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<mods:title id="C0CFC0F7A1293C69DDE468A956E2EF9F">Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="52C3BD2EEC2882B45F3F08C324987C05">Ghate, Hemant V.</mods:namePart>
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<taxonomicName id="4C15FDD2FF8FFF87FF5E11D4FDFEFF1A" ID-CoL="Y3DC" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[151,518,151,178]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B9615A43FF8FFF87FF5E11D4FDFEFF1A" bold="true" box="[151,518,151,178]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF5E11D4FE90FF19" bold="true" box="[151,360,151,177]" italics="true" pageId="20" pageNumber="255">Copelatus oblitus</emphasis>
<bibRefCitation id="EF84FBA0FF8FFF87FEA611DBFDFEFF1A" box="[367,518,152,178]" pageId="20" pageNumber="255" refId="ref16437" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article">Sharp, 1882</bibRefCitation>
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<taxonomicName id="4C15FDD2FF8FFF87FF5E119FFDF8FF5A" ID-CoL="Y3DC" authority="Sharp, 1882: 582" authorityName="Sharp" authorityPageNumber="582" authorityYear="1882" box="[151,512,220,242]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B9615A43FF8FFF87FF5E119FFEB1FF5A" box="[151,329,220,242]" italics="true" pageId="20" pageNumber="255">Copelatus oblitus</emphasis>
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<bibRefCitation id="EF84FBA0FF8FFF87FE86119FFE35FF5A" box="[335,461,220,242]" pageId="20" pageNumber="255" refId="ref16437" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article">Sharp, 1882</bibRefCitation>
: 582
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.
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<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E11B9FBDEFEB8" box="[151,1062,250,272]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E11B9FD8AFEB8" ID-CoL="Y336" authority="Regimbart, 1899: 302" authorityName="Regimbart" authorityPageNumber="302" authorityYear="1899" box="[151,626,250,272]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="B9615A43FF8FFF87FF5E11B9FE73FEB8" box="[151,395,250,272]" italics="true" pageId="20" pageNumber="255">Copelatus andamanicus</emphasis>
<treatmentCitation id="0AB4A040FF8FFF87FE5811B9FD8AFEB8" author="" box="[401,626,250,272]" page="302" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FE5811B9FDC7FEB8" box="[401,575,250,272]" pageId="20" pageNumber="255" refId="ref16182" refString="Regimbart, M. (1899) Revision des Dytiscidae de la region Indo- Sino-Malaise. Annales de la Societe Entomologique de France, 68, 186 - 367." type="journal article">Régimbart, 1899</bibRefCitation>
: 302
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; synonymy by
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<bibRefCitation id="EF84FBA0FF8FFF87FCC211B9FC10FEB8" box="[779,1000,250,272]" pageId="20" pageNumber="255" refId="ref15752" refString="Hendrich, L., Balke, M. &amp; Yang, C. M. (2004) Aquatic Coleoptera of Singapore: Species richness, ecology and conservation. The Raffles Bulletin of Zoology, 52 (1), 97 - 145." type="journal article">
Hendrich
<emphasis id="B9615A43FF8FFF87FCB911B8FC64FEB8" box="[880,924,250,272]" italics="true" pageId="20" pageNumber="255">et al</emphasis>
. (2004
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: 118
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).
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<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E105AFBD1FE87" box="[151,1065,281,303]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E105AFD8BFE87" authority="Sato, 1961: 8" authorityName="Sato" authorityPageNumber="8" authorityYear="1961" box="[151,627,281,303]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="subSpecies" species="sociennus" subSpecies="ryukyuensis">
<emphasis id="B9615A43FF8FFF87FF5E105AFE1DFE87" box="[151,485,281,303]" italics="true" pageId="20" pageNumber="255">Copelatus sociennus ryukyuensis</emphasis>
<treatmentCitation id="0AB4A040FF8FFF87FE25105AFD8BFE87" author="" box="[492,627,281,303]" page="8" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FE25105AFDA3FE87" box="[492,603,281,303]" pageId="20" pageNumber="255" refId="ref16345" refString="Sato, M. (1961) Aquatic Coleoptera from Amami-Oshima of the Ryukyu Islands (II). Akitu, 10, 7 - 10." type="journal article">Satô, 1961</bibRefCitation>
: 8
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</taxonomicName>
; synonymy by
<treatmentCitation id="0AB4A040FF8FFF87FCC6105AFBE6FE87" author="" box="[783,1054,281,303]" page="118" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FCC6105AFC13FE87" box="[783,1003,281,303]" pageId="20" pageNumber="255" refId="ref15752" refString="Hendrich, L., Balke, M. &amp; Yang, C. M. (2004) Aquatic Coleoptera of Singapore: Species richness, ecology and conservation. The Raffles Bulletin of Zoology, 52 (1), 97 - 145." type="journal article">
Hendrich
<emphasis id="B9615A43FF8FFF87FCBA1059FC58FE87" box="[883,928,281,303]" italics="true" pageId="20" pageNumber="255">et al</emphasis>
. (2004
</bibRefCitation>
: 118
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).
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</paragraph>
<paragraph id="8BAA8651FF8FFF87FF5E107BFBD6FEE6" blockId="20.[151,1070,220,395]" box="[151,1070,312,334]" pageId="20" pageNumber="255">
<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E107BFBD6FEE6" box="[151,1070,312,334]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E107BFD82FEE6" authority="Zimmermann, 1919: 76" authorityName="Zimmermann" authorityPageNumber="76" authorityYear="1919" box="[151,634,312,334]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="subfasciatus">
<emphasis id="B9615A43FF8FFF87FF5E107BFE79FEE6" box="[151,385,312,334]" italics="true" pageId="20" pageNumber="255">Copelatus subfasciatus</emphasis>
<treatmentCitation id="0AB4A040FF8FFF87FE4E107BFD82FEE6" author="" box="[391,634,312,334]" page="76" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FE4E107BFDACFEE6" box="[391,596,312,334]" pageId="20" pageNumber="255" refId="ref16820" refString="Zimmermann, A. (1919) H. Sauter's Formosa-Ausbeute: Haliplidae et Dytiscidae (Col.). Entomologische Mitteilungen, 8, 75 - 77. https: // doi. org / 10.5962 / bhl. part. 27264" type="journal article">Zimmermann, 1919</bibRefCitation>
: 76
</treatmentCitation>
</taxonomicName>
; synonymy by
<treatmentCitation id="0AB4A040FF8FFF87FCDD107BFBDAFEE6" author="" box="[788,1058,312,334]" page="118" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FCDD107BFC08FEE6" box="[788,1008,312,334]" pageId="20" pageNumber="255" refId="ref15752" refString="Hendrich, L., Balke, M. &amp; Yang, C. M. (2004) Aquatic Coleoptera of Singapore: Species richness, ecology and conservation. The Raffles Bulletin of Zoology, 52 (1), 97 - 145." type="journal article">
Hendrich
<emphasis id="B9615A43FF8FFF87FCB1107AFC5DFEE6" box="[888,933,312,334]" italics="true" pageId="20" pageNumber="255">et al</emphasis>
. (2004
</bibRefCitation>
: 118
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E1015FC11FEC4" box="[151,1001,342,365]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E1015FDCDFEC4" authority="Nakane, 1963: 25" authorityName="Nakane" authorityPageNumber="25" authorityYear="1963" box="[151,565,342,364]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="tokaraensis">
<emphasis id="B9615A43FF8FFF87FF5E1015FE81FEC4" box="[151,377,342,364]" italics="true" pageId="20" pageNumber="255">Copelatus tokaraensis</emphasis>
<treatmentCitation id="0AB4A040FF8FFF87FEB61015FDCDFEC4" author="" box="[383,565,342,364]" page="25" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FEB61015FDE8FEC4" box="[383,528,342,364]" pageId="20" pageNumber="255" refId="ref16058" refString="Nakane, T. (1963) New or little-known Coleoptera from Japan and its adjacent regions. XVIII. Fragmenta Coleopterologica, 6, 23 - 26." type="journal article">Nakane, 1963</bibRefCitation>
: 25
</treatmentCitation>
</taxonomicName>
; synonymy by
<treatmentCitation id="0AB4A040FF8FFF87FD061015FC26FEC4" author="" box="[719,990,342,365]" page="118" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FD061015FC53FEC4" box="[719,939,342,365]" pageId="20" pageNumber="255" refId="ref15752" refString="Hendrich, L., Balke, M. &amp; Yang, C. M. (2004) Aquatic Coleoptera of Singapore: Species richness, ecology and conservation. The Raffles Bulletin of Zoology, 52 (1), 97 - 145." type="journal article">
Hendrich
<emphasis id="B9615A43FF8FFF87FCFA101BFC98FEC4" box="[819,864,342,365]" italics="true" pageId="20" pageNumber="255">et al</emphasis>
. (2004
</bibRefCitation>
: 118
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF5E1036FCA0FE23" blockId="20.[151,1070,220,395]" box="[151,856,373,395]" pageId="20" pageNumber="255">
<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E1036FCA0FE23" box="[151,856,373,395]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E1036FD51FE23" ID-CoL="8L47D" authority="Holmen &amp; Vazirani, 1990: 27" authorityName="Holmen &amp; Vazirani" authorityPageNumber="27" authorityYear="1990" box="[151,681,373,395]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="karnatakus">
<emphasis id="B9615A43FF8FFF87FF5E1036FE8CFE23" box="[151,372,373,395]" italics="true" pageId="20" pageNumber="255">Copelatus karnatakus</emphasis>
<treatmentCitation id="0AB4A040FF8FFF87FEB21036FD51FE23" author="" box="[379,681,373,395]" page="27" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FEB21036FD7BFE23" box="[379,643,373,395]" pageId="20" pageNumber="255" refId="ref15798" refString="Holmen, M. &amp; Vazirani, T. G. (1990) Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 60, 19 - 32." type="journal article">Holmen &amp; Vazirani, 1990</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
;
<emphasis id="B9615A43FF8FFF87FD7D1036FCAAFE22" bold="true" box="[692,850,373,394]" pageId="20" pageNumber="255">new synonymy</emphasis>
.
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</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8FFF87FF5E10F6FA62FD3E" pageId="20" pageNumber="255" type="materials_examined">
<paragraph id="8BAA8651FF8FFF87FF5E10F6FB0EFE66" blockId="20.[151,1436,437,895]" box="[151,1270,437,462]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF5E10F6FEB0FE66" bold="true" box="[151,328,437,462]" pageId="20" pageNumber="255">
<typeStatus id="54AE38F3FF8FFF87FF5E10F6FF2AFE66" box="[151,210,437,462]" pageId="20" pageNumber="255">Type</typeStatus>
localities.
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<taxonomicName id="4C15FDD2FF8FFF87FE8710F6FDEBFE66" authorityName="Sharp" authorityYear="1882" box="[334,531,437,462]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B9615A43FF8FFF87FE8710F6FDEBFE66" box="[334,531,437,462]" italics="true" pageId="20" pageNumber="255">Copelatus oblitus</emphasis>
</taxonomicName>
: &quot;
<collectingCountry id="F302C6C1FF8FFF87FDE210F6FD5DFE66" box="[555,677,437,462]" name="Singapore" pageId="20" pageNumber="255">Singapore</collectingCountry>
&quot;.
<taxonomicName id="4C15FDD2FF8FFF87FD7110F5FCADFE66" authorityName="Holmen &amp; Vazirani" authorityYear="1990" box="[696,853,437,462]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="karnatakus">
<emphasis id="B9615A43FF8FFF87FD7110F5FCADFE66" box="[696,853,437,462]" italics="true" pageId="20" pageNumber="255">C. karnatakus</emphasis>
</taxonomicName>
: &quot;
<collectingCountry id="F302C6C1FF8FFF87FCA410F6FC54FE66" box="[877,940,437,462]" name="India" pageId="20" pageNumber="255">India</collectingCountry>
,
<collectingRegion id="49D148B3FF8FFF87FC7E10F6FBD5FE66" box="[951,1069,437,462]" country="India" name="Karnataka" pageId="20" pageNumber="255">Karnataka</collectingRegion>
, Mudigere Area&quot;.
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF0E1095FA7AFDB8" blockId="20.[151,1436,437,895]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF0E1095FE80FE47" bold="true" box="[199,376,470,495]" pageId="20" pageNumber="255">
<typeStatus id="54AE38F3FF8FFF87FF0E1095FEFAFE47" box="[199,258,470,495]" pageId="20" pageNumber="255">Type</typeStatus>
material.
</emphasis>
<taxonomicName id="4C15FDD2FF8FFF87FE481095FDB1FE47" authorityName="Sharp" authorityYear="1882" box="[385,585,470,495]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B9615A43FF8FFF87FE481095FDB1FE47" box="[385,585,470,495]" italics="true" pageId="20" pageNumber="255">Copelatus oblitus</emphasis>
</taxonomicName>
:
<typeStatus id="54AE38F3FF8FFF87FD921095FD3BFE47" box="[603,707,470,495]" pageId="20" pageNumber="255" type="holotype">Holotype</typeStatus>
<specimenCount id="9D134DD8FF8FFF87FD041094FD1AFE46" box="[717,738,471,494]" pageId="20" pageNumber="255" type="male">♂</specimenCount>
(
<collectionCode id="ED041E94FF8FFF87FD3D1094FCB5FE47" box="[756,845,471,495]" pageId="20" pageNumber="255">BMNH</collectionCode>
), labelled: &quot;
<typeStatus id="54AE38F3FF8FFF87FC1F1094FBE8FE47" box="[982,1040,471,495]" pageId="20" pageNumber="255">Type</typeStatus>
[p, round label with red frame] //
<collectingCountry id="F302C6C1FF8FFF87FF5E10B4FEF1FDB8" box="[151,265,503,528]" name="Singapore" pageId="20" pageNumber="255">Singapore</collectingCountry>
/ 695 [hw] // Sharp Coll. / 1905-313. [p] //
<typeStatus id="54AE38F3FF8FFF87FD2710BBFCDDFDB8" box="[750,805,504,528]" pageId="20" pageNumber="255">Type</typeStatus>
695 DS. /
<taxonomicName id="4C15FDD2FF8FFF87FC5710B4FBF6FDB8" authority="Sheth &amp; Ghate &amp; Hájek, 2018" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[926,1038,503,528]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus" status="sp. nov.">Copelatus</taxonomicName>
/
<taxonomicName id="4C15FDD2FF8FFF87FBED10B4FB88FDB8" authority="Sheth &amp; Ghate &amp; Hájek, 2018" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[1060,1136,503,528]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus" status="sp. nov.">oblitus</taxonomicName>
<taxonomicNameLabel id="A252E738FF8FFF87FBB110BBFB55FDA7" box="[1144,1197,504,527]" pageId="20" pageNumber="255" rank="species">n.sp.</taxonomicNameLabel>
/
<collectingCountry id="F302C6C1FF8FFF87FB0B10B4FACBFDB8" box="[1218,1331,503,528]" name="Singapore" pageId="20" pageNumber="255">Singapore</collectingCountry>
[hw]&quot;.
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF5E135AFB05FDDC" blockId="20.[151,1436,437,895]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E135AFE6BFD9A" authorityName="Holmen &amp; Vazirani" authorityYear="1990" box="[151,403,537,562]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="karnatakus">
<emphasis id="B9615A43FF8FFF87FF5E135AFE6BFD9A" box="[151,403,537,562]" italics="true" pageId="20" pageNumber="255">Copelatus karnatakus</emphasis>
</taxonomicName>
: Holotype ♂ (ZMUC), labelled: &quot;S.India: Karnataka / Mudigere area, c.
<quantity id="4CED2BB4FF8FFF87FAC1135AFAA3FD99" box="[1288,1371,537,561]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="255" unit="m" value="900.0">900 m</quantity>
/ 2– 1o.
<date id="FFABA091FF8FFF87FF7A1379FEF7FDFB" box="[179,271,570,595]" pageId="20" pageNumber="255" value="1977-11">xi.1977</date>
/ Zool.Mus.Copenhagen Exp. [p] // Holo- / type [p, round label with red margin] //
<taxonomicName id="4C15FDD2FF8FFF87FADE1379FA7FFDFB" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[1303,1415,570,595]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/
<taxonomicName id="4C15FDD2FF8FFF87FF5E131FFEE8FDDD" authorityName="Holmen &amp; Vazirani" authorityYear="1990" box="[151,272,604,629]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="karnatakus">karnatakus</taxonomicName>
/
<emphasis id="B9615A43FF8FFF87FEEF131FFE9DFDDD" bold="true" box="[294,357,604,629]" pageId="20" pageNumber="255">sp. n.</emphasis>
[hw] / det. T.G. Vazirani, 198 [p] 2 [hw] / m. Holmen [hw] // zmuc00020126 [p]&quot;.
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF0E133EFA62FD3E" blockId="20.[151,1436,437,895]" box="[199,1434,637,662]" pageId="20" pageNumber="255">
<materialsCitation id="3B7D8C0CFF8FFF87FF0E133EFA62FD3E" ID-GBIF-Occurrence="1917355419" box="[199,1434,637,662]" collectionCode="BMNH" collectorName="Haldwani, H. G" country="India" location="Kumaon district" pageId="20" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Uttarakhand">
<emphasis id="B9615A43FF8FFF87FF0E133EFD90FD3E" bold="true" box="[199,616,637,662]" pageId="20" pageNumber="255">
Additional material studied.
<collectingCountry id="F302C6C1FF8FFF87FDD7133EFD9AFD3E" box="[542,610,637,662]" name="India" pageId="20" pageNumber="255">India</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8FFF87FDA6133DFD6BFD3E" box="[623,659,638,662]" pageId="20" pageNumber="255" type="male">1♂</specimenCount>
,
<collectingRegion id="49D148B3FF8FFF87FD69133EFCD5FD3E" box="[672,813,637,662]" country="India" name="Uttarakhand" pageId="20" pageNumber="255">Uttarakhand</collectingRegion>
,
<location id="8ECAD08AFF8FFF87FCF1133DFC08FD3E" LSID="urn:lsid:plazi:treatment:03BC3747FF8FFF87FF5E11D4FABBFCD7:8ECAD08AFF8FFF87FCF1133DFC08FD3E" box="[824,1008,637,662]" country="India" name="Kumaon district" pageId="20" pageNumber="255" stateProvince="Uttarakhand">Kumaon district</location>
,
<collectorName id="26E0E387FF8FFF87FC32133EFB5EFD3E" box="[1019,1190,637,662]" pageId="20" pageNumber="255">Haldwani, H.G</collectorName>
. Champion (
<collectionCode id="ED041E94FF8FFF87FAFD133DFA76FD3E" box="[1332,1422,638,662]" pageId="20" pageNumber="255">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8FFF87FF0E13DDFC3BFC95" pageId="20" pageNumber="255" type="discussion">
<paragraph id="8BAA8651FF8FFF87FF0E13DDFC3BFC95" blockId="20.[151,1436,437,895]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF0E13DDFDE4FD1F" bold="true" box="[199,540,670,695]" pageId="20" pageNumber="255">Comments on classification.</emphasis>
<bibRefCitation id="EF84FBA0FF8FFF87FDEE13DDFC98FD1F" box="[551,864,670,695]" pageId="20" pageNumber="255" refId="ref15798" refString="Holmen, M. &amp; Vazirani, T. G. (1990) Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 60, 19 - 32." type="journal article">Holmen &amp; Vazirani (1990)</bibRefCitation>
diagnosed
<taxonomicName id="4C15FDD2FF8FFF87FC2E13DCFB70FD1F" authorityName="Holmen &amp; Vazirani" authorityYear="1990" box="[999,1160,670,695]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="karnatakus">
<emphasis id="B9615A43FF8FFF87FC2E13DCFB70FD1F" box="[999,1160,670,695]" italics="true" pageId="20" pageNumber="255">C. karnatakus</emphasis>
</taxonomicName>
as a member of the
<taxonomicName id="4C15FDD2FF8FFF87FA4B13DCFF20FD70" authorityName="Régimbart" authorityYear="1899" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="irinus">
<emphasis id="B9615A43FF8FFF87FA4B13DCFF20FD70" italics="true" pageId="20" pageNumber="255">C. irinus</emphasis>
</taxonomicName>
species group with characteristic shape of median lobe with characteristic dorsal (ventral in current concept) projection. Apparently, they were not aware of the widespread
<emphasis id="B9615A43FF8FFF87FC4E13A1FBF9FD52" box="[903,1025,737,762]" italics="true" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FC4E13A1FC05FD52" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[903,1021,737,762]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus">C. oblitus</taxonomicName>
.
</emphasis>
The direct comparison of the two
<typeStatus id="54AE38F3FF8FFF87FF5E1241FEFCFCB3" box="[151,260,770,795]" pageId="20" pageNumber="255" type="holotype">holotypes</typeStatus>
undoubtedly proved that they are identical. Therefore, we establish the following synonymy:
<taxonomicName id="4C15FDD2FF8FFF87FAE31241FE89FC94" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B9615A43FF8FFF87FAE31241FF1BFC95" italics="true" pageId="20" pageNumber="255">Copelatus oblitus</emphasis>
<bibRefCitation id="EF84FBA0FF8FFF87FF231267FE89FC94" box="[234,369,804,829]" pageId="20" pageNumber="255" refId="ref16437" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article">Sharp, 1882</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C15FDD2FF8FFF87FE461266FCADFC94" authority="Holmen &amp; Vazirani, 1990" authorityName="Holmen &amp; Vazirani" authorityYear="1990" box="[399,853,804,829]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="karnatakus">
<emphasis id="B9615A43FF8FFF87FE461266FDD5FC95" box="[399,557,804,829]" italics="true" pageId="20" pageNumber="255">C. karnatakus</emphasis>
<bibRefCitation id="EF84FBA0FF8FFF87FDFA1267FCADFC94" box="[563,853,804,829]" pageId="20" pageNumber="255" refId="ref15798" refString="Holmen, M. &amp; Vazirani, T. G. (1990) Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 60, 19 - 32." type="journal article">Holmen &amp; Vazirani, 1990</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A252E738FF8FFF87FC951267FC3BFC95" box="[860,963,804,829]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FC951267FC3BFC95" bold="true" box="[860,963,804,829]" pageId="20" pageNumber="255">syn. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8FFF87FF0E1206FABBFCD7" pageId="20" pageNumber="255" type="distribution">
<paragraph id="8BAA8651FF8FFF87FF0E1206FABBFCD7" blockId="20.[151,1436,437,895]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF0E1206FE98FCF6" bold="true" box="[199,352,837,862]" pageId="20" pageNumber="255">Distribution.</emphasis>
Widely distributed species ranging in the
<collectingRegion id="49D148B3FF8FFF87FC941206FC41FCF6" box="[861,953,837,862]" country="Democratic Republic of the Congo" name="Orientale" pageId="20" pageNumber="255">Oriental</collectingRegion>
region from
<collectingCountry id="F302C6C1FF8FFF87FB921206FB6CFCF6" box="[1115,1172,837,862]" name="India" pageId="20" pageNumber="255">India</collectingCountry>
to Sulawesi, and also reaching southern
<collectingCountry id="F302C6C1FF8FFF87FEAC1225FE50FCD7" box="[357,424,870,895]" name="China" pageId="20" pageNumber="255">China</collectingCountry>
and
<collectingCountry id="F302C6C1FF8FFF87FE161224FDE7FCD6" box="[479,543,871,894]" name="Japan" pageId="20" pageNumber="255">Japan</collectingCountry>
in the Palaearctic region. First record from Indian state of
<collectingRegion id="49D148B3FF8FFF87FB791225FAC6FCD7" box="[1200,1342,870,895]" country="India" name="Uttarakhand" pageId="20" pageNumber="255">Uttarakhand</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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