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<document id="B8D83A3AC39DABF6E9606E5C5A44880C" ID-DOI="10.1080/0022290310001657892" ID-ISSN="1464-5262" ID-Zenodo-Dep="5214577" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629228154861" checkinUser="felipe" docAuthor="Brown, G. R." docDate="2005" docId="734787BFFFAA5A43D92BFCA5FDE2B110" docLanguage="en" docName="JNatHist.39.2.197-239.pdf" docOrigin="Journal of Natural History 39 (2)" docSource="http://dx.doi.org/10.1080/0022290310001657892" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Heligmothynnus sabronensis Brown 2005, comb. nov." docType="treatment" docVersion="4" lastPageNumber="213" masterDocId="8F7EFFC7FFB85A50D81EFF89FF9CB34F" masterDocTitle="A revision of Tachyphron Brown and description of two new genera within the Ariphron group (Hymenoptera: Tiphiidae)" masterLastPageNumber="239" masterPageNumber="197" pageNumber="213" updateTime="1699065075339" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="1D320FD5E3720ED793B814D87990EF9E">A revision of Tachyphron Brown and description of two new genera within the Ariphron group (Hymenoptera: Tiphiidae)</mods:title>
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<mods:namePart id="123DF0177913C9DF2A98DC879C6CA6DC">Brown, G. R.</mods:namePart>
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<mods:title id="C2BCBFDF005404584AF94E574359747C">Journal of Natural History</mods:title>
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<mods:date id="4E4FA2E59D6027E78F7733C10A6DD448">2005</mods:date>
<mods:detail id="FDBC357C3451F034BD463C843D1DE014" type="pubDate">
<mods:number id="AB78EEA73F663B6F2AC8C00AF69CB9DC">2010-12-06</mods:number>
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<mods:number id="822B60CFC0B06FF3A0F71BBA390A836C">39</mods:number>
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<mods:number id="22A922F757F334D90B9E0DF0D83B1C59">2</mods:number>
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<mods:classification id="4A29EF35D4757796338294B8AC415230">journal article</mods:classification>
<mods:identifier id="7AF739C4EF331173084F040FDCE2DA1B" type="DOI">10.1080/0022290310001657892</mods:identifier>
<mods:identifier id="2A692774868BCD02204C5F242D78E087" type="ISSN">1464-5262</mods:identifier>
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<treatment id="734787BFFFAA5A43D92BFCA5FDE2B110" ID-DOI="http://doi.org/10.5281/zenodo.5214737" ID-Zenodo-Dep="5214737" LSID="urn:lsid:plazi:treatment:734787BFFFAA5A43D92BFCA5FDE2B110" httpUri="http://treatment.plazi.org/id/734787BFFFAA5A43D92BFCA5FDE2B110" lastPageId="19" lastPageNumber="213" pageId="18" pageNumber="213">
<subSubSection id="B3F46522FFAA5A42D92BFCA5FC6AB00B" box="[309,1014,812,836]" pageId="18" pageNumber="213" type="nomenclature">
<paragraph id="FB5136A9FFAA5A42D92BFCA5FC6AB00B" blockId="18.[138,1014,812,934]" box="[309,1014,812,836]" pageId="18" pageNumber="213">
<heading id="A01981C5FFAA5A42D92BFCA5FC6AB00B" box="[309,1014,812,836]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="213" reason="2">
<taxonomicName id="3CEE4D2AFFAA5A42D92BFCA5FCF5B00B" authority="(Kimsey 1996)" authorityName="Brown" authorityYear="2005" baseAuthorityName="Kimsey" baseAuthorityYear="1996" box="[309,873,812,836]" class="Insecta" family="Tiphiidae" genus="Heligmothynnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="213" phylum="Arthropoda" rank="species" species="sabronensis" status="comb. nov.">
<emphasis id="C99AEABBFFAA5A42D92BFCA5FD2DB00B" bold="true" box="[309,689,812,836]" italics="true" pageId="18" pageNumber="213">Heligmothynnus sabronensis</emphasis>
(
<bibRefCitation id="9F7F4B58FFAA5A42DADFFCA5FCFDB00B" author="Kimsey LS" box="[705,865,812,836]" pageId="18" pageNumber="213" pagination="303 - 311" refId="ref23386" refString="Kimsey LS. 1996. Re-examination of the Ariphron generic group of thynnine wasps (Hymenoptera: Tiphiidae). Journal of the Australian Entomological Society 35: 303 - 311." type="journal article" year="1996">Kimsey 1996</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="D2A957C0FFAA5A42DB6CFCA5FC6AB00B" box="[882,1014,812,836]" pageId="18" pageNumber="213" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B3F46522FFAA5A42DA0FFCC5FC86B02B" box="[529,794,844,868]" pageId="18" pageNumber="213" type="description">
<paragraph id="FB5136A9FFAA5A42DA0FFCC5FC86B02B" blockId="18.[138,1014,812,934]" box="[529,794,844,868]" pageId="18" pageNumber="213">
(
<figureCitation id="63D52A2CFFAA5A42DA07FCC5FD16B02B" box="[537,650,844,868]" captionStart="Figures 1–4" captionStartId="6.[138,207,905,924]" captionTargetBox="[183,1140,133,869]" captionTargetId="figure-200@6.[183,1140,133,869]" captionTargetPageId="6" captionText="Figures 1–4. Tachyphron spp. and Heligmothynnus sabronensis. (1) T. subtriangularus, male gena. (2) T. aculeatus, male habitus. (3) H. sabronensis, male gena. (4) T. subtriangularus, female habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5214579" httpUri="https://zenodo.org/record/5214579/files/figure.png" pageId="18" pageNumber="213">Figures 3</figureCitation>
,
<figureCitation id="63D52A2CFFAA5A42DA84FCC5FD79B02B" box="[666,741,844,868]" captionStart-0="Figures 17–24" captionStart-1="Figures 25–28" captionStartId-0="14.[138,207,885,904]" captionStartId-1="14.[138,207,1638,1657]" captionTargetBox-0="[183,1138,134,845]" captionTargetBox-1="[182,1137,1318,1599]" captionTargetId-0="figure-128@14.[182,1140,133,849]" captionTargetId-1="figure-100@14.[182,1140,1318,1602]" captionTargetPageId-0="14" captionTargetPageId-1="14" captionText-0="Figures 17–24. Heligmothynnus spp., male genitalia, lateral and dorsal views. (17, 18) H. microspinus. (19, 20) H. atratus. (21, 22) H. neoaratus. (23, 24) H. sabronensis (note: basal ring not shown). Scale bar: 0.25 mm." captionText-1="Figures 25–28. Heligmothynnus spp., male hypopygia. (25) H. sabronensis. (26) H. microspinus. (27) H. atratus. (28) H. neoaratus. Scale bar: 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5214585" figureDoi-1="http://doi.org/10.5281/zenodo.5214587" httpUri-0="https://zenodo.org/record/5214585/files/figure.png" httpUri-1="https://zenodo.org/record/5214587/files/figure.png" pageId="18" pageNumber="213">23–25</figureCitation>
,
<figureCitation id="63D52A2CFFAA5A42DAECFCC5FC8DB02B" box="[754,785,844,868]" captionStart="Figure 59" captionStartId="27.[125,185,910,929]" captionTargetBox="[170,1125,138,870]" captionTargetId="figure-194@27.[170,1128,133,874]" captionTargetPageId="27" captionText="Figure 59. Distribution of Deuterothynnus spp. and Heligmothynnus spp. (note the coincident collecting of D. fulvicentratus with H. microspinus, and D. fulvisetatus with D. parallelus)." figureDoi="http://doi.org/10.5281/zenodo.5214599" httpUri="https://zenodo.org/record/5214599/files/figure.png" pageId="18" pageNumber="213">59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B3F46522FFAA5A42D894FCE4FDF1B0E9" pageId="18" pageNumber="213" type="reference_group">
<paragraph id="FB5136A9FFAA5A42D894FCE4FDEBB0CA" blockId="18.[138,1014,812,934]" box="[138,631,877,901]" pageId="18" pageNumber="213">
<taxonomicName id="3CEE4D2AFFAA5A42D894FCE4FDEFB0CA" authority="Kimsey, 1966: 310" authorityName="Kimsey" authorityPageNumber="310" authorityYear="1966" box="[138,627,877,901]" class="Insecta" family="Tiphiidae" genus="Takyomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="213" phylum="Arthropoda" rank="species" species="sabronensis">
<emphasis id="C99AEABBFFAA5A42D894FCE4FE14B0CA" box="[138,392,877,901]" italics="true" pageId="18" pageNumber="213">Takyomyia sabronensis</emphasis>
Kimsey, 1966: 310
</taxonomicName>
.
</paragraph>
<paragraph id="FB5136A9FFAA5A42D894FC07FDF1B0E9" blockId="18.[138,1014,812,934]" box="[138,621,910,934]" pageId="18" pageNumber="213">
<treatmentCitationGroup id="DBFE1187FFAA5A42D894FC07FDF1B0E9" box="[138,621,910,934]" pageId="18" pageNumber="213">
<treatmentCitation id="7A4F10B8FFAA5A42D894FC07FDF4B0E9" author="Brown GR" box="[138,616,910,934]" page="40" pageId="18" pageNumber="213" year="2001">
<taxonomicName id="3CEE4D2AFFAA5A42D894FC07FDF4B0E9" authority=": Brown, 2001: 40" authorityName="Brown" authorityPageNumber="40" authorityYear="2001" box="[138,616,910,934]" class="Insecta" family="Tiphiidae" genus="Tachyphron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="213" phylum="Arthropoda" rank="species" species="sabronensis">
<emphasis id="C99AEABBFFAA5A42D894FC07FE11B0E9" box="[138,397,910,934]" italics="true" pageId="18" pageNumber="213">Tachyphron sabronensis</emphasis>
:
<bibRefCitation id="9F7F4B58FFAA5A42D980FC07FDF4B0E9" author="Brown GR" box="[414,616,910,934]" pageId="18" pageNumber="213" pagination="23 - 40" refId="ref23101" refString="Brown GR. 2001. Status of the Ariphron generic group (Hymenoptera: Tiphiidae): a critical review. Australian Journal of Entomology 40: 23 - 40." type="journal article" year="2001">Brown, 2001: 40</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B3F46522FFAA5A43D894FC60FDE2B110" lastPageId="19" lastPageNumber="214" pageId="18" pageNumber="213" type="materials_examined">
<paragraph id="FB5136A9FFAA5A42D894FC60FCB9B76E" blockId="18.[138,1184,1001,1057]" pageId="18" pageNumber="213">
<emphasis id="C99AEABBFFAA5A42D894FC60FEFAB74E" box="[138,358,1001,1025]" italics="true" pageId="18" pageNumber="213">Material examined.</emphasis>
<materialsCitation id="4B863CF4FFAA5A42D96CFC60FCB9B76E" collectingDate="1936-05" collectionCode="BMNH" collectorName="L. E. Cheesman" country="Indonesia" elevation="283" location="Sabron" municipality="Cyclops Mountains" pageId="18" pageNumber="213" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="2455880BFFAA5A42D96CFC60FE72B74F" box="[370,494,1001,1025]" pageId="18" pageNumber="213" type="holotype">HOLOTYPE</typeStatus>
:
<emphasis id="C99AEABBFFAA5A42DA1AFC63FD8AB74E" box="[516,534,1002,1025]" italics="true" pageId="18" pageNumber="213">„</emphasis>
.
<emphasis id="C99AEABBFFAA5A42DA35FC60FD34B74E" bold="true" box="[555,680,1001,1025]" pageId="18" pageNumber="213">
<collectingCountry id="83F97639FFAA5A42DA35FC60FD34B74E" box="[555,680,1001,1025]" name="Indonesia" pageId="18" pageNumber="213">Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion id="392AF84BFFAA5A42DAA0FC60FCABB74E" box="[702,823,1001,1025]" country="Indonesia" name="Papua Barat" pageId="18" pageNumber="213">Irian Jaya</collectingRegion>
:
<collectingMunicipality id="1B35ACD3FFAA5A42DB56FC60FBA8B74E" box="[840,1076,1001,1025]" pageId="18" pageNumber="213">Cyclops Mountains</collectingMunicipality>
,
<location id="FE316072FFAA5A42DC58FC60FB00B74E" LSID="urn:lsid:plazi:treatment:734787BFFFAA5A43D92BFCA5FDE2B110:FE316072FFAA5A42DC58FC60FB00B74E" box="[1094,1180,1001,1025]" country="Indonesia" municipality="Cyclops Mountains" name="Sabron" pageId="18" pageNumber="213" stateProvince="Papua Barat">Sabron</location>
,
<quantity id="3C169B4CFFAA5A42D894FB80FF4AB76E" box="[138,214,1033,1057]" metricMagnitude="2" metricUnit="m" metricValue="2.83" pageId="18" pageNumber="213" unit="m" value="283.0">
<elevation id="70C3D19AFFAA5A42D894FB80FF4AB76E" box="[138,214,1033,1057]" metricMagnitude="2" metricUnit="m" metricValue="2.83" pageId="18" pageNumber="213" unit="m" value="283.0">283 m</elevation>
</quantity>
(
<quantity id="3C169B4CFFAA5A42D8F9FB80FEA8B76E" box="[231,308,1033,1057]" metricMagnitude="2" metricUnit="m" metricValue="2.83464" pageId="18" pageNumber="213" unit="ft" value="930.0">
<elevation id="70C3D19AFFAA5A42D8F9FB80FEA8B76E" box="[231,308,1033,1057]" metricMagnitude="2" metricUnit="m" metricValue="2.83464" pageId="18" pageNumber="213" unit="ft" value="930.0">930 ft</elevation>
</quantity>
),
<date id="8F501069FFAA5A42D954FB80FE5AB76E" box="[330,454,1033,1057]" pageId="18" pageNumber="213" value="1936-05">
<collectingDate id="9F14E981FFAA5A42D954FB80FE5AB76E" box="[330,454,1033,1057]" pageId="18" pageNumber="213" value="1936-05">May 1936</collectingDate>
</date>
,
<collectorName id="561B537FFFAA5A42D9CDFB80FD0AB76E" box="[467,662,1033,1057]" pageId="18" pageNumber="213">L. E. Cheesman</collectorName>
, in
<collectionCode id="9DFFAE6CFFAA5A42DADCFB80FCBCB76E" box="[706,800,1033,1057]" pageId="18" pageNumber="213">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="FB5136A9FFAA5A42D894FBECFB11B732" blockId="18.[138,1165,1125,1149]" box="[138,1165,1125,1149]" pageId="18" pageNumber="213">
<emphasis id="C99AEABBFFAA5A42D894FBECFE88B732" box="[138,276,1125,1149]" italics="true" pageId="18" pageNumber="213">Distribution.</emphasis>
Known from the Cyclops Mountains, north-eastern
<collectingRegion id="392AF84BFFAA5A42DB98FBECFC6AB732" box="[902,1014,1125,1149]" country="Indonesia" name="Papua Barat" pageId="18" pageNumber="213">Irian Jaya</collectingRegion>
(
<figureCitation id="63D52A2CFFAA5A42DC19FBECFB1CB732" box="[1031,1152,1125,1149]" captionStart="Figure 59" captionStartId="27.[125,185,910,929]" captionTargetBox="[170,1125,138,870]" captionTargetId="figure-194@27.[170,1128,133,874]" captionTargetPageId="27" captionText="Figure 59. Distribution of Deuterothynnus spp. and Heligmothynnus spp. (note the coincident collecting of D. fulvicentratus with H. microspinus, and D. fulvisetatus with D. parallelus)." figureDoi="http://doi.org/10.5281/zenodo.5214599" httpUri="https://zenodo.org/record/5214599/files/figure.png" pageId="18" pageNumber="213">Figure 59</figureCitation>
).
</paragraph>
<paragraph id="FB5136A9FFAA5A42D894FB49FCFBB616" blockId="18.[138,1185,1216,1369]" pageId="18" pageNumber="213">
<emphasis id="C99AEABBFFAA5A42D894FB49FF60B797" box="[138,252,1216,1240]" italics="true" pageId="18" pageNumber="213">Diagnosis.</emphasis>
Male: hypopygium with lateral margins straight and diverging posteriorly, and the apical spine broadly triangular and as long as the lateral spines. Gena less than quarter length of minimum eye diameter. Fore wing without a preapical spot. Metasoma red-brown to black, but not uniformly black. Dorsal margin of parameres slightly sinusoidal and dentate slightly before apex of basiparameres (
<figureCitation id="63D52A2CFFAA5A42DAB6FAC8FCC6B616" box="[680,858,1345,1369]" captionStart="Figures 17–24" captionStartId="14.[138,207,885,904]" captionTargetBox="[183,1138,134,845]" captionTargetId="figure-128@14.[182,1140,133,849]" captionTargetPageId="14" captionText="Figures 17–24. Heligmothynnus spp., male genitalia, lateral and dorsal views. (17, 18) H. microspinus. (19, 20) H. atratus. (21, 22) H. neoaratus. (23, 24) H. sabronensis (note: basal ring not shown). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5214585" httpUri="https://zenodo.org/record/5214585/files/figure.png" pageId="18" pageNumber="213">Figures 23, 24</figureCitation>
).
</paragraph>
<paragraph id="FB5136A9FFAA5A43D894FA15FDE2B110" blockId="18.[138,1185,1436,1685]" lastBlockId="19.[125,1173,136,607]" lastPageId="19" lastPageNumber="214" pageId="18" pageNumber="213">
<emphasis id="C99AEABBFFAA5A42D894FA15FE90B6FB" box="[138,268,1436,1460]" italics="true" pageId="18" pageNumber="213">Description.</emphasis>
Male: black; anterior margin of pronotum (narrowly interrupted medially) and disc of metanotum yellow; metasoma red-brown to black; tegulae testaceous suffused with yellow basally; wings weakly suffused with orange with slightly darker preapical spot on fore wing; veins and stigma orange; setae white. Clypeus closely and finely punctate, apically truncate, sagittal carina obscure dorsally. Frons closely to rugosely punctate. Vertex rugosely punctate. Gena (
<figureCitation id="63D52A2CFFAA5A42D9AAF9B4FD85B51A" box="[436,537,1597,1621]" captionStart="Figures 1–4" captionStartId="6.[138,207,905,924]" captionTargetBox="[183,1140,133,869]" captionTargetId="figure-200@6.[183,1140,133,869]" captionTargetPageId="6" captionText="Figures 1–4. Tachyphron spp. and Heligmothynnus sabronensis. (1) T. subtriangularus, male gena. (2) T. aculeatus, male habitus. (3) H. sabronensis, male gena. (4) T. subtriangularus, female habitus. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5214579" httpUri="https://zenodo.org/record/5214579/files/figure.png" pageId="18" pageNumber="213">Figure 3</figureCitation>
) finely punctate, narrowly subparallel, less than quarter length of minimum eye diameter. Occipital carina visible dorsally. Genal cavity visible dorsally. Pronotum including anterior surface rugosely punctate. Mesoscutum rugosely punctate. Mesoscutellum closely punctate. Metanotum finely punctate. Propodeum closely to rugosely punctate. Mesopleura finely rugosely punctate.
<collectionCode id="9DFFAE6CFFAB5A43DB38FF21FCA6B38F" box="[806,826,168,192]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="214">T</collectionCode>
1–6 almost impunctate except posterolaterally;
<collectionCode id="9DFFAE6CFFAB5A43D95EFF41FEC8B3AF" box="[320,340,200,224]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="214">T</collectionCode>
7 closely and coarsely punctate, impunctate medially;
<collectionCode id="9DFFAE6CFFAB5A43DBC4FF41FC6DB3AF" box="[986,1009,200,224]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="214">T</collectionCode>
1 with length: width 2.0:1. S1 shallowly and sparsely punctate; S2–8 almost impunctate but becoming punctate posteriorly on posterior segments. Hypopygium (
<figureCitation id="63D52A2CFFAB5A43DB42FE81FC46B26F" box="[860,986,264,288]" captionStart="Figures 25–28" captionStartId="14.[138,207,1638,1657]" captionTargetBox="[182,1137,1318,1599]" captionTargetId="figure-100@14.[182,1140,1318,1602]" captionTargetPageId="14" captionText="Figures 25–28. Heligmothynnus spp., male hypopygia. (25) H. sabronensis. (26) H. microspinus. (27) H. atratus. (28) H. neoaratus. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5214587" httpUri="https://zenodo.org/record/5214587/files/figure.png" pageId="19" pageNumber="214">Figure 25</figureCitation>
) with margins divergent posteriorly, lateral spines short, triangular and divergent, apical spine broadly triangular and as long as lateral spines. Genitalia (
<figureCitation id="63D52A2CFFAB5A43DAC8FEC1FC14B22F" box="[726,904,328,352]" captionStart="Figures 17–24" captionStartId="14.[138,207,885,904]" captionTargetBox="[183,1138,134,845]" captionTargetId="figure-128@14.[182,1140,133,849]" captionTargetPageId="14" captionText="Figures 17–24. Heligmothynnus spp., male genitalia, lateral and dorsal views. (17, 18) H. microspinus. (19, 20) H. atratus. (21, 22) H. neoaratus. (23, 24) H. sabronensis (note: basal ring not shown). Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5214585" httpUri="https://zenodo.org/record/5214585/files/figure.png" pageId="19" pageNumber="214">Figures 23, 24</figureCitation>
) with parameres long, narrow and curved over most of length, dorsal margin emarginate basally, strongly emarginate near apex of basiparameres, apices strongly down-turned, narrowly rounded, divergent, ending at apex of aedeagus (when coiled); basiparameres in dorsal view subtriangular, apex emarginate and ending beyond level of apices of cuspides, broad in lateral view; aedeagus with basal and apical sections, basal section short, swollen and hoodlike with a pair of ventral lobes that originate from inside (rather than the margin of) the basal section, apical section extremely long, filamentous and coiled, much longer than length of parameres. BL: 9; FW: 7; HW: 5.
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</treatment>
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