247 lines
23 KiB
XML
247 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.376.6047" ID-GBIF-Dataset="ea01dfa1-5c94-4a89-804b-aa8037d4dacc" ID-PMC="PMC3935421" ID-Pensoft-Pub="1313-2970-376-1" ID-PubMed="24574849" ID-ZBK="2B58574A5FCF40D19A3AFB4D13D33A92" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-376-1" ModsDocOrigin="ZooKeys 376" ModsDocTitle="Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)" checkinTime="1451246380649" checkinUser="pensoft" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2014" docId="0EB01408DCCB453DABF6CBE673FC7F84" docLanguage="en" docName="ZooKeys 376: 1-83" docOrigin="ZooKeys 376" docSource="http://dx.doi.org/10.3897/zookeys.376.6047" docTitle="Oecetis angularis Blahnik & Holzenthal, 2014, sp. n." docType="treatment" docUuid="CBD87120-82F9-4DF7-9062-40993116FD69" docUuidSource="ZooBank" docVersion="4" lastPageNumber="15" masterDocId="3A31DE5CFF94FFE90671FF839B6AAD3F" masterDocTitle="Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="12" updateTime="1668157599346" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review and redescription of species in the Oecetis avara group, with the description of 15 new species (Trichoptera, Leptoceridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Blahnik, Roger J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Holzenthal, Ralph W.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>376</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.376.6047</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.376.6047</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-376-1</mods:identifier>
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<mods:identifier type="ZBK">2B58574A5FCF40D19A3AFB4D13D33A92</mods:identifier>
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<mods:identifier type="ZooBank">2B58574A5FCF40D19A3AFB4D13D33A92</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050827" LSID="urn:lsid:zoobank.org:act:CBD87120-82F9-4DF7-9062-40993116FD69" httpUri="http://treatment.plazi.org/id/0EB01408DCCB453DABF6CBE673FC7F84" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
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<subSubSection pageId="11" pageNumber="12" type="nomenclature">
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName LSID="http://zoobank.org/CBD87120-82F9-4DF7-9062-40993116FD69" class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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<taxonomicNameLabel pageId="11" pageNumber="12">sp. n.</taxonomicNameLabel>
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Figs 3, 33, 47, Map 1
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph pageId="11" pageNumber="12">Diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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sp. n. is diagnosed by its intermediate size, coloration (yellow, with distinct forewing spots), weakly protruding, but angularly demarcated ventral lobe of the inferior appendage (Fig. 3A), and by having a phallobase with its ventral apex acutely angled (V-shaped), as viewed caudally (Fig. 3C). The ventral margin of the phallobase is arched rather than strongly bent, as viewed laterally. Other species in which the apex of the phallobase is V-shaped in caudal view include
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis metlacensis" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="metlacensis">Oecetis metlacensis</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis uncata" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="uncata">Oecetis uncata</taxonomicName>
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sp. n., and
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis apache" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apache">Oecetis apache</taxonomicName>
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sp. n., all of which are slightly larger in size, darker in color (yellowish-brown or light brown), and have the phallobase more strongly arched or bent as viewed laterally, and
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis acciptrina" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="acciptrina">Oecetis acciptrina</taxonomicName>
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, which is much smaller in size and has the phallobase very strongly bent or hooked near its apex as viewed laterally.
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis metlacensis" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="metlacensis">Oecetis metlacensis</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis apache" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apache">Oecetis apache</taxonomicName>
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additionally differ by having much less prominent forewing spots (absent in
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis apache" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apache">Oecetis apache</taxonomicName>
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).
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</paragraph>
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<paragraph pageId="12" pageNumber="13">
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<pageBreakToken pageId="12" pageNumber="13" start="start">Despite</pageBreakToken>
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the resemblance of the genitalia to the above mentioned species, in overall appearance (size, color and forewing spotting)
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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probably more closely resembles
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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sp. n.,
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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sp. n., or
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis mexicana" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mexicana">Oecetis mexicana</taxonomicName>
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, sp. n., all of which differ in having the apex of the phallobase U-shaped in caudal view (although the shape of the apex may be somewhat difficult to discern in
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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due to its bulbous expansion). A distinction can be made on the basis that the ventral margin of the phallobase is not
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<normalizedToken originalValue="“keeled”">"keeled"</normalizedToken>
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preapically in species described as having a U-shaped apex. The general shape of the inferior appendage in
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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, with a weakly protruding ventral lobe, is most similar to
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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, which is also similar in size;
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis tumida" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tumida">Oecetis tumida</taxonomicName>
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differs in having the ventral margin the phallobase noticeably convexly rounded preapically, as viewed laterally (Fig. 24B), somewhat as in
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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. In Costa Rica, where
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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have been found together,
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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is generally somewhat larger in size than
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis protrusa" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="protrusa">Oecetis protrusa</taxonomicName>
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. In other parts of their ranges, however, size differences seem to be less apparent.
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</paragraph>
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<paragraph pageId="12" pageNumber="13">
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In size, color, and forewing spotting
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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is also nearly identical to
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis punctata" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="punctata">Oecetis punctata</taxonomicName>
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, with which it may co-occur. However, the differences in the shape of the inferior appendages of the males, as well as the arrangement of the crossveins of the chord, as discussed above for characters separating the
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis avara" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="avara">Oecetis avara</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis punctata" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="punctata">Oecetis punctata</taxonomicName>
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groups, should readily distinguish them.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="adult">
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<paragraph pageId="12" pageNumber="13">Adult.</paragraph>
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<paragraph lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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Forewing length: male (9.3-10.8 mm); female (6.3-9.0 mm). Color pale yellowish to brownish-yellow (similar to
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis mexicana" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mexicana">
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<pageBreakToken pageId="13" pageNumber="14" start="start">Oecetis</pageBreakToken>
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mexicana
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</taxonomicName>
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); forewing spots distinct, spots at base of discal and thyridial cells large or moderately large, other spots smaller; veins of forewing chord widely spaced, s and r-m veins slightly closer; chord with small spots at juncture of major veins, spots at opposite ends of chord larger; spots at apices of major veins small, pigmentation confined to vein. Setae along veins in apical part of forewing elongate, semi-prostrate, laterally diverging. Fringe of setae along costal margin of forewing moderate in length, somewhat projecting, not conspicuously erect.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="male genitalia">
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<paragraph pageId="13" pageNumber="14">Male genitalia.</paragraph>
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<paragraph pageId="13" pageNumber="14">Segment IX very short, with elongate setae along posterolateral margin. Preanal appendage moderately elongate, simple in structure, apical setae more elongate. Tergum X with elongate, narrow, deflexed mesal lobe, apex of lobe slightly enlarged, with numerous small sensilla; mesal lobe continuous basoventrally with short, paired lateral membranous or lightly sclerotized projections. Inferior appendage with prominent rounded dorsal process and angularly projecting ventral process; ventral process not strongly projecting, forming an approximate right angle with dorsal process; basomesal projection rounded, moderately prominent, with short, stout mesally directed setae; dorsal process with stout, ventrally curved setae; base of appendage, as viewed ventrally (Fig. 3E), distinctly angled. Phallobase short and tubular basally, with relatively elongate, arched ventral apex; apex, as viewed caudally, with ventral margin sharply V-shaped and slightly keeled (Fig. 3C); phallotremal sclerite elongate, prominent.</paragraph>
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<caption pageId="13" pageNumber="14">
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<paragraph pageId="13" pageNumber="14">
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Figure 3.
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<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
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sp. n., male genitalia: A lateral B phallic apparatus, lateral C phallic apparatus, caudal D inferior appendage, caudal E inferior appendage, ventral.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="holotype">
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<paragraph pageId="13" pageNumber="14">Holotype.</paragraph>
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<paragraph pageId="13" pageNumber="14">
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Male(pinned), COSTA RICA: Puntarenas:
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Bellavista, ca. 1.5 km NW Las Alturas,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.951">8°57.060'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-82.846">82°50.760'W</geoCoordinate>
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, el 1400 m, 16-17.iii.1991, Holzenthal,
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<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
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, Huisman (UMSP) (UMSP000208089).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="paratypes">
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<paragraph pageId="13" pageNumber="14">Paratypes.</paragraph>
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<paragraph lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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GUATEMALA: Suchitepequez: Finca Moca, 12.vi.1966, Flint & Ortiz, 1 male, 1 female (pinned) (NMNH); Puente Ixtacapa, 18-19.vi.1966, Flint & Ortiz, 1 male (pinned)(NMNH); COSTA RICA: Arenal, Quebrada Trondorancito, 24.vii.1967, O. S. Flint, Jr., 13 males, 9 females (pinned) (NMNH); La Suiza, 17.vi.1967, Flint & Ortiz, 1 male (pinned) (NMNH);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Seco, NW Esparta, 28.vii.1967, O. S. Flint, Jr., 1 male (alcohol) (NMNH); Alajuela: Reserva Forestal San Ramon,
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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San Lorencito and tribs.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="10.216">10°12.960'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-84.607">84°36.420'W</geoCoordinate>
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, el 980 m, 30.iii.-1.iv.1987, Holzenthal, Hamilton, Heyn, 1 male (alcohol) (UMSP);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Pizote, ca. 5 km N Dos
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<normalizedToken originalValue="Ríos">Rios</normalizedToken>
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,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="10.948">10°56.880'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-85.291">85°17.460'W</geoCoordinate>
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, el 470 m, 9.iii.1986, Holzenthal & Fasth, 1 male (pinned), 16 males, 15 females (alcohol) (UMSP); Cartago:
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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<normalizedToken originalValue="Reventazón">Reventazon</normalizedToken>
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in CATIE along Sendero Espaveles,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.893">9°53.580'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.651">83°39.060'W</geoCoordinate>
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, el 500 m, 22.iii.1991,
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<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
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, 5 males, 2 females (pinned), 1 male (alcohol) (UMSP);
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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:
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Banano, 16 km WSW Bomba,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.871333">9°52.280'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.167">83°10.020'W</geoCoordinate>
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, el 150 m, 26.iii.1987, Holzenthal, Hamilton, Heyn, 2 males, 2 females (pinned) (UMSP);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Telire and small trib., SE Suretka,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.554">9°33.240'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-82.892">82°53.520'W</geoCoordinate>
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, el 48 m, 1.ii.1986, 1 male, 1 female (pinned) (UMSP); Res. Biol. Barbilla,
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Dantas,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.994">9°59.640'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.443">83°26.580'W</geoCoordinate>
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, el 300 m, 27-30.i.1992, Holzenthal,
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<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
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, Kjer, 7 males (alcohol) (UMSP); Puntarenas: same data as holotype, 15 males, 8 females (pinned) (UMSP);
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|
<normalizedToken originalValue="Cotón">Coton</normalizedToken>
|
|
in Las Alturas,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.938">8°56.280'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-82.826">82°49.560'W</geoCoordinate>
|
|
, el 1360 m, 18.iii.1991, Holzenthal
|
|
<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
|
|
, Huisman, 18 males, 7 females (pinned), 10 males, 18 females (alcohol) (UMSP); same location, 12.viii.1990, Holzenthal, Blahnik,
|
|
<normalizedToken originalValue="Muñoz">Munoz</normalizedToken>
|
|
, 7 males, 1 female (pinned), 8 males (alcohol) (UMSP);
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|
Jaba, 2.4 km (air) NW San Vito,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.832">8°49.920'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-82.991">82°59.460'W</geoCoordinate>
|
|
, el 970 m, 13.vi.1986, Holzenthal, Heyn, Armitage, 1 male (pinned), 1 male, 1 female (alcohol) (UMSP);
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|
<pageBreakToken pageId="14" pageNumber="15" start="start">Ceibo</pageBreakToken>
|
|
, route 2, ca. 6 km W road to Buenos Aires,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.149">9°8.940'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.377">83°22.620'W</geoCoordinate>
|
|
, el 250 m, 20.ii.1986, Holzenthal, Morse, Fasth, 1 male (alcohol) (UMSP);
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|
Singri, ca. 2 km (air) S Finca Helechales,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="9.057">9°3.420'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.082">83°4.920'W</geoCoordinate>
|
|
, el 720 m, 21.ii.1986, Holzenthal, Morse, Fasth, 10 males, 20 females (alcohol) (UMSP); ECUADOR: Cotopaxi: Latacunga, 133 km W, 2.vii.1975, el 1080 m, A. Langley & J. Cohen, 1 male (pinned) (NMNH); Quevedo, 36 km NE, el 335 m, 2.vii.1976, J. Cohen, 9 males, 4 females (alcohol) (NMNH); Loja:
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|
Puyango, el 300 m, 17-18.viii.1977, L. E.
|
|
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
|
G., 2 males, 1 female (pinned) (NMNH); Pichincha: Santo Domingo de los Colorados, 14. km E, 5.vii.1975, Langley & Cohen, 1 male (pinned) (NMNH).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="14" pageNumber="15" type="etymology">
|
|
<paragraph pageId="14" pageNumber="15">Etymology.</paragraph>
|
|
<paragraph pageId="14" pageNumber="15">
|
|
This species is named
|
|
<taxonomicName class="Insecta" family="Leptoceridae" genus="Oecetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oecetis angularis" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="angularis">Oecetis angularis</taxonomicName>
|
|
for the acutely angled ventral apex of the phallobase, as seen in caudal view.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |