraph>
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<paragraph id="3B0936B3FF99BA2BC3A2F9E4FC519E3E" blockId="5.[151,921,1539,1561]" box="[151,921,1539,1561]" pageId="5" pageNumber="626">
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||
<treatmentCitationGroup id="1BA6119DFF99BA2BC3A2F9E4FC519E3E" box="[151,921,1539,1561]" pageId="5" pageNumber="626">
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<taxonomicName id="FCB64D30FF99BA2BC3A2F9E4FDFA9E3E" ID-CoL="84T7C" box="[151,562,1539,1561]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="626" phylum="Annelida" rank="genus">
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<emphasis id="09C2EAA1FF99BA2BC3A2F9E4FE919E3E" box="[151,345,1539,1561]" italics="true" pageId="5" pageNumber="626">Rhamphobrachium</emphasis>
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(
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<emphasis id="09C2EAA1FF99BA2BC252F9E4FDE19E3E" box="[359,553,1539,1561]" italics="true" pageId="5" pageNumber="626">Rhamphobrachium</emphasis>
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)
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</taxonomicName>
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sp.—
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<treatmentCitation id="BA1710A2FF99BA2BC159F9E3FCEA9E3E" author="Paxton" box="[620,802,1539,1561]" page="19" pageId="5" pageNumber="626" year="1986">
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<bibRefCitation id="5F274B42FF99BA2BC159F9E3FD359E3E" author="Paxton" box="[620,765,1539,1561]" pageId="5" pageNumber="626" refString="Paxton, H. (1986 a) Generic revision and relationships of the family Onuphidae (Annelida: Polychaeta). Records of the Australian Museum, 38, 1 - 74. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.175" type="journal article" year="1986" yearSuffix="a">Paxton 1986a</bibRefCitation>
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: 19
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</treatmentCitation>
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; 1986b: 87.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="73AC6538FF99BA2BC3A2F9A1FB339F85" pageId="5" pageNumber="626" type="materials_examined">
|
||
<paragraph id="3B0936B3FF99BA2BC3A2F9A1FBA39EEC" blockId="5.[151,1437,1606,1955]" pageId="5" pageNumber="626">
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||
<emphasis id="09C2EAA1FF99BA2BC3A2F9A1FE8C9E78" bold="true" box="[151,324,1606,1631]" pageId="5" pageNumber="626">
|
||
<typeStatus id="E40D8811FF99BA2BC3A2F9A1FF1A9E78" box="[151,210,1606,1631]" pageId="5" pageNumber="626">Type</typeStatus>
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material.
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</emphasis>
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||
<typeStatus id="E40D8811FF99BA2BC27EF9A1FE7F9E78" box="[331,439,1606,1631]" pageId="5" pageNumber="626" type="holotype">Holotype</typeStatus>
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||
: AM W.198965, Victoria,
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||
<quantity id="FC4E9B56FF99BA2BC1E8F9A0FCF89E78" box="[733,816,1606,1631]" metricMagnitude="5" metricUnit="m" metricValue="1.1199999999999999" pageId="5" pageNumber="626" unit="km" value="112.0">112 km</quantity>
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||
S of Lakes Entrance,
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||
<geoCoordinate id="5E825074FF99BA2BC710F9A0FB629E78" box="[1061,1194,1607,1631]" direction="south" orientation="latitude" pageId="5" pageNumber="626" precision="15" value="-39.0">39º00’00”S</geoCoordinate>
|
||
,
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||
<geoCoordinate id="5E825074FF99BA2BC781F9A1FA819E78" box="[1204,1353,1606,1631]" direction="east" orientation="longitude" pageId="5" pageNumber="626" precision="15" value="148.41388">148º24’50”E</geoCoordinate>
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||
, sta. 20 ‘Esso-Gipps’, sand,
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<quantity id="FC4E9B56FF99BA2BC2BBF98DFE1B9EA5" box="[398,467,1642,1667]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="626" unit="m" value="95.0">95 m</quantity>
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||
, coll. C. Phipps,
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||
<date id="4F081073FF99BA2BC18FF98CFCF19EA4" box="[698,825,1642,1667]" pageId="5" pageNumber="626" value="1969-05">May 1969</date>
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||
.
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||
<typeStatus id="E40D8811FF99BA2BC07BF98CFC089EA4" box="[846,960,1643,1667]" pageId="5" pageNumber="626" type="paratype">Paratypes</typeStatus>
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||
: AM W.47833 (2); AM W.47834.001, W.47835.001, W.47836.001,
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||
<specimenCount id="2DB0FD3AFF99BA2BC2D1F968FDB99E80" box="[484,625,1678,1703]" pageId="5" pageNumber="626" type="generic">3 specimens</specimenCount>
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on SEM pins, same data as
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||
<typeStatus id="E40D8811FF99BA2BC0F4F969FBEE9E80" box="[961,1062,1678,1703]" pageId="5" pageNumber="626" type="holotype">holotype</typeStatus>
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||
; AM W.198964 (2), Bass Strait,
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||
<geoCoordinate id="5E825074FF99BA2BC3A2F955FED39EED" box="[151,283,1714,1738]" direction="south" orientation="latitude" pageId="5" pageNumber="626" precision="15" value="-39.0">39º00’00”S</geoCoordinate>
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||
,
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||
<geoCoordinate id="5E825074FF99BA2BC212F955FE739EED" box="[295,443,1714,1738]" direction="east" orientation="longitude" pageId="5" pageNumber="626" precision="15" value="148.5">148º30’00”E</geoCoordinate>
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||
, sta. 19 ‘Esso-Gipps’,
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||
<quantity id="FC4E9B56FF99BA2BC18DF955FCC99EED" box="[696,769,1714,1739]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="5" pageNumber="626" unit="m" value="126.0">126 m</quantity>
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||
, coll. C. Phipps,
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<date id="4F081073FF99BA2BC0F5F955FBAE9EEC" box="[960,1126,1714,1739]" pageId="5" pageNumber="626" value="1969-05-07" valueMax="1969-05-09" valueMin="1969-05-07">7–9 May 1969</date>
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||
.
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||
</paragraph>
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||
<paragraph id="3B0936B3FF99BA2BC3F2F931FB339F85" blockId="5.[151,1437,1606,1955]" pageId="5" pageNumber="626">
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||
<emphasis id="09C2EAA1FF99BA2BC3F2F931FDC19EC8" bold="true" box="[199,521,1750,1775]" pageId="5" pageNumber="626">Other material examined.</emphasis>
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MV
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<accessionNumber id="24E5AB50FF99BA2BC165F931FD679EC8" box="[592,687,1750,1775]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F43626" pageId="5" pageNumber="626" type="EnaNcbi">F43626</accessionNumber>
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||
(1), Tasmania, eastern Bass Strait,
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||
<quantity id="FC4E9B56FF99BA2BC77FF930FB509EC8" box="[1098,1176,1750,1775]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="626" unit="km" value="42.0">42 km</quantity>
|
||
SW of Babel Island,
|
||
<geoCoordinate id="5E825074FF99BA2BC3A2F91DFEC09F35" box="[151,264,1786,1810]" direction="south" orientation="latitude" pageId="5" pageNumber="626" precision="92" value="-40.24">40º14.4’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5E825074FF99BA2BC220F91DFE5D9F35" box="[277,405,1786,1810]" direction="east" orientation="longitude" pageId="5" pageNumber="626" precision="92" value="148.66667">148º40.0’E</geoCoordinate>
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||
, sta. BSS 165 ‘Tangaroa’, fine sand,
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||
<quantity id="FC4E9B56FF99BA2BC077F91DFCB69F35" box="[834,894,1786,1811]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="626" unit="m" value="60.0">60 m</quantity>
|
||
, Smith-McIntyre grab, coll. R. Wilson,
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||
<date id="4F081073FF99BA2BC672F91DFF1A9F10" pageId="5" pageNumber="626" value="1981-11-14">14 Nov 1981</date>
|
||
; MV
|
||
<accessionNumber id="24E5AB50FF99BA2BC22AF8F9FEBC9F10" box="[287,372,1822,1847]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F43628" pageId="5" pageNumber="626" type="EnaNcbi">F43628</accessionNumber>
|
||
(9), Tasmania, eastern Bass Strait,
|
||
<quantity id="FC4E9B56FF99BA2BC012F8F9FCBC9F10" box="[807,884,1822,1847]" metricMagnitude="4" metricUnit="m" metricValue="6.3" pageId="5" pageNumber="626" unit="km" value="63.0">63 km</quantity>
|
||
E of North Point, Flinders Island,
|
||
<geoCoordinate id="5E825074FF99BA2BC612F8F8FA509F10" box="[1319,1432,1823,1847]" direction="south" orientation="latitude" pageId="5" pageNumber="626" precision="92" value="-39.746666">39º44.8’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5E825074FF99BA2BC3A2F8A5FED09F7C" box="[151,280,1858,1883]" direction="east" orientation="longitude" pageId="5" pageNumber="626" precision="92" value="148.67667">148º40.6’E</geoCoordinate>
|
||
, sta. BSS 167 ‘Tangaroa’, muddy sand,
|
||
<quantity id="FC4E9B56FF99BA2BC1DDF8A5FCFB9F7D" box="[744,819,1858,1882]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="626" unit="m" value="124.0">124 m</quantity>
|
||
, Smith-McIntyre grab, coll. R. Wilson,
|
||
<date id="4F081073FF99BA2BC7CBF8A5FA5F9F7D" box="[1278,1431,1858,1882]" pageId="5" pageNumber="626" value="1981-11-14">14 Nov 1981</date>
|
||
; MV
|
||
<accessionNumber id="24E5AB50FF99BA2BC3F9F881FEEA9F58" box="[204,290,1894,1919]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/F43629" pageId="5" pageNumber="626" type="EnaNcbi">F43629</accessionNumber>
|
||
(13), Tasmania, eastern Bass Strait,
|
||
<quantity id="FC4E9B56FF99BA2BC1F6F881FCC49F58" box="[707,780,1894,1919]" metricMagnitude="4" metricUnit="m" metricValue="6.3" pageId="5" pageNumber="626" unit="km" value="63.0">63 km</quantity>
|
||
E of North Point, Flinders Island,
|
||
<geoCoordinate id="5E825074FF99BA2BC7ACF880FAC39F58" box="[1177,1291,1895,1919]" direction="south" orientation="latitude" pageId="5" pageNumber="626" precision="92" value="-39.746666">39º44.8’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5E825074FF99BA2BC622F881FA509F58" box="[1303,1432,1894,1919]" direction="east" orientation="longitude" pageId="5" pageNumber="626" precision="92" value="148.67667">148º40.6’E</geoCoordinate>
|
||
, sta. BSS 167 ‘Tangaroa’, muddy sand,
|
||
<quantity id="FC4E9B56FF99BA2BC179F86DFD5D9F85" box="[588,661,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="626" unit="m" value="124.0">124 m</quantity>
|
||
,
|
||
<collectionCode id="5DA7AE76FF99BA2BC194F86DFD259F85" box="[673,749,1930,1954]" httpUri="http://grbio.org/cool/btw5-epas" name="Woods Hole Oceanographic Institution" pageId="5" pageNumber="626">WHOI</collectionCode>
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||
epibenthic sled, coll. R. Wilson,
|
||
<date id="4F081073FF99BA2BC757F86DFB3F9F85" box="[1122,1271,1930,1954]" pageId="5" pageNumber="626" value="1981-11-14">14 Nov 1981</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6FC9663BFF9ABA28C3A2FB1EFE2A9D4C" httpUri="https://zenodo.org/record/233300/files/figure.png" pageId="6" pageNumber="627" targetBox="[151,1436,193,1252]" targetPageId="6">
|
||
<paragraph id="3B0936B3FF9ABA28C3A2FB1EFE2A9D4C" blockId="6.[151,1436,1273,1387]" pageId="6" pageNumber="627">
|
||
<emphasis id="09C2EAA1FF9ABA28C3A2FB1EFEDC9D28" bold="true" box="[151,276,1273,1295]" pageId="6" pageNumber="627">FIGURE 4.</emphasis>
|
||
Juvenile of
|
||
<taxonomicName id="FCB64D30FF9ABA28C2A1FB1EFC8B9D28" box="[404,835,1273,1295]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="627" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9ABA28C2A1FB1EFD9E9D28" box="[404,598,1273,1295]" italics="true" pageId="6" pageNumber="627">Rhamphobrachium</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF9ABA28C153FB1EFD3F9D28" box="[614,759,1273,1295]" italics="true" pageId="6" pageNumber="627">Minibrachium</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF9ABA28C032FB1DFC8B9D28" box="[775,835,1274,1295]" italics="true" pageId="6" pageNumber="627">nutrix</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF9ABA28C079FB1DFC4D9D28" box="[844,901,1274,1295]" pageId="6" pageNumber="627" rank="species">n. sp.</taxonomicNameLabel>
|
||
, AM W.47875. A. Ventral view; B. Anterior part of body, ventral view; C. Parapodia from chaetigers 3 and 4; D. Parapodium from chaetiger 6; E. Subacicular hooks from chaetiger 9; F. The same from chaetiger 12. Abbreviations: fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, ul = upper lip.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="73AC6538FF9ABA28C3F2FA5BFB1E9E42" pageId="6" pageNumber="627" type="diagnosis">
|
||
<paragraph id="3B0936B3FF9ABA28C3F2FA5BFB1E9E42" blockId="6.[151,1437,1468,1997]" pageId="6" pageNumber="627">
|
||
<emphasis id="09C2EAA1FF9ABA28C3F2FA5BFE899DF2" bold="true" box="[199,321,1468,1493]" pageId="6" pageNumber="627">Diagnosis.</emphasis>
|
||
Small, up to 1.0 mm wide; diagnosis and description based on
|
||
<typeStatus id="E40D8811FF9ABA28C720FA5BFBBF9DF2" box="[1045,1143,1468,1493]" pageId="6" pageNumber="627" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="E40D8811FF9ABA28C785FA5AFAD49DF2" box="[1200,1308,1469,1493]" pageId="6" pageNumber="627" type="paratype">paratypes</typeStatus>
|
||
wider than
|
||
<quantity id="FC4E9B56FF9ABA28C3A2FA06FF269DDF" box="[151,238,1505,1529]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="627" unit="mm" value="0.4">0.4 mm</quantity>
|
||
at chaetiger 10 excluding parapodia. Unidentate distally recurved spiny hooks on chaetigers 1 and 2; four to eight compound to pseudocompound subacicular hooks with spiny hoods on chaetiger 3, five to eight simple subacicular hooks from chaetiger 4, hoods becoming gradually less spiny, resembling typical subacicular hooks by about chaetiger 20; pectinate chaetae with 8–10 long teeth. Pygidium with two pairs of anal cirri.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF9ABA26C3F2F997FE069A1B" lastPageId="8" lastPageNumber="629" pageId="6" pageNumber="627" type="description">
|
||
<paragraph id="3B0936B3FF9ABA28C3F2F997FD9A9ED3" blockId="6.[151,1437,1468,1997]" pageId="6" pageNumber="627">
|
||
<emphasis id="09C2EAA1FF9ABA28C3F2F997FE919EAE" bold="true" box="[199,345,1648,1673]" pageId="6" pageNumber="627">Description.</emphasis>
|
||
<typeStatus id="E40D8811FF9ABA28C257F997FE029EAE" box="[354,458,1648,1673]" pageId="6" pageNumber="627" type="holotype">Holotype</typeStatus>
|
||
incomplete, measuring
|
||
<quantity id="FC4E9B56FF9ABA28C1D4F997FC8F9EAF" box="[737,839,1648,1673]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="627" unit="mm" value="11.5">11.5 mm</quantity>
|
||
in length for 38 chaetigers,
|
||
<quantity id="FC4E9B56FF9ABA28C7A0F996FB269EAF" box="[1173,1262,1649,1673]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="627" unit="mm" value="0.8">0.8 mm</quantity>
|
||
in width; only complete
|
||
<typeStatus id="E40D8811FF9ABA28C230F972FEAD9E8A" box="[261,357,1685,1709]" pageId="6" pageNumber="627" type="paratype">paratype</typeStatus>
|
||
<quantity id="FC4E9B56FF9ABA28C259F972FE739E8B" box="[364,443,1685,1709]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="627" unit="mm" value="10.0">10 mm</quantity>
|
||
long for 58 chaetigers,
|
||
<quantity id="FC4E9B56FF9ABA28C1F3F973FCD39E8B" box="[710,795,1684,1709]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="627" unit="mm" value="0.5">0.5 mm</quantity>
|
||
wide, incomplete
|
||
<typeStatus id="E40D8811FF9ABA28C0DFF972FB9D9E8A" box="[1002,1109,1685,1709]" pageId="6" pageNumber="627" type="paratype">paratypes</typeStatus>
|
||
<quantity id="FC4E9B56FF9ABA28C768F973FB2C9E8B" box="[1117,1252,1684,1709]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="6" pageNumber="627" unit="mm" value="0.7" valueMax="0.9" valueMin="0.5">0.5–0.9 mm</quantity>
|
||
wide. Complete non-type specimens
|
||
<quantity id="FC4E9B56FF9ABA28C2B0F95EFE3C9EF7" box="[389,500,1721,1745]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="2.0" metricValueMin="0.4" pageId="6" pageNumber="627" unit="mm" value="12.0" valueMax="20.0" valueMin="4.0">4–20 mm</quantity>
|
||
long for 35–77 chaetigers, 0.3–1.0 mm wide. Alcohol stored specimens overall cream-coloured, lacking colour pattern.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9ABA28C3F2F8E7FD1E9FEA" blockId="6.[151,1437,1468,1997]" pageId="6" pageNumber="627">
|
||
Prostomium (
|
||
<figureCitation id="A38D2A36FF9ABA28C254F8E7FE6D9F3E" box="[353,421,1792,1817]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="6" pageNumber="627">Fig. 5</figureCitation>
|
||
A) of
|
||
<typeStatus id="E40D8811FF9ABA28C2D8F8E7FD879F3E" box="[493,591,1792,1817]" pageId="6" pageNumber="627" type="holotype">holotype</typeStatus>
|
||
and most
|
||
<typeStatus id="E40D8811FF9ABA28C1FFF8E6FCFD9F3E" box="[714,821,1793,1817]" pageId="6" pageNumber="627" type="paratype">paratypes</typeStatus>
|
||
anteriorly shovel-nosed, two
|
||
<typeStatus id="E40D8811FF9ABA28C7B9F8E6FB3F9F3E" box="[1164,1271,1793,1817]" pageId="6" pageNumber="627" type="paratype">paratypes</typeStatus>
|
||
with indented frontal margin, representing partly formed frontal lips (
|
||
<figureCitation id="A38D2A36FF9ABA28C039F8C3FC999F1A" box="[780,849,1828,1853]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="6" pageNumber="627">Fig. 2</figureCitation>
|
||
C). Ceratophores of palps and antennae with one to two proximal rings and one longer distal ring; ceratostyles short and subulate, palps reaching to peristomium or chaetiger 1, lateral and median antennae reaching to chaetiger 1. Nuchal grooves curved laterally, with narrow middorsal separation. One pair of small eyespots between bases of palps and lateral antennae. Peristomium slightly shorter than first chaetiger; peristomial cirri absent.
|
||
</paragraph>
|
||
<caption id="6FC9663BFF9BBA29C3A2FA71FC029E01" httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="7" pageNumber="628" targetBox="[151,1435,193,1407]" targetPageId="7">
|
||
<paragraph id="3B0936B3FF9BBA29C3A2FA71FC029E01" blockId="7.[151,1436,1430,1574]" pageId="7" pageNumber="628">
|
||
<emphasis id="09C2EAA1FF9BBA29C3A2FA71FEDC9D8B" bold="true" box="[151,276,1430,1452]" pageId="7" pageNumber="628">FIGURE 5.</emphasis>
|
||
<taxonomicName id="FCB64D30FF9BBA29C229FA71FD169D8B" box="[284,734,1430,1452]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="7" pageNumber="628" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9BBA29C229FA71FE169D8B" box="[284,478,1430,1452]" italics="true" pageId="7" pageNumber="628">Rhamphobrachium</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF9BBA29C2DBFA71FDB79D8B" box="[494,639,1430,1452]" italics="true" pageId="7" pageNumber="628">Minibrachium</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF9BBA29C1A5FA71FD169D8B" box="[656,734,1430,1452]" italics="true" pageId="7" pageNumber="628">fractum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF9BBA29C1D3FA70FCE89D8B" box="[742,800,1431,1452]" pageId="7" pageNumber="628" rank="species">n. sp.</taxonomicNameLabel>
|
||
, AM W.47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3–5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3B0936B3FF9BBA29C3F2F990FAE39EDB" blockId="7.[151,1436,1655,2004]" pageId="7" pageNumber="628">
|
||
First two pairs of parapodia modified, projecting anterolaterally, directed ventrally (
|
||
<figureCitation id="A38D2A36FF9BBA29C79EF990FB3B9EB7" box="[1195,1267,1655,1680]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="7" pageNumber="628">Fig. 5</figureCitation>
|
||
A). Neither of parapodia 1 and 2 fully extended, anterior ends inverted, hiding parapodial lobes. Unmodified parapodia lacking distinct parapodial lobes. Dorsal cirri subulate on anterior chaetigers, more posteriorly becoming digitate. Ventral cirri subulate on chaetiger 1 and 2, replaced by rounded ventral pads from chaetiger 3. Branchiae absent.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9BBA29C3F2F8E0FEB29FAB" blockId="7.[151,1436,1655,2004]" pageId="7" pageNumber="628">
|
||
Modified parapodia with three long, pseudocompound recurved hooks (
|
||
<figureCitation id="A38D2A36FF9BBA29C0D3F8E0FBE59F38" box="[998,1069,1799,1824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="7" pageNumber="628">Figs 2</figureCitation>
|
||
D, 5A). Shafts of hooks with two rows of long moveable spines below pseudoarticulation and irregularly distributed small spines along whole length of shaft, but more dense above pseudoarticulation; slight swelling before distal curvature of hook. Internal chaetal sacs to chaetiger 30.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9BBA26C3F2F870FEE699F7" blockId="7.[151,1436,1655,2004]" lastBlockId="8.[151,1437,151,1328]" lastPageId="8" lastPageNumber="629" pageId="7" pageNumber="628">
|
||
Unmodified parapodia from chaetiger 3, supported by two to three very thick aciculae. Limbate chaetae from chaetiger 3, decreasing in length from upper to lower position (
|
||
<figureCitation id="A38D2A36FF9BBA29C06EF85CFC579FF3" box="[859,927,1979,2004]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="7" pageNumber="628">Fig. 5</figureCitation>
|
||
B). Spinigers absent. Four to eight compound (
|
||
<figureCitation id="A38D2A36FF94BA26C3AAFF70FF299897" box="[159,225,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="8" pageNumber="629">Fig. 2</figureCitation>
|
||
E) to pseudocompound (
|
||
<figureCitation id="A38D2A36FF94BA26C2C7FF70FDFC9897" box="[498,564,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="8" pageNumber="629">Fig. 2</figureCitation>
|
||
F) subacicular hooks with spiny hoods on chaetiger 3 (
|
||
<figureCitation id="A38D2A36FF94BA26C7A1FF70FB1D9897" box="[1172,1237,151,176]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="8" pageNumber="629">Fig. 5</figureCitation>
|
||
C) (
|
||
<typeStatus id="E40D8811FF94BA26C635FF70FAAC9897" box="[1280,1380,151,176]" pageId="8" pageNumber="629" type="holotype">holotype</typeStatus>
|
||
with five compound and two pseudocompound hooks); five to eight simple subacicular hooks from chaetiger 4 (
|
||
<figureCitation id="A38D2A36FF94BA26C60BFF5BFA4B98F3" box="[1342,1411,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="8" pageNumber="629">Figs 2</figureCitation>
|
||
G, 5D) (except in juveniles, measuring less than
|
||
<quantity id="FC4E9B56FF94BA26C1A0FF07FD2398D0" box="[661,747,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="629" unit="mm" value="0.3">0.3 mm</quantity>
|
||
in width). Hoods initially enclosing distal part of hooks more closely, gradually opening up, becoming smoother and changing to divided hoods, one on either side of distal part of hook by about chaetiger 20, resembling typical subacicular hooks; concurrent with morphological change, number of hooks becoming gradually reduced to two by chaetiger 20–25 (
|
||
<figureCitation id="A38D2A36FF94BA26C0DFFEABFBE79942" box="[1002,1071,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="8" pageNumber="629">Fig. 5</figureCitation>
|
||
E, F). Slightly oblique pectinate chaetae from chaetiger 13; with 8–10 long teeth, one lateral tooth twice as long, as other teeth (
|
||
<figureCitation id="A38D2A36FF94BA26C7F4FE88FACC99AF" box="[1217,1284,367,392]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="8" pageNumber="629">Fig. 5</figureCitation>
|
||
G). Pygidium with two pairs of anal cirri; dorsal pair longer than ventral one (
|
||
<figureCitation id="A38D2A36FF94BA26C0F6FE73FBC6998A" box="[963,1038,404,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="8" pageNumber="629">Fig. 5</figureCitation>
|
||
H). Fragile mucous tube without incrustations.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF94BA26C3F2FE3BFE069A1B" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="629">Mandibles decalcified, only protomandibles visible. Maxillae very delicate, elements transparent, attachment areas between plates only areas of slight sclerotisation. Maxillary formula: MxI = 1+1; MxII = 5+8; MxIII = 6+0; MxIV = 7+10; MxV = 1+1.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF94BA26C3F2FDA0FF019B82" pageId="8" pageNumber="629" type="discussion">
|
||
<paragraph id="3B0936B3FF94BA26C3F2FDA0FF019B82" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="629">
|
||
<emphasis id="09C2EAA1FF94BA26C3F2FDA0FEF39A47" bold="true" box="[199,315,583,608]" pageId="8" pageNumber="629">Remarks.</emphasis>
|
||
<taxonomicName id="FCB64D30FF94BA26C274FDAFFD649A47" box="[321,684,584,608]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C274FDAFFDD09A47" box="[321,536,584,608]" italics="true" pageId="8" pageNumber="629">Rhamphobrachium</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C112FDAEFD8E9A47" box="[551,582,585,608]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C160FDAFFD649A47" box="[597,684,584,608]" italics="true" pageId="8" pageNumber="629">fractum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C186FDAFFD269A78" box="[691,750,584,607]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="FCB64D30FF94BA26C011FDAEFC0B9A47" box="[804,963,584,608]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="talboti" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C011FDAEFCF49A47" box="[804,828,585,608]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C079FDAEFCA39A47" box="[844,875,585,608]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C04FFDAFFC0B9A47" box="[890,963,584,608]" italics="true" pageId="8" pageNumber="629">talboti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C0FCFDAFFBCD9A78" box="[969,1029,584,607]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
are much more similar to each other than to
|
||
<taxonomicName id="FCB64D30FF94BA26C3DFFD8AFE4C9AA3" box="[234,388,620,644]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C3DFFD8AFECB9AA3" box="[234,259,621,644]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C226FD8AFEF99AA3" box="[275,305,621,644]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C274FD8AFE4C9AA3" box="[321,388,621,644]" italics="true" pageId="8" pageNumber="629">nutrix</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C2BEFD8AFE0F9AA3" box="[395,455,621,644]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
described above. They are almost twice as large as the latter, have unidentate instead of bidentate distally recurved spiny hooks, multiple subacicular hooks from chaetiger 3 instead of only two from chaetiger 5, have two pairs instead of one pair of anal cirri and have pectinate chaetae with much longer teeth. The former two species differ from each other in that the subacicular hooks of chaetiger 3 are always simple in
|
||
<taxonomicName id="FCB64D30FF94BA26C679FD3EFF289B33" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="talboti" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C679FD3EFAAC9AD7" box="[1356,1380,729,752]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C641FD3EFA5B9AD7" box="[1396,1427,729,752]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C3A2FD1BFF289B33" box="[151,224,764,788]" italics="true" pageId="8" pageNumber="629">talboti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C3DDFD1AFEED9B33" box="[232,293,765,788]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="A38D2A36FF94BA26C200FD1BFEB19B32" box="[309,377,764,789]" captionStart="FIGURE 6" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,301,1759]" captionTargetId="figure@9.[151,1436,301,1761]" captionTargetPageId="9" captionText="FIGURE 6. Rhamphobrachium (Minibrachium) talboti n. sp., AM W. 47838.001. A. Anterior part of body, ventral view; B. The same, lateral view; C. Parapodium from chaetiger 3; D. The same, enlarged; E. Parapodium from chaetiger 9; F. The same, enlarged; G. Distal part of recurved hook from chaetiger 1; H – J. Shafts of recurved hooks with movable spines; K. Pectinate chaeta from chaetiger 13. Abbreviations: la = lateral antenna, ll = lower lip, ma = median antenna, mx = maxillae, p = palp, per = peristomium, ul = upper lip, vc = ventral cirrus." httpUri="https://zenodo.org/record/233302/files/figure.png" pageId="8" pageNumber="629">Fig. 6</figureCitation>
|
||
F) but compound to pseudocompound in
|
||
<taxonomicName id="FCB64D30FF94BA26C07BFD1AFC369B33" box="[846,1022,764,788]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C07BFD1AFCAE9B33" box="[846,870,765,788]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C042FD1AFC5E9B33" box="[887,918,765,788]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C092FD1BFC369B33" box="[935,1022,764,788]" italics="true" pageId="8" pageNumber="629">fractum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C733FD1AFB8C9B33" box="[1030,1092,765,788]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="A38D2A36FF94BA26C761FD1BFB5F9B32" box="[1108,1175,764,789]" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="8" pageNumber="629">Fig. 5</figureCitation>
|
||
C). This is a consistent characteristic that is not growth dependent as the smallest
|
||
<taxonomicName id="FCB64D30FF94BA26C011FCC6FC0D9B1F" box="[804,965,800,824]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="talboti" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C011FCC6FCF59B1F" box="[804,829,801,824]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C078FCC6FCA49B1F" box="[845,876,801,824]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C049FCC7FC0D9B1F" box="[892,965,800,824]" italics="true" pageId="8" pageNumber="629">talboti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C0F9FCC7FBC09B10" box="[972,1032,800,823]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
(
|
||
<quantity id="FC4E9B56FF94BA26C72DFCC7FBAF9B10" box="[1048,1127,800,824]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="629" unit="mm" value="11.0">11 mm</quantity>
|
||
for 67 chaetigers,
|
||
<quantity id="FC4E9B56FF94BA26C60DFCF8FA549B10" box="[1336,1436,799,824]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="629" unit="mm" value="0.45">0.45 mm</quantity>
|
||
wide) presented only simple subacicular hooks on chaetiger 3. Furthermore, the pectinate chaetae of
|
||
<taxonomicName id="FCB64D30FF94BA26C7C9FCA2FA549B7B" box="[1276,1436,836,860]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="talboti" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C7C9FCA2FADC9B7B" box="[1276,1300,837,860]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C611FCA2FA8B9B7B" box="[1316,1347,837,860]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C666FCA3FA549B7B" box="[1363,1436,836,860]" italics="true" pageId="8" pageNumber="629">talboti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C3A2FC8FFF1B9B58" box="[151,211,872,895]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
have 9–13 very long teeth (
|
||
<figureCitation id="A38D2A36FF94BA26C13BFC80FD999BA7" box="[526,593,871,896]" captionStart="FIGURE 6" captionStartId="9.[151,250,1782,1804]" captionTargetBox="[151,1436,301,1759]" captionTargetId="figure@9.[151,1436,301,1761]" captionTargetPageId="9" captionText="FIGURE 6. Rhamphobrachium (Minibrachium) talboti n. sp., AM W. 47838.001. A. Anterior part of body, ventral view; B. The same, lateral view; C. Parapodium from chaetiger 3; D. The same, enlarged; E. Parapodium from chaetiger 9; F. The same, enlarged; G. Distal part of recurved hook from chaetiger 1; H – J. Shafts of recurved hooks with movable spines; K. Pectinate chaeta from chaetiger 13. Abbreviations: la = lateral antenna, ll = lower lip, ma = median antenna, mx = maxillae, p = palp, per = peristomium, ul = upper lip, vc = ventral cirrus." httpUri="https://zenodo.org/record/233302/files/figure.png" pageId="8" pageNumber="629">Fig. 6</figureCitation>
|
||
K), while those of
|
||
<taxonomicName id="FCB64D30FF94BA26C016FC8EFC1A9BA7" box="[803,978,872,896]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="629" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF94BA26C016FC8EFCF39BA7" box="[803,827,873,896]" italics="true" pageId="8" pageNumber="629">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF94BA26C079FC8EFCA39BA7" box="[844,875,873,896]" italics="true" pageId="8" pageNumber="629">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF94BA26C04EFC8FFC1A9BA7" box="[891,978,872,896]" italics="true" pageId="8" pageNumber="629">fractum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF94BA26C0EFFC8FFBDE9B58" box="[986,1046,872,895]" pageId="8" pageNumber="629" rank="species">n. sp.</taxonomicNameLabel>
|
||
have 8–10 long teeth instead (
|
||
<figureCitation id="A38D2A36FF94BA26C65AFC80FF6D9B82" captionStart="FIGURE 5" captionStartId="7.[151,250,1430,1452]" captionTargetBox="[151,1435,193,1407]" captionTargetId="figure@7.[151,1437,193,1408]" captionTargetPageId="7" captionText="FIGURE 5. Rhamphobrachium (Minibrachium) fractum n. sp., AM W. 47834.001. A. Anterior part of body, lateral view; B. Parapodia from chaetigers 3 – 5; C. Compound subacicular hooks from chaetiger 3; D. Simple subacicular hooks from chaetiger 4; E. Parapodium from chaetiger 19; F. Simple subacicular hook from chaetiger 23; G. Pectinate chaeta from chaetiger 13; H. Posterior end, dorsolateral view, showing pygidium with anus and anal cirri. Abbreviations: ac = anal cirrus, dc = dorsal cirrus, la = lateral antenna, ma = median antenna, p = palp, vgp = ventral glandular pad." httpUri="https://zenodo.org/record/233301/files/figure.png" pageId="8" pageNumber="629">Fig. 5</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF94BA26C3F2FC48FDC59BCB" pageId="8" pageNumber="629" type="etymology">
|
||
<paragraph id="3B0936B3FF94BA26C3F2FC48FDC59BCB" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="629">
|
||
<emphasis id="09C2EAA1FF94BA26C3F2FC48FE869BEF" bold="true" box="[199,334,943,968]" pageId="8" pageNumber="629">Etymology.</emphasis>
|
||
The name is derived from
|
||
<emphasis id="09C2EAA1FF94BA26C1ADFC57FD2E9BEF" box="[664,742,944,968]" italics="true" pageId="8" pageNumber="629">fractus</emphasis>
|
||
(break) in Latin and refers to the characteristic compound subacicular hooks on chaetiger 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF94BA26C3F2FC10FDB59CCF" pageId="8" pageNumber="629" type="biology_ecology">
|
||
<paragraph id="3B0936B3FF94BA26C3F2FC10FDB59CCF" blockId="8.[151,1437,151,1328]" pageId="8" pageNumber="629">
|
||
<emphasis id="09C2EAA1FF94BA26C3F2FC10FEEF9C37" bold="true" box="[199,295,1015,1040]" pageId="8" pageNumber="629">Biology.</emphasis>
|
||
No brooding specimens were encountered. The smallest specimens in the mixed samples measured
|
||
<quantity id="FC4E9B56FF94BA26C3A2FBFBFF3C9C13" box="[151,244,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="8" pageNumber="629" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
in length for 35–40 chaetigers,
|
||
<quantity id="FC4E9B56FF94BA26C16FFBFBFD799C13" box="[602,689,1052,1076]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="629" unit="mm" value="0.3">0.3 mm</quantity>
|
||
in width and are considered to be recently settled juveniles. These juveniles differ from the larger specimens described above in having compound to pseudocompound subacicular hooks not only on chaetiger 3, but also up to about chaetiger 20, and thereafter only simple ones. In other characteristics they are like larger specimens, except for having even less developed frontal lips and shorter modified parapodia. The distally recurved spiny hooks are not visible externally but are present internally with the chaetal sacs reaching to about chaetiger 20.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF94BA26C3F2FB13FB209D2A" box="[199,1256,1268,1293]" pageId="8" pageNumber="629" type="materials_examined">
|
||
<paragraph id="3B0936B3FF94BA26C3F2FB13FB209D2A" blockId="8.[151,1437,151,1328]" box="[199,1256,1268,1293]" pageId="8" pageNumber="629">
|
||
<emphasis id="09C2EAA1FF94BA26C3F2FB13FEAD9D2A" bold="true" box="[199,357,1268,1293]" pageId="8" pageNumber="629">
|
||
<typeStatus id="E40D8811FF94BA26C3F2FB13FECA9D2A" box="[199,258,1268,1293]" pageId="8" pageNumber="629">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Pacific Ocean,
|
||
<quantity id="FC4E9B56FF94BA26C123FB13FDA29D2A" box="[534,618,1268,1293]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="629" unit="km" value="110.0">110 km</quantity>
|
||
S of Lake Entrance, Victoria;
|
||
<geoCoordinate id="5E825074FF94BA26C08AFB13FB8C9D2B" box="[959,1092,1268,1292]" direction="south" orientation="latitude" pageId="8" pageNumber="629" precision="15" value="-39.0">39º00’00”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5E825074FF94BA26C77AFB13FB2C9D2A" box="[1103,1252,1268,1293]" direction="east" orientation="longitude" pageId="8" pageNumber="629" precision="15" value="148.41388">148º24’50”E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF94BA26C3F2FAF0FB8F9D17" box="[199,1095,1303,1328]" pageId="8" pageNumber="629" type="distribution">
|
||
<paragraph id="3B0936B3FF94BA26C3F2FAF0FB8F9D17" blockId="8.[151,1437,151,1328]" box="[199,1095,1303,1328]" pageId="8" pageNumber="629">
|
||
<emphasis id="09C2EAA1FF94BA26C3F2FAF0FEA89D17" bold="true" box="[199,352,1303,1328]" pageId="8" pageNumber="629">Distribution.</emphasis>
|
||
Bass Strait, Victoria and Tasmania,
|
||
<collectingCountry id="43A17623FF94BA26C1C2FAF0FCA99D17" box="[759,865,1303,1328]" name="Australia" pageId="8" pageNumber="629">Australia</collectingCountry>
|
||
; in
|
||
<quantity id="FC4E9B56FF94BA26C0BCFAF0FC329D08" box="[905,1018,1303,1328]" metricMagnitude="1" metricUnit="m" metricValue="9.3" metricValueMax="12.6" metricValueMin="6.0" pageId="8" pageNumber="629" unit="m" value="93.0" valueMax="126.0" valueMin="60.0">60–126 m</quantity>
|
||
depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |