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<document id="CAE161A4855719AEB546BAD4021F321F" ID-CLB-Dataset="29024" ID-DOI="10.11646/zootaxa.4459.2.2" ID-GBIF-Dataset="634e23da-84c4-46a4-8bb8-94fcf1637160" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458548" ID-ZooBank="73510C3B-6482-4760-8173-B6E701839940" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288809625" checkinUser="plazi" docAuthor="Sheth, Sayali D., Ghate, Hemant V. &amp; Hájek, Jiří" docDate="2018" docId="03BC3747FF97FF9FFF5E12BEFB88F831" docLanguage="en" docName="zootaxa.4459.2.2.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus indicus Sharp 1882" docType="treatment" docVersion="7" lastPageNumber="247" masterDocId="FF854F3FFF9BFF93FFC91143FFF8FFA8" masterDocTitle="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="260" masterPageNumber="235" pageNumber="247" updateTime="1698578320975" updateUser="ExternalLinkService">
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<mods:title id="BA4E036D1BEB7A3A3096A73C58E3F23F">Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="3AA93E4BA145DAF76841AC47BAB29200">Sheth, Sayali D.</mods:namePart>
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<mods:namePart id="BC42BC7CC4CD212F1B044AFE2D48B790">Ghate, Hemant V.</mods:namePart>
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<mods:namePart id="7D24F2E18E380CD0D43FC2D99BE246BA">Hájek, Jiří</mods:namePart>
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<mods:title id="8284E477E5FD4B245178C399ADACA41F">Zootaxa</mods:title>
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<mods:part id="8D892B886B524E26DD445AD8A7F4DC91">
<mods:date id="C70223ADCEBCB34A94C2D37DAB98EB91">2018</mods:date>
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<mods:number id="B26CCBBEC00CC4034CAC0AD0A21C6386">2018-08-15</mods:number>
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<mods:number id="59BC66A98FEACFE69C19211E76ECE8D8">4459</mods:number>
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<mods:identifier id="34E7E8C9D59E09C56641093A48F5D807" type="CLB-Dataset">29024</mods:identifier>
<mods:identifier id="BF68158FA65E186343E5BBFFF6B35976" type="DOI">10.11646/zootaxa.4459.2.2</mods:identifier>
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<treatment id="03BC3747FF97FF9FFF5E12BEFB88F831" ID-DOI="http://doi.org/10.5281/zenodo.5966533" ID-GBIF-Taxon="148692674" ID-Zenodo-Dep="5966533" LSID="urn:lsid:plazi:treatment:03BC3747FF97FF9FFF5E12BEFB88F831" httpUri="http://treatment.plazi.org/id/03BC3747FF97FF9FFF5E12BEFB88F831" lastPageNumber="247" pageId="12" pageNumber="247">
<subSubSection id="C30FD5DAFF97FF9FFF5E12BEFEA7FB91" pageId="12" pageNumber="247" type="nomenclature">
<paragraph id="8BAA8651FF97FF9FFF5E12BEFDF4FBBF" blockId="12.[151,524,1021,1081]" box="[151,524,1021,1047]" pageId="12" pageNumber="247">
<heading id="D0E2313DFF97FF9FFF5E12BEFDF4FBBF" bold="true" box="[151,524,1021,1047]" fontSize="11" level="1" pageId="12" pageNumber="247" reason="1">
<taxonomicName id="4C15FDD2FF97FF9FFF5E12BEFDF4FBBF" ID-CoL="Y39K" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[151,524,1021,1047]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9615A43FF97FF9FFF5E12BEFDF4FBBF" bold="true" box="[151,524,1021,1047]" pageId="12" pageNumber="247">
<emphasis id="B9615A43FF97FF9FFF5E12BEFE97FBBF" bold="true" box="[151,367,1021,1047]" italics="true" pageId="12" pageNumber="247">Copelatus indicus</emphasis>
<bibRefCitation id="EF84FBA0FF97FF9FFEBF12BEFDF4FBBF" box="[374,524,1021,1047]" pageId="12" pageNumber="247" refId="ref16437" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAA8651FF97FF9FFF5E1563FEA7FB91" blockId="12.[151,524,1021,1081]" box="[151,351,1056,1081]" pageId="12" pageNumber="247">
(
<figureCitation id="132E9AD4FF97FF9FFF561563FEFCFB90" box="[159,260,1056,1081]" captionStart="FIGURES 5–8" captionStartId="4.[151,263,1970,1993]" captionTargetBox="[174,1372,207,1947]" captionTargetId="figure@4.[151,1435,193,1949]" captionTargetPageId="4" captionText="FIGURES 5–8. Habitus of Copelatus. 5–6. C. bezdeki sp. nov.: 5. male holotype; 6. female paratype (Lonavala); 7–8. C. indicus: 7. male (Lonavala); 8. female (Mahabaleshwar)." httpUri="https://zenodo.org/record/1458552/files/figure.png" pageId="12" pageNumber="247">Figs 7–8</figureCitation>
,
<figureCitation id="132E9AD4FF97FF9FFEC61563FEAFFB91" box="[271,343,1056,1081]" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="12" pageNumber="247">25–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF97FF9FFF5E1520FDF0FBD1" box="[151,520,1123,1145]" pageId="12" pageNumber="247" type="reference_group">
<paragraph id="8BAA8651FF97FF9FFF5E1520FDF0FBD1" blockId="12.[151,520,1123,1145]" box="[151,520,1123,1145]" pageId="12" pageNumber="247">
<treatmentCitationGroup id="AB05A17FFF97FF9FFF5E1520FDF0FBD1" box="[151,520,1123,1145]" pageId="12" pageNumber="247">
<taxonomicName id="4C15FDD2FF97FF9FFF5E1520FDFCFBD1" ID-CoL="Y39K" authority="Sharp, 1882: 582" authorityName="Sharp" authorityPageNumber="582" authorityYear="1882" box="[151,516,1123,1145]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9615A43FF97FF9FFF5E1520FEB5FBD1" box="[151,333,1123,1145]" italics="true" pageId="12" pageNumber="247">Copelatus indicus</emphasis>
<treatmentCitation id="0AB4A040FF97FF9FFE9A1520FDFCFBD1" author="" box="[339,516,1123,1145]" page="582" pageId="12" pageNumber="247">
<bibRefCitation id="EF84FBA0FF97FF9FFE9A1520FE29FBD1" box="[339,465,1123,1145]" pageId="12" pageNumber="247" refId="ref16437" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article">Sharp, 1882</bibRefCitation>
: 582
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF97FF9FFF5E15E1FA7AF9E3" pageId="12" pageNumber="247" type="materials_examined">
<paragraph id="8BAA8651FF97FF9FFF5E15E1FE6CFB13" blockId="12.[151,1437,1186,1945]" box="[151,404,1186,1211]" pageId="12" pageNumber="247">
<emphasis id="B9615A43FF97FF9FFF5E15E1FECDFB13" bold="true" box="[151,309,1186,1211]" pageId="12" pageNumber="247">Type locality.</emphasis>
&quot;India&quot;.
</paragraph>
<paragraph id="8BAA8651FF97FF9FFF0E1587FD5EFA0D" blockId="12.[151,1437,1186,1945]" pageId="12" pageNumber="247">
<emphasis id="B9615A43FF97FF9FFF0E1587FE8DFB75" bold="true" box="[199,373,1220,1245]" pageId="12" pageNumber="247">Type material.</emphasis>
<taxonomicName id="4C15FDD2FF97FF9FFEB21587FDBDFB75" authorityName="Sharp" authorityYear="1882" box="[379,581,1220,1245]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B9615A43FF97FF9FFEB21587FDBDFB75" box="[379,581,1220,1245]" italics="true" pageId="12" pageNumber="247">Copelatus indicus</emphasis>
</taxonomicName>
: Lectotype ♂ (BMNH), designated by
<bibRefCitation id="EF84FBA0FF97FF9FFC351587FB1EFB74" box="[1020,1254,1220,1245]" pageId="12" pageNumber="247" refId="ref16506" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4, 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article">Vazirani (1970: 312)</bibRefCitation>
, labelled: &quot;Type [p, round label with red frame // India / 696 [hw] // Sharp coll. / 1905-313. [p] // Type 696 /
<taxonomicName id="4C15FDD2FF97FF9FFB5115A6FA99FB56" authority="Sheth &amp; Ghate &amp; Hájek, 2018" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[1176,1377,1253,1278]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">Copelatus indicus</taxonomicName>
<taxonomicNameLabel id="A252E738FF97FF9FFAAE15A5FA64FB55" box="[1383,1436,1254,1277]" pageId="12" pageNumber="247" rank="species">n.sp.</taxonomicNameLabel>
/ India [hw]&quot; Paralectotypes: 1♂, &quot;Co- / type [p, round label with yellow frame] // India 696 [hw] // Sharp coll. / 1905-313. [p] //
<taxonomicName id="4C15FDD2FF97FF9FFE87146BFE46FAE9" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[334,446,1320,1345]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/
<taxonomicName id="4C15FDD2FF97FF9FFE1C146BFDDEFAE9" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[469,550,1320,1345]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus">indicus</taxonomicName>
/ ♂ Sharp. / Co-type [hw]&quot;; 1♀, &quot;Co- / type [p, round label with yellow frame] // 696 [hw] // India [p, underlined yellow] // Sharp coll. / 1905-313. [p] //
<taxonomicName id="4C15FDD2FF97FF9FFC1E140AFBBFFACA" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[983,1095,1353,1378]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/
<taxonomicName id="4C15FDD2FF97FF9FFB97140AFB57FACA" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[1118,1199,1353,1378]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus">indicus</taxonomicName>
/ Sharp. / ♀ Co-type [hw]&quot;; 1♀, &quot;Co- / type [p, round label with yellow frame] // India bor. / 696 [hw] // Sharp coll. / 1905-313. [p] //
<taxonomicName id="4C15FDD2FF97FF9FFF5E14CFFEFFFA0D" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[151,263,1420,1445]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/
<taxonomicName id="4C15FDD2FF97FF9FFED414CFFE96FA0D" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[285,366,1420,1445]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="indicus">indicus</taxonomicName>
/ Sharp. / ♀ Co-type [hw]&quot;.
</paragraph>
<paragraph id="8BAA8651FF97FF9FFF5E14EDFDA7FA6E" blockId="12.[151,1437,1186,1945]" box="[151,607,1453,1478]" pageId="12" pageNumber="247">Notes. Median lobe of the lectotype lost.</paragraph>
<paragraph id="8BAA8651FF97FF9FFF0E148DFA7AF9E3" blockId="12.[151,1437,1186,1945]" pageId="12" pageNumber="247">
<materialsCitation id="3B7D8C0CFF97FF9FFF0E148DFDDFF9A0" ID-GBIF-Occurrence="1917355521" collectingDate="2005-09-24" collectingDateMax="2005-09-28" collectingDateMin="2005-09-24" collectionCode="NMPC" collectorName="J. Bezdek" country="India" elevation="500" location="Bhushi" pageId="12" pageNumber="247" specimenCount="2" specimenCount-male="2" stateProvince="Maharashtra">
<emphasis id="B9615A43FF97FF9FFF0E148DFDC2FA4F" bold="true" box="[199,570,1486,1511]" pageId="12" pageNumber="247">Additional material examined.</emphasis>
<collectingCountry id="F302C6C1FF97FF9FFD8D148DFD7AFA4F" box="[580,642,1486,1511]" name="India" pageId="12" pageNumber="247">India</collectingCountry>
:
<collectingRegion id="49D148B3FF97FF9FFD59148DFCD9FA4F" box="[656,801,1486,1511]" country="India" name="Maharashtra" pageId="12" pageNumber="247">Maharashtra</collectingRegion>
:
<specimenCount id="9D134DD8FF97FF9FFCE7148CFCACFA4F" box="[814,852,1487,1511]" pageId="12" pageNumber="247" type="male">2♂</specimenCount>
,
<locationDeviation id="9A189F73FF97FF9FFCAA148CFBD6FA4F" box="[867,1070,1486,1511]" pageId="12" pageNumber="247">
<quantity id="4CED2BB4FF97FF9FFCAA148CFC67FA4F" box="[867,927,1486,1511]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="247" unit="km" value="4.0">4 km</quantity>
S Lonavala
</locationDeviation>
,
<location id="8ECAD08AFF97FF9FFBF4148DFB74FA4F" LSID="urn:lsid:plazi:treatment:03BC3747FF97FF9FFF5E12BEFB88F831:8ECAD08AFF97FF9FFBF4148DFB74FA4F" box="[1085,1164,1486,1511]" country="India" name="Bhushi" pageId="12" pageNumber="247" stateProvince="Maharashtra">Bhushi</location>
dam env.,
<elevation id="00386162FF97FF9FFAD9148DFAA4FA4F" box="[1296,1372,1486,1511]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="247" unit="m" value="500.0">
<quantity id="4CED2BB4FF97FF9FFAD9148DFAA4FA4F" box="[1296,1372,1486,1511]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="247" unit="m" value="500.0">500 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF97FF9FFAA2148CFEE9F9A1" pageId="12" pageNumber="247" value="2005-09-24" valueMax="2005-09-28" valueMin="2005-09-24">
<date id="FFABA091FF97FF9FFAA2148CFEE9F9A1" pageId="12" pageNumber="247" value="2005-09-24" valueMax="2005-09-28" valueMin="2005-09-24">24.– 28.ix.2005</date>
</collectingDate>
,
<collectorName id="26E0E387FF97FF9FFED714B2FE74F9A1" box="[286,396,1520,1545]" pageId="12" pageNumber="247">J. Bezděk</collectorName>
lgt. (
<collectionCode id="ED041E94FF97FF9FFE0014B3FDE7F9A0" box="[457,543,1520,1544]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="12" pageNumber="247">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF97FF9FFDFA14B3FABAF9A0" ID-GBIF-Occurrence="1917355262" box="[563,1346,1520,1545]" collectingDate="2005-10-02" collectingDateMax="2005-10-07" collectingDateMin="2005-10-02" collectionCode="NMPC" collectorName="J. Bezdek" country="India" location="Wai" municipality="Pune" pageId="12" pageNumber="247" specimenCount="1" specimenCount-male="1" stateProvince="Maharashtra">
<specimenCount id="9D134DD8FF97FF9FFDFA14B3FDAFF9A0" box="[563,599,1520,1544]" pageId="12" pageNumber="247" type="male">1♂</specimenCount>
,
<quantity id="4CED2BB4FF97FF9FFDAC14B3FD55F9A1" box="[613,685,1520,1545]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="247" unit="km" value="70.0">70 km</quantity>
S of
<collectingMunicipality id="6BCE1C2BFF97FF9FFD2514B2FCDFF9A0" box="[748,807,1521,1544]" pageId="12" pageNumber="247">Pune</collectingMunicipality>
,
<location id="8ECAD08AFF97FF9FFCFA14B3FCA7F9A1" LSID="urn:lsid:plazi:treatment:03BC3747FF97FF9FFF5E12BEFB88F831:8ECAD08AFF97FF9FFCFA14B3FCA7F9A1" box="[819,863,1520,1545]" country="India" municipality="Pune" name="Wai" pageId="12" pageNumber="247" stateProvince="Maharashtra">Wai</location>
env.,
<collectingDate id="EFEF5979FF97FF9FFC6C14B3FBD4F9A1" box="[933,1068,1520,1545]" pageId="12" pageNumber="247" value="2005-10-02" valueMax="2005-10-07" valueMin="2005-10-02">
<date id="FFABA091FF97FF9FFC6C14B3FBD4F9A1" box="[933,1068,1520,1545]" pageId="12" pageNumber="247" value="2005-10-02" valueMax="2005-10-07" valueMin="2005-10-02">2.–7.x.2005</date>
</collectingDate>
,
<collectorName id="26E0E387FF97FF9FFBF014B2FB5FF9A1" box="[1081,1191,1520,1545]" pageId="12" pageNumber="247">J. Bezděk</collectorName>
lgt. (
<collectionCode id="ED041E94FF97FF9FFB2D14B3FAC2F9A0" box="[1252,1338,1520,1544]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="12" pageNumber="247">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF97FF9FFA8714B3FF05F9E3" ID-GBIF-Occurrence="1917355251" collectingDate="2006-06-03" collectingDateMax="2006-06-06" collectingDateMin="2006-06-03" collectionCode="NMPC" collectorName="V. Ryjacek" country="India" elevation="1280" latitude="17.916666" location="Panchgani" longLatPrecision="1277" longitude="73.816666" municipality="Mahabaleshwar" pageId="12" pageNumber="247" specimenCount="1" specimenCount-female="1" stateProvince="Maharashtra">
<specimenCount id="9D134DD8FF97FF9FFA8714B3FA89F9A0" box="[1358,1393,1520,1544]" pageId="12" pageNumber="247" type="female">1♀</specimenCount>
,
<quantity id="4CED2BB4FF97FF9FFA4914B3FF43F982" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="247" unit="km" value="15.0">15 km</quantity>
E of
<collectingMunicipality id="6BCE1C2BFF97FF9FFECA1752FE4EF982" box="[259,438,1553,1578]" pageId="12" pageNumber="247">Mahabaleshwar</collectingMunicipality>
, E of
<location id="8ECAD08AFF97FF9FFDCA1752FD82F982" LSID="urn:lsid:plazi:treatment:03BC3747FF97FF9FFF5E12BEFB88F831:8ECAD08AFF97FF9FFDCA1752FD82F982" box="[515,634,1553,1578]" country="India" latitude="17.916666" longLatPrecision="1277" longitude="73.816666" municipality="Mahabaleshwar" name="Panchgani" pageId="12" pageNumber="247" stateProvince="Maharashtra">Panchgani</location>
, table land,
<geoCoordinate id="EE21E096FF97FF9FFCC51752FC94F982" box="[780,876,1553,1578]" direction="north" orientation="latitude" pageId="12" pageNumber="247" precision="925" value="17.916666">17°55′N</geoCoordinate>
,
<geoCoordinate id="EE21E096FF97FF9FFCB21752FC21F982" box="[891,985,1553,1578]" direction="east" orientation="longitude" pageId="12" pageNumber="247" precision="925" value="73.816666">73°49′E</geoCoordinate>
,
<elevation id="00386162FF97FF9FFC2E1751FBBAF981" box="[999,1090,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="12" pageNumber="247" unit="m" value="1280.0">
<quantity id="4CED2BB4FF97FF9FFC2E1751FBBAF981" box="[999,1090,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="12" pageNumber="247" unit="m" value="1280.0">1280 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF97FF9FFB981752FB18F982" box="[1105,1248,1553,1578]" pageId="12" pageNumber="247" value="2006-06-03" valueMax="2006-06-06" valueMin="2006-06-03">
<date id="FFABA091FF97FF9FFB981752FB18F982" box="[1105,1248,1553,1578]" pageId="12" pageNumber="247" value="2006-06-03" valueMax="2006-06-06" valueMin="2006-06-03">3.–6.vi.2006</date>
</collectingDate>
,
<collectorName id="26E0E387FF97FF9FFB391751FA95F982" box="[1264,1389,1553,1578]" pageId="12" pageNumber="247">V. Ryjáček</collectorName>
lgt. (
<collectionCode id="ED041E94FF97FF9FFF561770FF0DF9E3" box="[159,245,1587,1611]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="12" pageNumber="247">NMPC</collectionCode>
)
</materialsCitation>
; 1♂, 1♀, Satara distr., Kaas area, 17°43.4′N, 73°49.5′E,
<quantity id="4CED2BB4FF97FF9FFCBC1770FC34F9E2" box="[885,972,1587,1611]" metricMagnitude="3" metricUnit="m" metricValue="1.237" pageId="12" pageNumber="247" unit="m" value="1237.0">1237 m</quantity>
,
<date id="FFABA091FF97FF9FFC111771FBA2F9E3" box="[984,1114,1586,1611]" pageId="12" pageNumber="247" value="2014-07-16">16.vii.2014</date>
, S. D. Sheth lgt. (HVGC).
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF97FF9FFF0E1717FB6EF8FE" pageId="12" pageNumber="247" type="diagnosis">
<paragraph id="8BAA8651FF97FF9FFF0E1717FF3BF89D" blockId="12.[151,1437,1186,1945]" pageId="12" pageNumber="247">
<emphasis id="B9615A43FF97FF9FFF0E1717FEB9F9C5" bold="true" box="[199,321,1620,1645]" pageId="12" pageNumber="247">Diagnosis.</emphasis>
Medium sized (TL:
<quantity id="4CED2BB4FF97FF9FFDE51717FD40F9C4" box="[556,696,1620,1645]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="12" pageNumber="247" unit="mm" value="5.1" valueMax="5.4" valueMin="4.8">4.8–5.4 mm</quantity>
), oblong oval species. Dorsal surface black, pronotal sides, elytra laterally from dorsal stria 4, and apex of elytra yellow (
<figureCitation id="132E9AD4FF97FF9FFCE51736FC8DF926" box="[812,885,1653,1678]" captionStart="FIGURES 5–8" captionStartId="4.[151,263,1970,1993]" captionTargetBox="[174,1372,207,1947]" captionTargetId="figure@4.[151,1435,193,1949]" captionTargetPageId="4" captionText="FIGURES 5–8. Habitus of Copelatus. 5–6. C. bezdeki sp. nov.: 5. male holotype; 6. female paratype (Lonavala); 7–8. C. indicus: 7. male (Lonavala); 8. female (Mahabaleshwar)." httpUri="https://zenodo.org/record/1458552/files/figure.png" pageId="12" pageNumber="247">Fig. 7</figureCitation>
). All six dorsal striae on elytra complete, well impressed; submarginal stria rather long, starting at about elytral midlength and terminates in approximately apical fifth of elytral length. Females with numerous short strioles covering basal two thirds of elytral length, except for base and lateral margin (
<figureCitation id="132E9AD4FF97FF9FFE65179AFE17F95A" box="[428,495,1753,1778]" captionStart="FIGURES 5–8" captionStartId="4.[151,263,1970,1993]" captionTargetBox="[174,1372,207,1947]" captionTargetId="figure@4.[151,1435,193,1949]" captionTargetPageId="4" captionText="FIGURES 5–8. Habitus of Copelatus. 5–6. C. bezdeki sp. nov.: 5. male holotype; 6. female paratype (Lonavala); 7–8. C. indicus: 7. male (Lonavala); 8. female (Mahabaleshwar)." httpUri="https://zenodo.org/record/1458552/files/figure.png" pageId="12" pageNumber="247">Fig. 8</figureCitation>
). Median lobe sickle-shaped in lateral view, in apical fourth gradually narrowing to obtusely pointed apex (
<figureCitation id="132E9AD4FF97FF9FFE5617B9FE0DF8BB" box="[415,501,1786,1811]" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="12" pageNumber="247">Fig. 25</figureCitation>
). Parameres 'D'-shaped; apex short and broad; apical lobes long, club-shaped (
<figureCitation id="132E9AD4FF97FF9FFAA617B9FF4FF89D" captionStart="FIGURES 23–30" captionStartId="14.[151,263,1513,1536]" captionTargetBox="[159,1413,213,1460]" captionTargetId="figure@14.[151,1436,193,1492]" captionTargetPageId="14" captionText="FIGURES 23–30. Male genitalia of Copelatus. 23–24. C. bezdeki sp. nov.; 25–26. C. indicus; 27–28. C. neelumae; 29–30. C. schereri. 23, 25, 27, 29. median lobe in lateral view (27a. redrawn from original description of C. neelumae); 24, 26, 28, 30. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458560/files/figure.png" pageId="12" pageNumber="247">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="8BAA8651FF97FF9FFF0E167EFB6EF8FE" blockId="12.[151,1437,1186,1945]" box="[199,1174,1853,1878]" pageId="12" pageNumber="247">
<emphasis id="B9615A43FF97FF9FFF0E167EFE0FF8FE" bold="true" box="[199,503,1853,1878]" pageId="12" pageNumber="247">Collecting circumstances.</emphasis>
The species was found in a shallow, ephemeral water body.
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF97FF9FFF0E161DFB88F831" pageId="12" pageNumber="247" type="distribution">
<paragraph id="8BAA8651FF97FF9FFF0E161DFB88F831" blockId="12.[151,1437,1186,1945]" pageId="12" pageNumber="247">
<emphasis id="B9615A43FF97FF9FFF0E161DFE98F8DF" bold="true" box="[199,352,1886,1911]" pageId="12" pageNumber="247">Distribution.</emphasis>
<collectingCountry id="F302C6C1FF97FF9FFEAF161DFE5BF8DF" box="[358,419,1886,1911]" name="India" pageId="12" pageNumber="247">India</collectingCountry>
;
<bibRefCitation id="EF84FBA0FF97FF9FFE67161DFD38F8DF" box="[430,704,1886,1911]" pageId="12" pageNumber="247" refId="ref15475" refString="Ghosh, S. K. &amp; Nilsson, A. N. (2012) Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae). Skorvnopparn Supplement, 3, 1 - 77." type="journal article">Ghosh &amp; Nilsson (2012)</bibRefCitation>
listed the species from
<collectingRegion id="49D148B3FF97FF9FFC0F161CFBEDF8DF" box="[966,1045,1887,1911]" country="India" name="Assam" pageId="12" pageNumber="247">Assam</collectingRegion>
,
<collectingRegion id="49D148B3FF97FF9FFBD6161DFBA7F8DF" box="[1055,1119,1886,1911]" country="India" name="Bihar" pageId="12" pageNumber="247">Bihar</collectingRegion>
,
<collectingRegion id="49D148B3FF97FF9FFBA3161DFB53F8DF" box="[1130,1195,1886,1911]" country="India" name="Delhi" pageId="12" pageNumber="247">Delhi</collectingRegion>
,
<collectingRegion id="49D148B3FF97FF9FFB7C161DFAD3F8DF" box="[1205,1323,1886,1911]" country="India" name="Jharkhand" pageId="12" pageNumber="247">Jharkhand</collectingRegion>
,
<collectingRegion id="49D148B3FF97FF9FFAFF161DFA60F8DF" box="[1334,1432,1886,1911]" country="India" name="Manipur" pageId="12" pageNumber="247">Manipur</collectingRegion>
,
<collectingRegion id="49D148B3FF97FF9FFF5E16C3FF12F831" box="[151,234,1920,1945]" country="India" name="Tripura" pageId="12" pageNumber="247">Tripura</collectingRegion>
and
<collectingRegion id="49D148B3FF97FF9FFEE916C2FE45F831" box="[288,445,1920,1945]" country="India" name="Uttar Pradesh" pageId="12" pageNumber="247">Uttar Pradesh</collectingRegion>
. We add records from
<collectingRegion id="49D148B3FF97FF9FFD7016C3FCBDF831" box="[697,837,1920,1945]" country="India" name="Maharashtra" pageId="12" pageNumber="247">Maharashtra</collectingRegion>
(Satara District) (
<figureCitation id="132E9AD4FF97FF9FFBC616C3FB9CF831" box="[1039,1124,1920,1945]" captionStart="FIGURE 45" captionStartId="21.[151,249,1915,1937]" captionTargetBox="[190,1401,196,1869]" captionTargetId="figure@21.[153,1434,158,1905]" captionTargetPageId="21" captionText="FIGURE 45. Distributional map of Copelatus species in Maharashtra." httpUri="https://zenodo.org/record/1458568/files/figure.png" pageId="12" pageNumber="247">Fig. 45</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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