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<document id="8F1740C0BEDABB90C31B06CB53110B15" ID-CLB-Dataset="22067" ID-DOI="10.5852/ejt.2017.337" ID-GBIF-Dataset="5a7f73c5-5646-446f-a72c-3ea18d8f634f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832483" ID-ZooBank="2669CB69-5064-4B5D-AC25-856680E3B9AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589830819450" checkinUser="carolina" docAuthor="Budha, Prem B., Naggs, Fred &amp; Backeljau, Thierry" docDate="2017" docId="03C6CF41FFF1FFFDFE164196D439FAEF" docLanguage="en" docName="ejt-2017-337.pdf.imf" docOrigin="European Journal of Taxonomy 337" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Diplommatina syabrubesiensis Budha &amp; Naggs &amp; Backeljau 2017, sp. nov." docType="treatment" docUuid="1CFD0C16-38F3-489C-9A86-945A6475250B" docUuidSource="ZooBank" docVersion="6" lastPageNumber="23" masterDocId="FFFFB739FFE4FFEBFFAE4762D552FF83" masterDocTitle="The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="22" updateTime="1698840088040" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="9CB74D9AE1600AA1B58560E7101B418E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AE0A970F3A8F27676EBE442F539582A7">
<mods:title id="B273388BDD3861BF004866243DE07B71">The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species</mods:title>
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<mods:name id="630CB27C7A99F90823FCA049D426B04A" type="personal">
<mods:role id="69DBCFF2F02312CE7BA2FE31FB76ABE7">
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<mods:namePart id="6E8A7AC9E6F88695FE09521C14661A68">Budha, Prem B.</mods:namePart>
<mods:affiliation id="F19657311D1127D6A8F74F418377210B">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Central Department of Zoology, Tribhuvan University, Kirtipur, Kathmandu, Nepal. &amp; Corresponding author: prembudha @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="C0738FE15EE13358930E24FB650A5AAD" type="email">prembudha@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="6DD51C2DFD86F582BA5856096C7147B8" type="personal">
<mods:role id="66692B3B13F1B83F71B5E8F74D7871B0">
<mods:roleTerm id="3D4D84288DA7110D21E767515D74249B">Author</mods:roleTerm>
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<mods:namePart id="05CD528B3A08D67A773F71BE000B70A8">Naggs, Fred</mods:namePart>
<mods:affiliation id="F1EB010217BAA9BDE213A4E9EAD857C3">Natural History Museum, Cromwell Road, London, SW 7 5 BD, UK. &amp; Email: f. naggs @ nhm. ac. uk</mods:affiliation>
<mods:nameIdentifier id="147F79DB1D0F41292B6B6559B8770F10" type="email">f.naggs@nhm.ac.uk</mods:nameIdentifier>
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<mods:name id="74F4641BC37BA898E2850A9D18A58874" type="personal">
<mods:role id="99F8B95285857D129329CC6628780CAB">
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<mods:namePart id="B4A045209E65490D56C162AFA70F6493">Backeljau, Thierry</mods:namePart>
<mods:affiliation id="8A42E736A3A6E09E19B47A0F4713F188">University of Antwerp, Evolutionary Ecology Group, Universiteitsplein 1, B- 2610, Antwerp, Belgium. &amp; Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B- 1000, Brussels, Belgium. &amp; Email: thierry. backeljau @ naturalsciences. be</mods:affiliation>
<mods:nameIdentifier id="5851F890595A483B006B8935EF579A46" type="email">thierry.backeljau@naturalsciences.be</mods:nameIdentifier>
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<mods:typeOfResource id="37976C55D557D068DAD6655FFB9079C8">text</mods:typeOfResource>
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<mods:title id="250B17605340B3D62372A2A6CBEB6378">European Journal of Taxonomy</mods:title>
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<mods:part id="43E56DB992F0FB17C138ABC003208B17">
<mods:date id="189C805F4981A4A1C8D13B081E7A3DAF">2017</mods:date>
<mods:detail id="A62544E2D46690CDA17FC1B49C19BE2D" type="pubDate">
<mods:number id="07BC385ADD370224A4AA8C4F71F2E35F">2017-07-21</mods:number>
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<mods:start id="52E36B5BC928D85346A51EBF4CDC59EE">1</mods:start>
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<mods:classification id="3DA1FF08F0270AB005BEEA4DFB516914">journal article</mods:classification>
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<mods:identifier id="58758840C17B8C1219441C672007209B" type="DOI">10.5852/ejt.2017.337</mods:identifier>
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<treatment id="03C6CF41FFF1FFFDFE164196D439FAEF" ID-DOI="http://doi.org/10.5281/zenodo.3850686" ID-GBIF-Taxon="164167125" ID-Zenodo-Dep="3850686" LSID="urn:lsid:zoobank.org:act:1CFD0C16-38F3-489C-9A86-945A6475250B" httpUri="http://treatment.plazi.org/id/03C6CF41FFF1FFFDFE164196D439FAEF" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="C3752DDCFFF1FFFEFE164196D12EF88C" box="[440,1148,1780,1807]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BD07E57FFF1FFFEFE164196D12EF88C" blockId="21.[372,1216,1780,1878]" box="[440,1148,1780,1807]" pageId="21" pageNumber="22">
<heading id="D098C93BFFF1FFFEFE164196D12EF88C" box="[440,1148,1780,1807]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="22" reason="2">
<taxonomicName id="4C6F05D4FFF1FFFEFE164196D14BF88D" ID-CoL="36M46" authority="Budha &amp; Backeljau" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[440,1049,1780,1807]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="syabrubesiensis" status="sp. nov.">
<emphasis id="B91BA245FFF1FFFEFE164196D679F88C" bold="true" box="[440,811,1780,1807]" italics="true" pageId="21" pageNumber="22">Diplommatina syabrubesiensis</emphasis>
Budha &amp; Backeljau
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF1FFFEFB8F4197D12EF88C" box="[1057,1148,1781,1807]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3752DDCFFF1FFFEFEDA407AD633F8D5" pageId="21" pageNumber="22" type="description">
<paragraph id="8BD07E57FFF1FFFEFEDA407AD192F8B0" blockId="21.[372,1216,1780,1878]" box="[372,1216,1816,1843]" pageId="21" pageNumber="22">
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urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="8BD07E57FFF1FFFEFD7C405ED633F8D5" blockId="21.[372,1216,1780,1878]" box="[722,865,1852,1878]" pageId="21" pageNumber="22">
<figureCitation id="135462D2FFF1FFFEFD7C405ED663F8D5" box="[722,817,1852,1878]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="21" pageNumber="22">Figs 8C</figureCitation>
,
<figureCitation id="135462D2FFF1FFFEFC93405ED633F8D5" box="[829,865,1852,1878]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="21" pageNumber="22">9O</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF1FFFDFF13401CD662FDE8" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BD07E57FFF1FFFEFF13401CD464F81B" blockId="21.[189,310,1918,1944]" box="[189,310,1918,1944]" pageId="21" pageNumber="22">
<heading id="D098C93BFFF1FFFEFF13401CD464F81B" bold="true" box="[189,310,1918,1944]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B91BA245FFF1FFFEFF13401CD464F81B" bold="true" box="[189,310,1918,1944]" pageId="21" pageNumber="22">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFF1FFFDFF1340CFD662FDE8" blockId="21.[189,1399,1965,2027]" lastBlockId="22.[189,1399,267,619]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Measurements (n = 11): SH
<quantity id="4C97D3B2FFF1FFFEFDB740CFD7FFF844" box="[537,685,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="21" pageNumber="22" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 mm</quantity>
, SW
<quantity id="4C97D3B2FFF1FFFEFD5C40CFD6D7F844" box="[754,901,1965,1992]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="21" pageNumber="22" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
, Wh 6.5–7.0;
<typeStatus id="54D4C0F5FFF1FFFEFB9D40CFD1CCF844" box="[1075,1182,1965,1991]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
: SH
<quantity id="4C97D3B2FFF1FFFEFB7240CFD06EF844" box="[1244,1340,1965,1991]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="21" pageNumber="22" unit="mm" value="3.2">3.2 mm</quantity>
, SW
<quantity id="4C97D3B2FFF1FFFEFF1340B2D449F868" box="[189,283,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="22" unit="mm" value="1.5">1.5 mm</quantity>
, Wh 7.0. Dextral, whorls regularly increasing, penultimate whorl slightly wider than the body whorl, suture impressed, constriction nearly at the middle in front of the parietal wall (
<figureCitation id="135462D2FFF2FFFDFB614669D07DFEA6" box="[1231,1327,267,293]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="22" pageNumber="23">Fig. 9O</figureCitation>
). One weak parietalis (
<figureCitation id="135462D2FFF2FFFDFE2D464DD4B0FEC9" box="[387,482,303,330]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="22" pageNumber="23">Fig. 8C</figureCitation>
<figureCitation id="135462D2FFF2FFFDFE4F4652D4A2FEC9" box="[481,496,304,330]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[189,1393,274,1888]" captionTargetId="figure@6.[189,1398,269,1893]" captionTargetPageId="6" captionText="Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832490" httpUri="https://zenodo.org/record/3832490/files/figure.png" pageId="22" pageNumber="23">3</figureCitation>
). One vertical palatalis. Horizontal palatalis absent (
<figureCitation id="135462D2FFF2FFFDFBC9464DD19AFEC9" box="[1127,1224,303,330]" captionStart="Fig" captionStartId="24.[189,232,1789,1815]" captionTargetBox="[297,1297,294,1724]" captionTargetId="figure@24.[247,1340,265,1748]" captionTargetPageId="24" captionText="Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3832502" httpUri="https://zenodo.org/record/3832502/files/figure.png" pageId="22" pageNumber="23">Fig. 9O</figureCitation>
). Columellaris moderate, weak towards the aperture. Protoconch smooth, with minute pits evenly distributed, radial ribs absent (
<figureCitation id="135462D2FFF2FFFDFEE5461AD4FBFE11" box="[331,425,376,402]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="22" pageNumber="23">Fig. 8C</figureCitation>
<figureCitation id="135462D2FFF2FFFDFE07461AD4E6FE11" box="[425,436,376,402]" captionStart="Fig" captionStartId="2.[189,235,1999,2025]" captionTargetBox="[189,1399,1129,1958]" captionTargetId="figure@2.[187,1401,1126,1960]" captionTargetPageId="2" captionText="Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal." figureDoi="http://doi.org/10.5281/zenodo.3832486" httpUri="https://zenodo.org/record/3832486/files/figure.png" pageId="22" pageNumber="23">1</figureCitation>
). Teleoconch with distinct radial ribs; 9–11 ribs/
<quantity id="4C97D3B2FFF2FFFDFC44461AD111FE11" box="[1002,1091,376,403]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="0.5">0.5 mm</quantity>
on the penultimate whorl; 13–16 ribs/
<quantity id="4C97D3B2FFF2FFFDFEED46FED4CFFE35" box="[323,413,412,439]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="23" unit="mm" value="0.5">0.5 mm</quantity>
on the body whorl towards the aperture. Dorsal fold present, but weakly developed (
<figureCitation id="135462D2FFF2FFFDFF6846A2D473FE59" box="[198,289,448,474]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="22" pageNumber="23">Fig. 8C</figureCitation>
<figureCitation id="135462D2FFF2FFFDFE8E46A2D462FE59" box="[288,304,448,474]" captionStart="Fig" captionStartId="4.[189,232,1895,1921]" captionTargetBox="[189,1398,285,1853]" captionTargetId="figure@4.[189,1398,285,1853]" captionTargetPageId="4" captionText="Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two)." figureDoi="http://doi.org/10.5281/zenodo.3832488" httpUri="https://zenodo.org/record/3832488/files/figure.png" pageId="22" pageNumber="23">2</figureCitation>
, C
<figureCitation id="135462D2FFF2FFFDFEFC46A2D431FE59" box="[338,355,448,474]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="22" pageNumber="23">4</figureCitation>
). Spiral striation present, but hardly visible. Umbilicus closed. Aperture circular; parietal wall thick, raised and slightly oblique with regard to the coiling axis. Peristome with two slightly spaced lips; angular edge of the peristome straight; palatal lip rounded; basal lip along the columellar margin slightly curved; inner lip slightly thickened and forming a thick callus over the parietal wall. Operculum paucispiral, Wh 2.5, outer whorl comparatively wide.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF2FFFDFF1345F1D60DFD5F" box="[189,863,659,733]" pageId="22" pageNumber="23" type="etymology">
<paragraph id="8BD07E57FFF2FFFDFF1345F1D416FD2E" blockId="22.[189,324,659,685]" box="[189,324,659,685]" pageId="22" pageNumber="23">
<heading id="D098C93BFFF2FFFDFF1345F1D416FD2E" bold="true" box="[189,324,659,685]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B91BA245FFF2FFFDFF1345F1D416FD2E" bold="true" box="[189,324,659,685]" pageId="22" pageNumber="23">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFF2FFFDFF1345A0D60DFD5F" blockId="22.[189,863,706,733]" box="[189,863,706,733]" pageId="22" pageNumber="23">
The species name refers to the
<typeStatus id="54D4C0F5FFF2FFFDFD8645A1D709FD5E" box="[552,603,707,733]" pageId="22" pageNumber="23">type</typeStatus>
locality ‘Syabrubesi’.
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF2FFFDFF134467D7B6FBE8" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BD07E57FFF2FFFDFF134467D43EFC9C" blockId="22.[189,364,773,799]" box="[189,364,773,799]" pageId="22" pageNumber="23">
<heading id="D098C93BFFF2FFFDFF134467D43EFC9C" bold="true" box="[189,364,773,799]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B91BA245FFF2FFFDFF134467D43EFC9C" bold="true" box="[189,364,773,799]" pageId="22" pageNumber="23">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B07740AFFF2FFFDFF444457D7BAFC1B" ID-GBIF-Occurrence="2620517303" collectingDate="2008-05-13" collectionCode="CDZMTU" collectorName="P. B. Budha" country="Nepal" county="Langtang National Park" elevation="2635" latitude="28.197563" location="Syabrubesi" longLatPrecision="1" longitude="85.45512" municipality="Lama Hotel" pageId="22" pageNumber="23" specimenCode="CDZMTU051" specimenCount="1" typeStatus="holotype">
<paragraph id="8BD07E57FFF2FFFDFF444457D40EFCCC" blockId="22.[189,1399,821,920]" box="[234,348,821,847]" pageId="22" pageNumber="23">
<emphasis id="B91BA245FFF2FFFDFF444457D40EFCCC" bold="true" box="[234,348,821,847]" pageId="22" pageNumber="23">
<typeStatus id="54D4C0F5FFF2FFFDFF444457D40EFCCC" box="[234,348,821,847]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFF2FFFDFF13443BD7BAFC1B" blockId="22.[189,1399,821,920]" pageId="22" pageNumber="23">
<collectingCountry id="F3783EC7FFF2FFFDFF13443BD44CFCF0" box="[189,286,857,883]" name="Nepal" pageId="22" pageNumber="23">NEPAL</collectingCountry>
:
<location id="8EB0288CFFF2FFFDFE87443BD4FCFCF0" LSID="urn:lsid:plazi:treatment:03C6CF41FFF1FFFDFE164196D439FAEF:8EB0288CFFF2FFFDFE87443BD4FCFCF0" box="[297,430,857,883]" country="Nepal" county="Langtang National Park" latitude="28.197563" longLatPrecision="1" longitude="85.45512" municipality="Lama Hotel" name="Syabrubesi" pageId="22" pageNumber="23">Syabrubesi</location>
, above
<collectingMunicipality id="6BB4E42DFFF2FFFDFDA9443BD7C4FCF0" box="[519,662,857,883]" pageId="22" pageNumber="23">Lama Hotel</collectingMunicipality>
,
<collectingCounty id="62B106DBFFF2FFFDFD0C443BD6ECFCF0" box="[674,958,857,883]" pageId="22" pageNumber="23">Langtang National Park</collectingCounty>
,
<quantity id="4C97D3B2FFF2FFFDFC64443AD177FCF0" box="[970,1061,856,883]" metricMagnitude="3" metricUnit="m" metricValue="2.635" pageId="22" pageNumber="23" unit="m" value="2635.0">
<elevation id="00429964FFF2FFFDFC64443AD177FCF0" box="[970,1061,856,883]" metricMagnitude="3" metricUnit="m" metricValue="2.635" pageId="22" pageNumber="23" unit="m" value="2635.0">2635 m</elevation>
</quantity>
,
<geoCoordinate id="EE5B1890FFF2FFFDFB9C443AD19CFCF0" box="[1074,1230,856,883]" degrees="28.197564" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="28.197563">28.197564 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFF2FFFDFB74443AD023FCF0" box="[1242,1393,856,883]" degrees="85.455117" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="85.45512">85.455117 E</geoCoordinate>
,
<date id="FFD15897FFF2FFFDFF13441FD40FFC14" box="[189,349,893,919]" pageId="22" pageNumber="23" value="2008-05-13">
<collectingDate id="EF95A17FFFF2FFFDFF13441FD40FFC14" box="[189,349,893,919]" pageId="22" pageNumber="23" value="2008-05-13">13 May 2008</collectingDate>
</date>
,
<collectorName id="269A1B81FFF2FFFDFEC6441FD4BEFC14" box="[360,492,893,919]" pageId="22" pageNumber="23">P.B. Budha</collectorName>
leg. (
<specimenCode id="DBC9D62CFFF2FFFDFD83441FD78EFC1B" box="[557,732,893,920]" collectionCode="CDZMTU" pageId="22" pageNumber="23">CDZMTU051</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BD07E57FFF2FFFDFF4444DDD435FC5A" blockId="22.[189,1399,959,1131]" box="[234,359,959,985]" pageId="22" pageNumber="23">
<emphasis id="B91BA245FFF2FFFDFF4444DDD435FC5A" bold="true" box="[234,359,959,985]" pageId="22" pageNumber="23">
<typeStatus id="54D4C0F5FFF2FFFDFF4444DDD435FC5A" box="[234,359,959,985]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD07E57FFF2FFFDFF134486D7B6FBE8" blockId="22.[189,1399,959,1131]" pageId="22" pageNumber="23">
<collectingCountry id="F3783EC7FFF2FFFDFF134486D44CFC7D" box="[189,286,996,1022]" name="Nepal" pageId="22" pageNumber="23">NEPAL</collectingCountry>
: 7 shells from the type locality, same data as
<typeStatus id="54D4C0F5FFF2FFFDFCE54481D6E0FC7E" box="[843,946,995,1021]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7EE692FFF2FFFDFC6A4481D114FC7D" box="[964,1094,995,1022]" pageId="22" pageNumber="23">CDZMTU</collectionCode>
052); 27 shells, Dhunche, Langtang National Park,
<quantity id="4C97D3B2FFF2FFFDFE774365D762FBA1" box="[473,560,1031,1058]" metricMagnitude="3" metricUnit="m" metricValue="2.056" pageId="22" pageNumber="23" unit="m" value="2056.0">2056m</quantity>
,
<geoCoordinate id="EE5B1890FFF2FFFDFD96436AD783FBA1" box="[568,721,1032,1058]" degrees="28.108942" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="28.108942">28.108942 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFF2FFFDFD774365D63FFBA1" box="[729,877,1031,1058]" degrees="85.291547" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="85.29155">85.291547E</geoCoordinate>
,
<date id="FFD15897FFF2FFFDFCD8436AD6A8FBA1" box="[886,1018,1032,1058]" pageId="22" pageNumber="23" value="2007-06-03">3Jun. 2007</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFF2FFFDFB154365D06CFBA1" box="[1211,1342,1031,1058]" pageId="22" pageNumber="23">CDZMTU</collectionCode>
053); 2 shells, Baghdwar-Deurali trek, Shivapuri-Nagarjun National Park,
<quantity id="4C97D3B2FFF2FFFDFC734349D165FBC5" box="[989,1079,1067,1094]" metricMagnitude="3" metricUnit="m" metricValue="2.4059999999999997" pageId="22" pageNumber="23" unit="m" value="2406.0">2406 m</quantity>
,
<geoCoordinate id="EE5B1890FFF2FFFDFBED4349D18CFBC5" box="[1091,1246,1067,1094]" degrees="27.803775" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="27.803776">27.803775 N</geoCoordinate>
,
<geoCoordinate id="EE5B1890FFF2FFFDFB474349D023FBC5" box="[1257,1393,1067,1094]" degrees="85.38872" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="85.38872">85.38872 E</geoCoordinate>
,
<date id="FFD15897FFF2FFFDFF134332D40BFBE9" box="[189,345,1104,1131]" pageId="22" pageNumber="23" value="2008-04-25">25 Apr. 2008</date>
, P.B. Budha leg. (
<collectionCode id="ED7EE692FFF2FFFDFD874332D7F9FBE8" box="[553,683,1104,1131]" pageId="22" pageNumber="23">CDZMTU</collectionCode>
054).
</paragraph>
</subSubSection>
<subSubSection id="C3752DDCFFF2FFFDFF1343F0D439FAEF" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BD07E57FFF2FFFDFF1343F0D47DFB2F" blockId="22.[189,303,1170,1196]" box="[189,303,1170,1196]" pageId="22" pageNumber="23">
<heading id="D098C93BFFF2FFFDFF1343F0D47DFB2F" bold="true" box="[189,303,1170,1196]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B91BA245FFF2FFFDFF1343F0D47DFB2F" bold="true" box="[189,303,1170,1196]" pageId="22" pageNumber="23">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD07E57FFF2FFFDFF1343A0D439FAEF" blockId="22.[189,1399,1218,1388]" pageId="22" pageNumber="23">
The shell of
<taxonomicName id="4C6F05D4FFF2FFFDFEE143A1D77EFB5F" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[335,556,1218,1244]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="syabrubesiensis" status="sp. nov.">
<emphasis id="B91BA245FFF2FFFDFEE143A1D77EFB5F" box="[335,556,1218,1244]" italics="true" pageId="22" pageNumber="23">D. syabrubesiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF2FFFDFD9D43A0D7DFFB5F" box="[563,653,1218,1244]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is similar to that of
<taxonomicName id="4C6F05D4FFF2FFFDFCD843A1D6ABFB5F" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[886,1017,1218,1244]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="fistulata" status="sp. nov.">
<emphasis id="B91BA245FFF2FFFDFCD843A1D6ABFB5F" box="[886,1017,1218,1244]" italics="true" pageId="22" pageNumber="23">D. fistulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF2FFFDFC5143A0D10BFB5F" box="[1023,1113,1218,1244]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
However, the peristome of
<taxonomicName id="4C6F05D4FFF2FFFDFF754385D409FA83" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[219,347,1254,1280]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="fistulata" status="sp. nov.">
<emphasis id="B91BA245FFF2FFFDFF754385D409FA83" box="[219,347,1254,1280]" italics="true" pageId="22" pageNumber="23">D. fistulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF2FFFDFECE4384D4EBFA83" box="[352,441,1254,1280]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
is highly tilted towards the left on the inner wall of the parietal lip on the columellar axis, while it is round in
<taxonomicName id="4C6F05D4FFF2FFFDFE4C4269D792FAA7" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[482,704,1290,1316]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="syabrubesiensis" status="sp. nov.">
<emphasis id="B91BA245FFF2FFFDFE4C4269D792FAA7" box="[482,704,1290,1316]" italics="true" pageId="22" pageNumber="23">D. syabrubesiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF2FFFDFD664268D676FAA7" box="[712,804,1290,1316]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
Moreover,
<taxonomicName id="4C6F05D4FFF2FFFDFC034269D163FAA7" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[941,1073,1290,1316]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="fistulata" status="sp. nov.">
<emphasis id="B91BA245FFF2FFFDFC034269D163FAA7" box="[941,1073,1290,1316]" italics="true" pageId="22" pageNumber="23">D. fistulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF2FFFDFB964268D1C1FAA7" box="[1080,1171,1290,1316]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
has a strong dorsal fold, which is not distinct in
<taxonomicName id="4C6F05D4FFF2FFFDFDB5424DD7A9FACB" authority="Budha &amp; Naggs &amp; Backeljau, 2017" authorityName="Budha &amp; Naggs &amp; Backeljau" authorityYear="2017" box="[539,763,1326,1352]" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="syabrubesiensis" status="sp. nov.">
<emphasis id="B91BA245FFF2FFFDFDB5424DD7A9FACB" box="[539,763,1326,1352]" italics="true" pageId="22" pageNumber="23">D. syabrubesiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2281F3EFFF2FFFDFCAA424DD609FACA" box="[772,859,1327,1353]" pageId="22" pageNumber="23" rank="species">sp. nov</taxonomicNameLabel>
and only visible through careful observation (see
<figureCitation id="135462D2FFF2FFFDFF5A4230D41FFAEF" box="[244,333,1362,1388]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[190,1398,279,1890]" captionTargetId="figure@20.[190,1398,276,1893]" captionTargetPageId="20" captionText="Fig. 8. A. Diplommatina silvicola Godwin-Austen, 1886 (CDZMTU050). B. D. sperata W.T. Blanford, 1862 (CDZMTU062). C. D. syabrubesiensis sp. nov., holotype (CDZMTU051). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832500" httpUri="https://zenodo.org/record/3832500/files/figure.png" pageId="22" pageNumber="23">Fig. 8C</figureCitation>
<figureCitation id="135462D2FFF2FFFDFEE34230D40CFAEF" box="[333,350,1362,1388]" captionStart="Fig" captionStartId="9.[189,232,1930,1956]" captionTargetBox="[189,1398,274,1888]" captionTargetId="figure@9.[189,1398,266,1891]" captionTargetPageId="9" captionText="Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3832492" httpUri="https://zenodo.org/record/3832492/files/figure.png" pageId="22" pageNumber="23">4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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