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<document id="352545E6A637D75E0F5E64BA564D660C" ID-CLB-Dataset="29024" ID-DOI="10.11646/zootaxa.4459.2.2" ID-GBIF-Dataset="634e23da-84c4-46a4-8bb8-94fcf1637160" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458548" ID-ZooBank="73510C3B-6482-4760-8173-B6E701839940" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288809625" checkinUser="plazi" docAuthor="Sheth, Sayali D., Ghate, Hemant V. &amp; Hájek, Jiří" docDate="2018" docId="03BC3747FF8DFF8BFF5E1487FD41FE78" docLanguage="en" docName="zootaxa.4459.2.2.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus tenebrosus Regimbart 1880" docType="treatment" docVersion="7" lastPageNumber="259" masterDocId="FF854F3FFF9BFF93FFC91143FFF8FFA8" masterDocTitle="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="260" masterPageNumber="235" pageNumber="257" updateTime="1698578320975" updateUser="ExternalLinkService">
<mods:mods id="47BC652658C8A59A15DDAEBA4C522370" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4DC0537FF25031A6D99BEA828ED2E61E">
<mods:title id="12870217FDC9E0DF0CC28B2F243BB4B8">Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="B0849B6984278DAE75B1575CBE2285E5" type="personal">
<mods:role id="EA04DCFB6110475C5E501C48D07667E4">
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<mods:namePart id="FEF3815F0592813383ECC38D4CF2D7A1">Sheth, Sayali D.</mods:namePart>
</mods:name>
<mods:name id="BD1124BCBAD1EB8F39A9536F169D3C46" type="personal">
<mods:role id="DFE2D65243FB8B0C8FB274DB38CC415C">
<mods:roleTerm id="FB52106311821C42977B8E0649AAF95E">Author</mods:roleTerm>
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<mods:namePart id="46EBC100FD9BB4C7189B0803F2B4555D">Ghate, Hemant V.</mods:namePart>
</mods:name>
<mods:name id="4966D85B7AD38C06E086F5EDB7448995" type="personal">
<mods:role id="0CE5F1BB33F2EB2FAF4F7716CF1D7E21">
<mods:roleTerm id="BE81556FB8EAAE68E4CF56C67258E724">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="87033C64D1C815B370AB7A46854AF7FD">Hájek, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="7E09F04375EA40428F0A44380C41C551">text</mods:typeOfResource>
<mods:relatedItem id="D3EF1D3632E1911E8B02327E30285202" type="host">
<mods:titleInfo id="B23A6CB403D3685D170EA235E676F35D">
<mods:title id="DD534AFA373D1059CB0AD38A80003123">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D5D88B1606697A91FE85FD5A7B808628">
<mods:date id="12D97798EEF36C2F857F7EF7F9FB6EA8">2018</mods:date>
<mods:detail id="F1BDC060B43A0668CCC4047B1C4B01A4" type="pubDate">
<mods:number id="23A0A0152B9A2B147282C085627DEAD6">2018-08-15</mods:number>
</mods:detail>
<mods:detail id="B37186379BB51A4EBEE1B449DEC9FB07" type="volume">
<mods:number id="3B218CEA844BC9F04F9C5D49AF4EACE9">4459</mods:number>
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<mods:detail id="81A103D8194EE437E0F8062ABAE6ECBD" type="issue">
<mods:number id="868963CC99340EED5D8766711EA0294E">2</mods:number>
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<mods:start id="078F6BCBA3BEEF510E9CC0A44FDEC71A">235</mods:start>
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<mods:classification id="F9AF64E0A04ED14C7AFE91107B58FDCF">journal article</mods:classification>
<mods:identifier id="FCBA78996982977D4A0DA7841B76E338" type="CLB-Dataset">29024</mods:identifier>
<mods:identifier id="4EE613D69E8F32DC6F77411B25C2A45D" type="DOI">10.11646/zootaxa.4459.2.2</mods:identifier>
<mods:identifier id="F60CCC8E9CED5C92ADA827BDD128F8A8" type="GBIF-Dataset">634e23da-84c4-46a4-8bb8-94fcf1637160</mods:identifier>
<mods:identifier id="08DDE39C0558FEA72C0609ACEC804A71" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CB23ADAD8071CE77EE0117A45C9AFD44" type="Zenodo-Dep">1458548</mods:identifier>
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</mods:mods>
<treatment id="03BC3747FF8DFF8BFF5E1487FD41FE78" ID-DOI="http://doi.org/10.5281/zenodo.5966553" ID-GBIF-Taxon="148692683" ID-Zenodo-Dep="5966553" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78" httpUri="http://treatment.plazi.org/id/03BC3747FF8DFF8BFF5E1487FD41FE78" lastPageId="24" lastPageNumber="259" pageId="22" pageNumber="257">
<subSubSection id="C30FD5DAFF8DFF85FF5E1487FD88FA76" box="[151,624,1476,1502]" pageId="22" pageNumber="257" type="nomenclature">
<paragraph id="8BAA8651FF8DFF85FF5E1487FD88FA76" blockId="22.[151,624,1476,1502]" box="[151,624,1476,1502]" pageId="22" pageNumber="257">
<heading id="D0E2313DFF8DFF85FF5E1487FD88FA76" bold="true" box="[151,624,1476,1502]" fontSize="11" level="1" pageId="22" pageNumber="257" reason="1">
<taxonomicName id="4C15FDD2FF8DFF85FF5E1487FD88FA76" ID-CoL="Y3GY" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[151,624,1476,1502]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF8DFF85FF5E1487FD88FA76" bold="true" box="[151,624,1476,1502]" pageId="22" pageNumber="257">
<emphasis id="B9615A43FF8DFF85FF5E1487FE61FA76" bold="true" box="[151,409,1476,1502]" italics="true" pageId="22" pageNumber="257">Copelatus tenebrosus</emphasis>
<bibRefCitation id="EF84FBA0FF8DFF85FE691487FD88FA76" box="[416,624,1476,1502]" pageId="22" pageNumber="257" refId="ref16151" refString="Regimbart, M. (1880) The new Dytiscidae and Gyrinidae collected during the recent scientific Sumatra-expedition. Notes from the Leyden Museum, 2, 209 - 216." type="journal article">Régimbart, 1880</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8DFF85FF5E1748FD0CF933" pageId="22" pageNumber="257" type="reference_group">
<paragraph id="8BAA8651FF8DFF85FF5E1748FDA3F989" blockId="22.[151,914,1547,1691]" box="[151,603,1547,1569]" pageId="22" pageNumber="257">
<treatmentCitationGroup id="AB05A17FFF8DFF85FF5E1748FDA3F989" box="[151,603,1547,1569]" pageId="22" pageNumber="257">
<taxonomicName id="4C15FDD2FF8DFF85FF5E1748FDAFF989" ID-CoL="Y3GY" authority="Regimbart, 1880: 210" authorityName="Regimbart" authorityPageNumber="210" authorityYear="1880" box="[151,599,1547,1569]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF8DFF85FF5E1748FE89F989" box="[151,369,1547,1569]" italics="true" pageId="22" pageNumber="257">Copelatus tenebrosus</emphasis>
<treatmentCitation id="0AB4A040FF8DFF85FEBE1748FDAFF989" author="" box="[375,599,1547,1569]" page="210" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FEBE1748FDDCF989" box="[375,548,1547,1569]" pageId="22" pageNumber="257" refId="ref16151" refString="Regimbart, M. (1880) The new Dytiscidae and Gyrinidae collected during the recent scientific Sumatra-expedition. Notes from the Leyden Museum, 2, 209 - 216." type="journal article">Régimbart, 1880</bibRefCitation>
: 210
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA8651FF8DFF85FF5E176AFC6AF997" blockId="22.[151,914,1547,1691]" box="[151,914,1577,1599]" pageId="22" pageNumber="257">
<treatmentCitationGroup id="AB05A17FFF8DFF85FF5E176AFC6AF997" box="[151,914,1577,1599]" pageId="22" pageNumber="257">
<taxonomicName id="4C15FDD2FF8DFF85FF5E176AFDF2F997" authority="Sharp, 1882: 580" authorityName="Sharp" authorityPageNumber="580" authorityYear="1882" box="[151,522,1577,1599]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B9615A43FF8DFF85FF5E176AFEABF997" box="[151,339,1577,1599]" italics="true" pageId="22" pageNumber="257">Copelatus pusillus</emphasis>
<treatmentCitation id="0AB4A040FF8DFF85FE90176AFDF2F997" author="" box="[345,522,1577,1599]" page="580" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FE90176AFE2FF997" box="[345,471,1577,1599]" pageId="22" pageNumber="257" refId="ref16437" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article">Sharp, 1882</bibRefCitation>
: 580
</treatmentCitation>
</taxonomicName>
; synonymy by
<treatmentCitation id="0AB4A040FF8DFF85FD6A176AFC76F997" author="" box="[675,910,1577,1599]" page="296" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FD6A176AFC76F997" box="[675,910,1577,1599]" pageId="22" pageNumber="257" refId="ref16182" refString="Regimbart, M. (1899) Revision des Dytiscidae de la region Indo- Sino-Malaise. Annales de la Societe Entomologique de France, 68, 186 - 367." type="journal article">Régimbart (1899: 296)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA8651FF8DFF85FF5E170BFCC7F9F6" blockId="22.[151,914,1547,1691]" box="[151,831,1608,1630]" pageId="22" pageNumber="257">
<treatmentCitationGroup id="AB05A17FFF8DFF85FF5E170BFCC7F9F6" box="[151,831,1608,1630]" pageId="22" pageNumber="257">
<taxonomicName id="4C15FDD2FF8DFF85FF5E170BFDF8F9F6" authority="Sato, 1961: 8" authorityName="Sato" authorityPageNumber="8" authorityYear="1961" box="[151,512,1608,1630]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="hisamatsui">
<emphasis id="B9615A43FF8DFF85FF5E170BFE8AF9F6" box="[151,370,1608,1630]" italics="true" pageId="22" pageNumber="257">Copelatus hisamatsui</emphasis>
<treatmentCitation id="0AB4A040FF8DFF85FEB1170BFDF8F9F6" author="" box="[376,512,1608,1630]" page="8" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FEB1170BFE10F9F6" box="[376,488,1608,1630]" pageId="22" pageNumber="257" refId="ref16345" refString="Sato, M. (1961) Aquatic Coleoptera from Amami-Oshima of the Ryukyu Islands (II). Akitu, 10, 7 - 10." type="journal article">Satô, 1961</bibRefCitation>
: 8
</treatmentCitation>
</taxonomicName>
; synonymy by
<treatmentCitation id="0AB4A040FF8DFF85FD52170BFCC3F9F6" author="" box="[667,827,1608,1630]" page="36" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FD52170BFCC3F9F6" box="[667,827,1608,1630]" pageId="22" pageNumber="257" refId="ref16372" refString="Sato, M. (1983) Distribution of the genus Copelatus in Japan (Coleoptera: Dytiscidae). In: Sato, M. (Ed.), Special Issue Concerning the Aquatic Coleoptera Presented at the Workshop of the XUI International Congress of Entomology in Kyoto, Japan in 1980. Nagoya, 1983, pp. 35 - 41." type="journal article">Satô (1983: 36)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA8651FF8DFF85FF5E1725FD15F9D4" blockId="22.[151,914,1547,1691]" box="[151,749,1638,1660]" pageId="22" pageNumber="257">
<treatmentCitationGroup id="AB05A17FFF8DFF85FF5E1725FD15F9D4" box="[151,749,1638,1660]" pageId="22" pageNumber="257">
<taxonomicName id="4C15FDD2FF8DFF85FF5E1725FDC7F9D4" ID-CoL="8L474" authority="Vazirani, 1969: 402" authorityName="Vazirani" authorityPageNumber="402" authorityYear="1969" box="[151,575,1638,1660]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8DFF85FF5E1725FE97F9D4" box="[151,367,1638,1660]" italics="true" pageId="22" pageNumber="257">Copelatus ceylonicus</emphasis>
<treatmentCitation id="0AB4A040FF8DFF85FEBC1725FDC7F9D4" author="" box="[373,575,1638,1660]" page="402" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FEBC1725FDF4F9D4" box="[373,524,1638,1660]" pageId="22" pageNumber="257" refId="ref16474" refString="Vazirani, T. G. (1969) Two new species and notes on other species of aquatic Coleoptera from Ceylon. Spolia Zeylanica, 31, 399 - 403." type="journal article">Vazirani, 1969</bibRefCitation>
: 402
</treatmentCitation>
</taxonomicName>
;
<emphasis id="B9615A43FF8DFF85FD801724FD1FF9D4" bold="true" box="[585,743,1639,1660]" pageId="22" pageNumber="257">new synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA8651FF8DFF85FF5E17C6FD0CF933" blockId="22.[151,914,1547,1691]" box="[151,756,1669,1691]" pageId="22" pageNumber="257">
<treatmentCitationGroup id="AB05A17FFF8DFF85FF5E17C6FD0CF933" box="[151,756,1669,1691]" pageId="22" pageNumber="257">
<taxonomicName id="4C15FDD2FF8DFF85FF5E17C6FDBEF933" ID-CoL="8L46Y" authority="Vazirani, 1970: 316" authorityName="Vazirani" authorityPageNumber="316" authorityYear="1970" box="[151,582,1669,1691]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B9615A43FF8DFF85FF5E17C6FE8EF933" box="[151,374,1669,1691]" italics="true" pageId="22" pageNumber="257">Copelatus assamensis</emphasis>
<treatmentCitation id="0AB4A040FF8DFF85FEB517C6FDBEF933" author="" box="[380,582,1669,1691]" page="316" pageId="22" pageNumber="257">
<bibRefCitation id="EF84FBA0FF8DFF85FEB517C6FDEBF933" box="[380,531,1669,1691]" pageId="22" pageNumber="257" refId="ref16506" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4, 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article">Vazirani, 1970</bibRefCitation>
: 316
</treatmentCitation>
</taxonomicName>
;
<emphasis id="B9615A43FF8DFF85FD9917C5FD16F933" bold="true" box="[592,750,1670,1691]" pageId="22" pageNumber="257">new synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8DFF84FF5E178BFBA9FE04" lastPageId="23" lastPageNumber="258" pageId="22" pageNumber="257" type="materials_examined">
<paragraph id="8BAA8651FF8DFF85FF5E178BFA72F949" blockId="22.[151,1437,1736,2013]" box="[151,1418,1736,1761]" pageId="22" pageNumber="257">
<emphasis id="B9615A43FF8DFF85FF5E178BFEB0F949" bold="true" box="[151,328,1736,1761]" pageId="22" pageNumber="257">
<typeStatus id="54AE38F3FF8DFF85FF5E178BFF2AF949" box="[151,210,1736,1761]" pageId="22" pageNumber="257">Type</typeStatus>
localities.
</emphasis>
<taxonomicName id="4C15FDD2FF8DFF85FE87178BFDBDF948" authorityName="Vazirani" authorityYear="1970" box="[334,581,1736,1761]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B9615A43FF8DFF85FE87178BFDBDF948" box="[334,581,1736,1761]" italics="true" pageId="22" pageNumber="257">Copelatus assamensis</emphasis>
</taxonomicName>
: &quot;
<collectingCountry id="F302C6C1FF8DFF85FD94178BFD65F949" box="[605,669,1736,1761]" name="India" pageId="22" pageNumber="257">India</collectingCountry>
:
<collectingRegion id="49D148B3FF8DFF85FD61178AFD01F949" box="[680,761,1737,1761]" country="India" name="Assam" pageId="22" pageNumber="257">Assam</collectingRegion>
: Umrau near Shilong&quot;.
<taxonomicName id="4C15FDD2FF8DFF85FC36178AFB6EF949" authorityName="Vazirani" authorityYear="1969" box="[1023,1174,1736,1761]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8DFF85FC36178AFB6EF949" box="[1023,1174,1736,1761]" italics="true" pageId="22" pageNumber="257">C. ceylonicus</emphasis>
</taxonomicName>
: &quot;
<collectingCountry id="F302C6C1FF8DFF85FB64178BFAFBF949" box="[1197,1283,1736,1761]" name="Sri Lanka" pageId="22" pageNumber="257">Ceylon</collectingCountry>
, Colombo&quot;.
</paragraph>
<paragraph id="8BAA8651FF8DFF85FF0E17AFFA65F881" blockId="22.[151,1437,1736,2013]" pageId="22" pageNumber="257">
<emphasis id="B9615A43FF8DFF85FF0E17AFFE8FF8AD" bold="true" box="[199,375,1772,1797]" pageId="22" pageNumber="257">
<typeStatus id="54AE38F3FF8DFF85FF0E17AFFEFAF8AD" box="[199,258,1772,1797]" pageId="22" pageNumber="257">Type</typeStatus>
material.
</emphasis>
<taxonomicName id="4C15FDD2FF8DFF85FE4917AFFD81F8AC" authorityName="Vazirani" authorityYear="1970" box="[384,633,1772,1797]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B9615A43FF8DFF85FE4917AFFD81F8AC" box="[384,633,1772,1797]" italics="true" pageId="22" pageNumber="257">Copelatus assamensis</emphasis>
</taxonomicName>
:
<typeStatus id="54AE38F3FF8DFF85FD4317AFFD0AF8AD" box="[650,754,1772,1797]" pageId="22" pageNumber="257" type="holotype">Holotype</typeStatus>
<specimenCount id="9D134DD8FF8DFF85FD3217AEFCEAF8AC" box="[763,786,1773,1796]" pageId="22" pageNumber="257" type="male">♂</specimenCount>
, deposited in Zoological Survey of
<collectingCountry id="F302C6C1FF8DFF85FB7917AFFB14F8AD" box="[1200,1260,1772,1797]" name="India" pageId="22" pageNumber="257">India</collectingCountry>
, Kolkata,
<collectingCountry id="F302C6C1FF8DFF85FAAB17AFFA64F8AD" box="[1378,1436,1772,1797]" name="India" pageId="22" pageNumber="257">India</collectingCountry>
(not studied).
<taxonomicName id="4C15FDD2FF8DFF85FEFB1653FDD8F881" authorityName="Vazirani" authorityYear="1969" box="[306,544,1808,1833]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8DFF85FEFB1653FDD8F881" box="[306,544,1808,1833]" italics="true" pageId="22" pageNumber="257">Copelatus ceylonicus</emphasis>
</taxonomicName>
:
<typeStatus id="54AE38F3FF8DFF85FDE61653FD6FF881" box="[559,663,1808,1833]" pageId="22" pageNumber="257" type="holotype">Holotype</typeStatus>
<specimenCount id="9D134DD8FF8DFF85FD541652FD4AF880" box="[669,690,1809,1832]" pageId="22" pageNumber="257" type="female">♀</specimenCount>
, deposited in Colombo National Museum,
<collectingCountry id="F302C6C1FF8DFF85FB5A1653FAF9F881" box="[1171,1281,1808,1833]" name="Sri Lanka" pageId="22" pageNumber="257">
<collectingRegion id="49D148B3FF8DFF85FB5A1653FB4CF881" box="[1171,1204,1808,1833]" country="United Kingdom" name="Surrey" pageId="22" pageNumber="257">Sri</collectingRegion>
Lanka
</collectingCountry>
(not studied).
</paragraph>
<paragraph id="8BAA8651FF8DFF84FF0E1677FBA9FE04" blockId="22.[151,1437,1736,2013]" lastBlockId="23.[151,1436,151,861]" lastPageId="23" lastPageNumber="258" pageId="22" pageNumber="257">
<materialsCitation id="3B7D8C0CFF8DFF85FF0E1677FDD4F8D8" ID-GBIF-Occurrence="1917355473" collectingDate="2010-06-20" collectingDateMax="2010-06-27" collectingDateMin="2010-06-20" collectionCode="NMPC" country="Bhutan" location="Phrumsengla" municipality="Thrumshingla" pageId="22" pageNumber="257" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mongar">
<emphasis id="B9615A43FF8DFF85FF0E1677FD69F8E5" bold="true" box="[199,657,1844,1869]" pageId="22" pageNumber="257">
Additional material studied.
<collectingCountry id="F302C6C1FF8DFF85FDE61677FD74F8E5" box="[559,652,1844,1869]" name="Bhutan" pageId="22" pageNumber="257">Bhutan</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8DFF85FD541677FD38F8E4" box="[669,704,1844,1868]" pageId="22" pageNumber="257" type="male">1♂</specimenCount>
<specimenCount id="9D134DD8FF8DFF85FD041677FD17F8E4" box="[717,751,1844,1868]" pageId="22" pageNumber="257" type="female">1♀</specimenCount>
,
<collectingRegion id="49D148B3FF8DFF85FCCA1676FCA4F8E5" box="[771,860,1845,1869]" country="Bhutan" name="Mongar" pageId="22" pageNumber="257">Mongar</collectingRegion>
[district],
<collectingMunicipality id="6BCE1C2BFF8DFF85FC151677FB81F8E5" box="[988,1145,1844,1869]" pageId="22" pageNumber="257">Thrumshingla</collectingMunicipality>
[
<location id="8ECAD08AFF8DFF85FB421677FADBF8E5" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8DFF85FB421677FADBF8E5" box="[1163,1315,1844,1869]" country="Bhutan" municipality="Thrumshingla" name="Phrumsengla" pageId="22" pageNumber="257" stateProvince="Mongar">Phrumsengla</location>
] NP,
<collectingDate id="EFEF5979FF8DFF85FAA21677FEE9F8D9" pageId="22" pageNumber="257" value="2010-06-20" valueMax="2010-06-27" valueMin="2010-06-20">
<date id="FFABA091FF8DFF85FAA21677FEE9F8D9" pageId="22" pageNumber="257" value="2010-06-20" valueMax="2010-06-27" valueMin="2010-06-20">20.– 27.vi.2010</date>
</collectingDate>
, local collector (
<collectionCode id="ED041E94FF8DFF85FE07161BFDDCF8D8" box="[462,548,1880,1904]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="22" pageNumber="257">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D8C0CFF8DFF85FDFE161BFAF3F8D8" ID-GBIF-Occurrence="1917355457" box="[567,1291,1880,1905]" collectingDate="1943-06" collectionCode="GWCW" collectorName="D. E. Hardy" country="India" location="Chabua" pageId="22" pageNumber="257" specimenCount="1" specimenCount-female="1" stateProvince="Assam">
<emphasis id="B9615A43FF8DFF85FDFE161BFD78F8D9" bold="true" box="[567,640,1880,1905]" pageId="22" pageNumber="257">
<collectingCountry id="F302C6C1FF8DFF85FDFE161BFD83F8D9" box="[567,635,1880,1905]" name="India" pageId="22" pageNumber="257">India</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8DFF85FD41161BFD53F8D8" box="[648,683,1880,1904]" pageId="22" pageNumber="257" type="female">1♀</specimenCount>
,
<collectingRegion id="49D148B3FF8DFF85FD73161AFCF1F8D9" box="[698,777,1881,1905]" country="India" name="Assam" pageId="22" pageNumber="257">Assam</collectingRegion>
,
<location id="8ECAD08AFF8DFF85FCDC161BFC95F8D9" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8DFF85FCDC161BFC95F8D9" box="[789,877,1880,1905]" country="India" name="Chabua" pageId="22" pageNumber="257" stateProvince="Assam">Chabua</location>
,
<collectingDate id="EFEF5979FF8DFF85FCB3161BFC29F8D9" box="[890,977,1880,1905]" pageId="22" pageNumber="257" value="1943-06">
<date id="FFABA091FF8DFF85FCB3161BFC29F8D9" box="[890,977,1880,1905]" pageId="22" pageNumber="257" value="1943-06">vi.1943</date>
</collectingDate>
,
<collectorName id="26E0E387FF8DFF85FC17161AFBA7F8D9" box="[990,1119,1880,1905]" pageId="22" pageNumber="257">D.E. Hardy</collectorName>
leg. (
<collectionCode id="ED041E94FF8DFF85FB69161BFAFBF8D8" box="[1184,1283,1880,1904]" pageId="22" pageNumber="257">GWCW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8DFF85FADE161BFC62F83C" ID-GBIF-Occurrence="1917355401" collectingDate="1976-05-07" collectingDateMax="1976-05-09" collectingDateMin="1976-05-07" collectionCode="GWCW" collectorName="Wittmer &amp; Baroni U." country="India" elevation="75" location="Kaziranga" pageId="22" pageNumber="257" specimenCount="1" specimenCount-female="1" stateProvince="Assam">
<specimenCount id="9D134DD8FF8DFF85FADE161BFAC2F8D8" box="[1303,1338,1880,1904]" pageId="22" pageNumber="257" type="female">1♀</specimenCount>
,
<collectingRegion id="49D148B3FF8DFF85FA80161AFA60F8D9" box="[1353,1432,1881,1905]" country="India" name="Assam" pageId="22" pageNumber="257">Assam</collectingRegion>
,
<location id="8ECAD08AFF8DFF85FF5E163FFEF5F83D" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8DFF85FF5E163FFEF5F83D" box="[151,269,1916,1941]" country="India" name="Kaziranga" pageId="22" pageNumber="257" stateProvince="Assam">Kaziranga</location>
,
<elevation id="00386162FF8DFF85FED3163FFEAFF83C" box="[282,343,1916,1941]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="22" pageNumber="257" unit="m" value="75.0">
<quantity id="4CED2BB4FF8DFF85FED3163FFEAFF83C" box="[282,343,1916,1941]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="22" pageNumber="257" unit="m" value="75.0">75 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF8DFF85FEAC163FFE12F83D" box="[357,490,1916,1941]" pageId="22" pageNumber="257" value="1976-05-07" valueMax="1976-05-09" valueMin="1976-05-07">
<date id="FFABA091FF8DFF85FEAC163FFE12F83D" box="[357,490,1916,1941]" pageId="22" pageNumber="257" value="1976-05-07" valueMax="1976-05-09" valueMin="1976-05-07">7.–9.v.1976</date>
</collectingDate>
,
<collectorName id="26E0E387FF8DFF85FE31163FFDACF83D" box="[504,596,1916,1941]" pageId="22" pageNumber="257">Wittmer</collectorName>
&amp;
<collectorName id="26E0E387FF8DFF85FDB5163FFD15F83C" box="[636,749,1916,1941]" pageId="22" pageNumber="257">Baroni U.</collectorName>
leg. (
<collectionCode id="ED041E94FF8DFF85FCE6163FFC69F83C" box="[815,913,1916,1940]" pageId="22" pageNumber="257">GWCW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8DFF85FC6E163FFCB2F810" ID-GBIF-Occurrence="1917355425" collectingDate="2006-05-26" collectingDateMax="2006-06-03" collectingDateMin="2006-05-26" collectionCode="LHCM" collectorName="L. Dembicky" country="India" elevation="150" latitude="27.033333" location="Bhalukpong" longLatPrecision="1239" longitude="92.583336" pageId="22" pageNumber="257" specimenCount="5" stateProvince="Assam">
<specimenCount id="9D134DD8FF8DFF85FC6E163FFBCDF83D" box="[935,1077,1916,1941]" pageId="22" pageNumber="257" type="generic">5 specimens</specimenCount>
,
<collectingRegion id="49D148B3FF8DFF85FB8A163FFB6AF83C" box="[1091,1170,1916,1940]" country="India" name="Assam" pageId="22" pageNumber="257">Assam</collectingRegion>
,
<location id="8ECAD08AFF8DFF85FB56163FFAD3F83D" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8DFF85FB56163FFAD3F83D" box="[1183,1323,1916,1941]" country="India" latitude="27.033333" longLatPrecision="1239" longitude="92.583336" name="Bhalukpong" pageId="22" pageNumber="257" stateProvince="Assam">Bhalukpong</location>
,
<geoCoordinate id="EE21E096FF8DFF85FAF1163FFA60F83D" box="[1336,1432,1916,1941]" direction="north" orientation="latitude" pageId="22" pageNumber="257" precision="925" value="27.033333">27°02′N</geoCoordinate>
,
<geoCoordinate id="EE21E096FF8DFF85FF5E16E3FF0CF811" box="[151,244,1952,1977]" direction="east" orientation="longitude" pageId="22" pageNumber="257" precision="925" value="92.583336">92°35′E</geoCoordinate>
,
<elevation id="00386162FF8DFF85FEC916E3FEB3F810" box="[256,331,1952,1977]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="257" unit="m" value="150.0">
<quantity id="4CED2BB4FF8DFF85FEC916E3FEB3F810" box="[256,331,1952,1977]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="257" unit="m" value="150.0">150 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF8DFF85FE9E16E3FDF0F811" box="[343,520,1952,1977]" pageId="22" pageNumber="257" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">
<date id="FFABA091FF8DFF85FE9E16E3FDF0F811" box="[343,520,1952,1977]" pageId="22" pageNumber="257" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">26.v.–3.vi.2006</date>
</collectingDate>
,
<collectorName id="26E0E387FF8DFF85FDDC16E2FD52F811" box="[533,682,1952,1977]" pageId="22" pageNumber="257">L. Dembický</collectorName>
leg. (
<collectionCode id="ED041E94FF8DFF85FD2216E3FCBAF810" box="[747,834,1952,1976]" pageId="22" pageNumber="257">LHCM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8DFF85FC9E16E3FA95F810" ID-GBIF-Occurrence="1917355467" box="[855,1389,1952,1977]" collectingDate="2006-05-26" collectingDateMax="2006-06-03" collectingDateMin="2006-05-26" collectionCode="LHCM" collectorName="P. Pacholatko" country="India" elevation="150" latitude="27.033333" location="Bhalukpong" longLatPrecision="1239" longitude="92.583336" pageId="22" pageNumber="257" specimenCount="1" specimenCount-female="1" stateProvince="Assam">
<specimenCount id="9D134DD8FF8DFF85FC9E16E3FC81F810" box="[855,889,1952,1976]" pageId="22" pageNumber="257" type="female">1♀</specimenCount>
, same data, but
<collectorName id="26E0E387FF8DFF85FBFD16E2FB35F811" box="[1076,1229,1952,1977]" pageId="22" pageNumber="257">P. Pacholátko</collectorName>
leg. (
<collectionCode id="ED041E94FF8DFF85FAC716E3FA9DF810" box="[1294,1381,1952,1976]" pageId="22" pageNumber="257">LHCM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8DFF84FAB016E3FDC0FF18" ID-GBIF-Occurrence="1917355382" collectingDate="2008-04-16" collectingDateMax="2008-04-19" collectingDateMin="2008-04-16" collectionCode="NMPC" collectorName="M. Fikacek &amp; H. Podskalska &amp; P. Sipek" country="India" county="Kohora" elevation="160" lastPageId="23" lastPageNumber="258" latitude="25.583334" location="Reed" longLatPrecision="1245" longitude="93.433334" municipality="Kaziranga" pageId="22" pageNumber="257" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Assam">
<specimenCount id="9D134DD8FF8DFF85FAB016E3FA64F810" box="[1401,1436,1952,1976]" pageId="22" pageNumber="257" type="male">2♂</specimenCount>
<specimenCount id="9D134DD8FF8DFF85FF5E1687FF43F874" box="[151,187,1988,2012]" pageId="22" pageNumber="257" type="female">4♀</specimenCount>
,
<collectingCounty id="62CBFEDDFF8DFF85FF021687FEE7F875" box="[203,287,1988,2013]" pageId="22" pageNumber="257">Kohora</collectingCounty>
(=
<collectingMunicipality id="6BCE1C2BFF8DFF85FE841687FE38F875" box="[333,448,1988,2013]" pageId="22" pageNumber="257">Kaziranga</collectingMunicipality>
vill.) at green
<location id="8ECAD08AFF8DFF85FDB81687FD52F875" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8DFF85FDB81687FD52F875" box="[625,682,1988,2013]" country="India" county="Kohora" latitude="25.583334" longLatPrecision="1245" longitude="93.433334" municipality="Kaziranga" name="Reed" pageId="22" pageNumber="257" stateProvince="Assam">Reed</location>
hotel,
<geoCoordinate id="EE21E096FF8DFF85FD361687FCA6F875" box="[767,862,1988,2013]" direction="north" orientation="latitude" pageId="22" pageNumber="257" precision="925" value="25.583334">25°35′N</geoCoordinate>
,
<geoCoordinate id="EE21E096FF8DFF85FCA41687FC33F875" box="[877,971,1988,2013]" direction="east" orientation="longitude" pageId="22" pageNumber="257" precision="925" value="93.433334">93°26′E</geoCoordinate>
,
<elevation id="00386162FF8DFF85FC101687FBDDF874" box="[985,1061,1988,2013]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="257" unit="m" value="160.0">
<quantity id="4CED2BB4FF8DFF85FC101687FBDDF874" box="[985,1061,1988,2013]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="257" unit="m" value="160.0">160 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF8DFF85FBFD1687FB26F875" box="[1076,1246,1988,2013]" pageId="22" pageNumber="257" value="2008-04-16" valueMax="2008-04-19" valueMin="2008-04-16">
<date id="FFABA091FF8DFF85FBFD1687FB26F875" box="[1076,1246,1988,2013]" pageId="22" pageNumber="257" value="2008-04-16" valueMax="2008-04-19" valueMin="2008-04-16">16.–19.iv.2008</date>
</collectingDate>
,
<collectorName id="26E0E387FF8DFF85FB241686FA8BF875" box="[1261,1395,1988,2013]" pageId="22" pageNumber="257">M. Fikáček</collectorName>
,
<collectorName id="26E0E387FF8DFF84FA481686FEEDFF18" lastPageId="23" lastPageNumber="258" pageId="22" pageNumber="257">H. Podskalská</collectorName>
&amp;
<collectorName id="26E0E387FF8CFF84FEF511DBFE60FF18" box="[316,408,151,176]" pageId="23" pageNumber="258">P. Šípek</collectorName>
leg. (
<collectionCode id="ED041E94FF8CFF84FE1311DBFDC8FF18" box="[474,560,152,176]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="258">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8CFF84FD8C11DBFCBEFF7C" ID-GBIF-Occurrence="1917355429" collectingDate="2012-05-01" collectingDateMax="2012-05-08" collectingDateMin="2012-05-01" collectionCode="NMPC, ZMFK" collectorName="L. Dembicky" country="India" elevation="150" latitude="27.013334" location="Bhalukpong" longLatPrecision="123" longitude="92.651665" municipality="Assam-Arunachal" pageId="23" pageNumber="258" specimenCount="3" stateProvince="Assam">
<specimenCount id="9D134DD8FF8CFF84FD8C11DBFD2BFF18" box="[581,723,151,176]" pageId="23" pageNumber="258" type="generic">3 specimens</specimenCount>
,
<collectingMunicipality id="6BCE1C2BFF8CFF84FD2911D4FC52FF18" box="[736,938,151,176]" pageId="23" pageNumber="258">Assam-Arunachal</collectingMunicipality>
border,
<location id="8ECAD08AFF8CFF84FBC211D4FB6FFF18" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FBC211D4FB6FFF18" box="[1035,1175,151,176]" country="India" latitude="27.013334" longLatPrecision="123" longitude="92.651665" municipality="Assam-Arunachal" name="Bhalukpong" pageId="23" pageNumber="258" stateProvince="Assam">Bhalukpong</location>
,
<geoCoordinate id="EE21E096FF8CFF84FB6A11D4FAE1FF18" box="[1187,1305,151,176]" direction="north" orientation="latitude" pageId="23" pageNumber="258" precision="92" value="27.013334">27°00.8′N</geoCoordinate>
,
<geoCoordinate id="EE21E096FF8CFF84FAEF11D4FA6FFF18" box="[1318,1431,151,176]" direction="east" orientation="longitude" pageId="23" pageNumber="258" precision="92" value="92.651665">92°39.1′E</geoCoordinate>
,
<elevation id="00386162FF8CFF84FF5E11FFFF10FF7C" box="[151,232,188,213]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="258" unit="m" value="150.0">
<quantity id="4CED2BB4FF8CFF84FF5E11FFFF10FF7C" box="[151,232,188,213]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="258" unit="m" value="150.0">150 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF8CFF84FF3411FFFE7AFF7C" box="[253,386,188,212]" pageId="23" pageNumber="258" value="2012-05-01" valueMax="2012-05-08" valueMin="2012-05-01">
<date id="FFABA091FF8CFF84FF3411FFFE7AFF7C" box="[253,386,188,212]" pageId="23" pageNumber="258" value="2012-05-01" valueMax="2012-05-08" valueMin="2012-05-01">1.–8.v.2012</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FE5E11FEFDCBFF7D" box="[407,563,188,213]" pageId="23" pageNumber="258">L. Dembický</collectorName>
leg. (
<collectionCode id="ED041E94FF8CFF84FD4A11FFFD2FFF7C" box="[643,727,188,212]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="258">NMPC</collectionCode>
,
<collectionCode id="ED041E94FF8CFF84FD2211FFFCC6FF7C" box="[747,830,188,212]" pageId="23" pageNumber="258">ZMFK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8CFF84FC9311FFFDC8FF50" ID-GBIF-Occurrence="1917355417" collectingDate="1984-11-09" collectingDateMax="1984-11-15" collectingDateMin="1984-11-09" collectionCode="NHMB" collectorName="B. Bhakta" country="India" county="Darjeeling" location="Mahahandra river" municipality="Sonapur" pageId="23" pageNumber="258" specimenCount="1" specimenCount-male="1" stateProvince="Assam">
<specimenCount id="9D134DD8FF8CFF84FC9311FFFC86FF7C" box="[858,894,188,212]" pageId="23" pageNumber="258" type="male">1♂</specimenCount>
,
<collectingCounty id="62CBFEDDFF8CFF84FC5A11FFFBF4FF7D" box="[915,1036,188,213]" pageId="23" pageNumber="258">Darjeeling</collectingCounty>
,
<collectingMunicipality id="6BCE1C2BFF8CFF84FBE911FFFB87FF7C" box="[1056,1151,188,212]" pageId="23" pageNumber="258">Sonapur</collectingMunicipality>
,
<location id="8ECAD08AFF8CFF84FB5A11FFFA9EFF7D" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FB5A11FFFA9EFF7D" box="[1171,1382,188,213]" country="India" county="Darjeeling" municipality="Sonapur" name="Mahahandra river" pageId="23" pageNumber="258" stateProvince="Assam">Mahahandra river</location>
,
<collectingDate id="EFEF5979FF8CFF84FAB011FFFEE8FF50" pageId="23" pageNumber="258" value="1984-11-09" valueMax="1984-11-15" valueMin="1984-11-09">
<date id="FFABA091FF8CFF84FAB011FFFEE8FF50" pageId="23" pageNumber="258" value="1984-11-09" valueMax="1984-11-15" valueMin="1984-11-09">9.– 15.xi.1984</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FED411A3FE75FF50" box="[285,397,223,248]" pageId="23" pageNumber="258">B. Bhakta</collectorName>
leg. (
<collectionCode id="ED041E94FF8CFF84FE0711A3FDD0FF50" box="[462,552,224,248]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="23" pageNumber="258">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D8C0CFF8CFF84FDF2119CFD87FEB4" ID-GBIF-Occurrence="1917355398" collectingDate="2000-07-21" collectingDateMax="2000-07-27" collectingDateMin="2000-07-21" collectionCode="NMPC" collectorName="J. Schneider" country="Nepal" elevation="166" latitude="27.35" location="Sauraha" longLatPrecision="742" longitude="84.3" municipality="Chitwan Roy." pageId="23" pageNumber="258" specimenCount="1" specimenCount-female="1">
<emphasis id="B9615A43FF8CFF84FDF2119CFD72FF50" bold="true" box="[571,650,223,248]" pageId="23" pageNumber="258">
<collectingCountry id="F302C6C1FF8CFF84FDF2119CFD7DFF50" box="[571,645,223,248]" name="Nepal" pageId="23" pageNumber="258">Nepal</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8CFF84FD5811A3FD4CFF50" box="[657,692,224,248]" pageId="23" pageNumber="258" type="female">1♀</specimenCount>
,
<collectingMunicipality id="6BCE1C2BFF8CFF84FD0A119CFCA6FF50" box="[707,862,223,248]" pageId="23" pageNumber="258">Chitwan Roy.</collectingMunicipality>
NP,
<location id="8ECAD08AFF8CFF84FC5D119CFC16FF50" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FC5D119CFC16FF50" box="[916,1006,223,248]" country="Nepal" latitude="27.35" longLatPrecision="742" longitude="84.3" municipality="Chitwan Roy." name="Sauraha" pageId="23" pageNumber="258">Sauraha</location>
vill.,
<geoCoordinate id="EE21E096FF8CFF84FBF9119CFB7EFF50" box="[1072,1158,223,248]" direction="north" orientation="latitude" pageId="23" pageNumber="258" precision="555" value="27.35">27.35N</geoCoordinate>
,
<geoCoordinate id="EE21E096FF8CFF84FB5B11A3FB1DFF50" box="[1170,1253,224,248]" direction="east" orientation="longitude" pageId="23" pageNumber="258" precision="555" value="84.3">84.30E</geoCoordinate>
,
<quantity id="4CED2BB4FF8CFF84FB39119CFAC2FF5F" box="[1264,1338,223,248]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="23" pageNumber="258" unit="m" value="166.0">
<elevation id="00386162FF8CFF84FB39119CFAC2FF5F" box="[1264,1338,223,248]" metricMagnitude="2" metricUnit="m" metricValue="1.66" pageId="23" pageNumber="258" unit="m" value="166.0">166 m</elevation>
</quantity>
, at light,
<collectingDate id="EFEF5979FF8CFF84FF5E1047FEB2FEB5" box="[151,330,260,285]" pageId="23" pageNumber="258" value="2000-07-21" valueMax="2000-07-27" valueMin="2000-07-21">
<date id="FFABA091FF8CFF84FF5E1047FEB2FEB5" box="[151,330,260,285]" pageId="23" pageNumber="258" value="2000-07-21" valueMax="2000-07-27" valueMin="2000-07-21">21.–27.vii.2000</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FE9F1046FE18FEB5" box="[342,480,260,285]" pageId="23" pageNumber="258">J. Schneider</collectorName>
leg. (
<collectionCode id="ED041E94FF8CFF84FDE81047FD8FFEB4" box="[545,631,260,284]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="258">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8CFF84FD451047FBA5FEB4" ID-GBIF-Occurrence="1917355414" box="[652,1117,260,285]" collectingDate="2000-07-21" collectingDateMax="2000-07-27" collectingDateMin="2000-07-21" collectionCode="NMPC" collectorName="D. Kral" country="Nepal" elevation="166" latitude="27.35" location="Sauraha" longLatPrecision="742" longitude="84.3" municipality="Chitwan Roy." pageId="23" pageNumber="258" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D134DD8FF8CFF84FD451047FD56FEB4" box="[652,686,260,284]" pageId="23" pageNumber="258" type="female">1♀</specimenCount>
, same data, but
<collectorName id="26E0E387FF8CFF84FCA01046FC46FEB5" box="[873,958,260,285]" pageId="23" pageNumber="258">D. Král</collectorName>
leg. (
<collectionCode id="ED041E94FF8CFF84FC361047FBADFEB4" box="[1023,1109,260,284]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="23" pageNumber="258">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7D8C0CFF8CFF84FBA11047FB13FEE8" ID-GBIF-Occurrence="1917355390" collectingDate="1979-12-04" collectingDateMax="1979-12-07" collectingDateMin="1979-12-04" collectionCode="ZMUC" collectorName="V. Mahler Jensen" country="Sri Lanka" determinerName="T. G. Vazirani" location="Sinharaya" pageId="23" pageNumber="258" specimenCount="1" specimenCount-female="1" stateProvince="Sri">
<emphasis id="B9615A43FF8CFF84FBA11047FB12FEB5" bold="true" box="[1128,1258,260,285]" pageId="23" pageNumber="258">
<collectingCountry id="F302C6C1FF8CFF84FBA11047FB1EFEB5" box="[1128,1254,260,285]" name="Sri Lanka" pageId="23" pageNumber="258">
<collectingRegion id="49D148B3FF8CFF84FBA11047FB74FEB5" box="[1128,1164,260,285]" country="United Kingdom" name="Surrey" pageId="23" pageNumber="258">Sri</collectingRegion>
Lanka
</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8CFF84FB3A1047FAEEFEB4" box="[1267,1302,260,284]" pageId="23" pageNumber="258" type="female">1♀</specimenCount>
,
<location id="8ECAD08AFF8CFF84FAEC1047FA60FEB5" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FAEC1047FA60FEB5" box="[1317,1432,260,285]" country="Sri Lanka" name="Sinharaya" pageId="23" pageNumber="258" stateProvince="Sri">Sinharaya</location>
,
<collectingDate id="EFEF5979FF8CFF84FF5E1064FED6FEE8" box="[151,302,295,320]" pageId="23" pageNumber="258" value="1979-12-04" valueMax="1979-12-07" valueMin="1979-12-04">
<date id="FFABA091FF8CFF84FF5E1064FED6FEE8" box="[151,302,295,320]" pageId="23" pageNumber="258" value="1979-12-04" valueMax="1979-12-07" valueMin="1979-12-04">4.–7.xii.1979</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FEF2106BFE06FE97" box="[315,510,295,320]" pageId="23" pageNumber="258">V. Mahler Jensen</collectorName>
lgt.,
<taxonomicName id="4C15FDD2FF8CFF84FDF31064FC93FEE8" authority="Reg." authorityName="Regimbart" authorityYear="1880" box="[570,875,295,320]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF8CFF84FDF31064FCD4FEE8" box="[570,812,295,320]" italics="true" pageId="23" pageNumber="258">Copelatus tenebrosus</emphasis>
Rég.
</taxonomicName>
, det.
<determinerName id="7E766A95FF8CFF84FC61106BFBC3FEE8" box="[936,1083,295,320]" pageId="23" pageNumber="258">T.G. Vazirani</determinerName>
1981 (
<collectionCode id="ED041E94FF8CFF84FB45106BFB1BFEE8" box="[1164,1251,296,320]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="23" pageNumber="258">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8CFF84FB3E106BFB34FECC" ID-GBIF-Occurrence="1917355437" collectingDate="1979-12-20" collectionCode="ZMUC" collectorName="V. Mahler Jensen" country="Sri Lanka" determinerName="T. G. Vazirani" location="Hanwella" pageId="23" pageNumber="258" specimenCount="1" specimenCount-male="1" stateProvince="Sri">
<specimenCount id="9D134DD8FF8CFF84FB3E106BFAE4FEE8" box="[1271,1308,296,320]" pageId="23" pageNumber="258" type="male">1♂</specimenCount>
,
<location id="8ECAD08AFF8CFF84FAE31064FA60FEE8" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FAE31064FA60FEE8" box="[1322,1432,295,320]" country="Sri Lanka" name="Hanwella" pageId="23" pageNumber="258" stateProvince="Sri">Hanwella</location>
,
<collectingDate id="EFEF5979FF8CFF84FF5E100FFEE1FECD" box="[151,281,332,357]" pageId="23" pageNumber="258" value="1979-12-20">
<date id="FFABA091FF8CFF84FF5E100FFEE1FECD" box="[151,281,332,357]" pageId="23" pageNumber="258" value="1979-12-20">20.xii.1979</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FEEC100EFE10FECC" box="[293,488,332,357]" pageId="23" pageNumber="258">V. Mahler Jensen</collectorName>
lgt.,
<taxonomicName id="4C15FDD2FF8CFF84FDEB100FFCB6FECC" authority="Vaz." authorityName="Vazirani" authorityYear="1969" box="[546,846,332,357]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8CFF84FDEB100FFCE9FECD" box="[546,785,332,357]" italics="true" pageId="23" pageNumber="258">Copelatus ceylonicus</emphasis>
Vaz.
</taxonomicName>
, det.
<determinerName id="7E766A95FF8CFF84FC43100FFBE5FECD" box="[906,1053,332,357]" pageId="23" pageNumber="258">T.G. Vazirani</determinerName>
1981 (
<collectionCode id="ED041E94FF8CFF84FBA5100FFB3BFECC" box="[1132,1219,332,356]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="23" pageNumber="258">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8CFF84FB1E100FFCAAFE20" ID-GBIF-Occurrence="1917355418" collectingDate="1979-12-14" collectionCode="ZMUC" collectorName="V. Mahler Jensen" country="Sri Lanka" location="Inginiyagala" pageId="23" pageNumber="258" specimenCount="2" specimenCount-male="2" stateProvince="Sri">
<specimenCount id="9D134DD8FF8CFF84FB1E100FFB05FECC" box="[1239,1277,332,356]" pageId="23" pageNumber="258" type="male">2♂</specimenCount>
,
<location id="8ECAD08AFF8CFF84FAC0100FFA60FECD" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FAC0100FFA60FECD" box="[1289,1432,332,357]" country="Sri Lanka" name="Inginiyagala" pageId="23" pageNumber="258" stateProvince="Sri">Inginiyagala</location>
,
<collectingDate id="EFEF5979FF8CFF84FF5E102CFEE1FE20" box="[151,281,367,392]" pageId="23" pageNumber="258" value="1979-12-14">
<date id="FFABA091FF8CFF84FF5E102CFEE1FE20" box="[151,281,367,392]" pageId="23" pageNumber="258" value="1979-12-14">14.xii.1979</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FEEC1033FE1FFE2F" box="[293,487,367,392]" pageId="23" pageNumber="258">V. Mahler Jensen</collectorName>
lgt. [as
<taxonomicName id="4C15FDD2FF8CFF84FD8B1033FD22FE20" authorityName="Vazirani" authorityYear="1969" box="[578,730,367,392]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8CFF84FD8B1033FD22FE20" box="[578,730,367,392]" italics="true" pageId="23" pageNumber="258">C. ceylonicus</emphasis>
</taxonomicName>
] (
<collectionCode id="ED041E94FF8CFF84FD3B1033FCB1FE20" box="[754,841,368,392]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="23" pageNumber="258">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8CFF84FC941033FE59FE04" ID-GBIF-Occurrence="1917355389" collectingDate="1980-11-08" collectionCode="NHMW, GWCW" collectorName="M. A. Jach" country="Sri Lanka" location="Colombo" pageId="23" pageNumber="258" specimenCount="20" stateProvince="Sri">
<specimenCount id="9D134DD8FF8CFF84FC941033FC00FE20" box="[861,1016,367,392]" pageId="23" pageNumber="258" type="generic">20 specimens</specimenCount>
,
<location id="8ECAD08AFF8CFF84FBCD102CFB94FE20" LSID="urn:lsid:plazi:treatment:03BC3747FF8DFF8BFF5E1487FD41FE78:8ECAD08AFF8CFF84FBCD102CFB94FE20" box="[1028,1132,367,392]" country="Sri Lanka" name="Colombo" pageId="23" pageNumber="258" stateProvince="Sri">Colombo</location>
env.,
<collectingDate id="EFEF5979FF8CFF84FB66102CFAE4FE20" box="[1199,1308,367,392]" pageId="23" pageNumber="258" value="1980-11-08">
<date id="FFABA091FF8CFF84FB66102CFAE4FE20" box="[1199,1308,367,392]" pageId="23" pageNumber="258" value="1980-11-08">8.xi.1980</date>
</collectingDate>
,
<collectorName id="26E0E387FF8CFF84FAE11033FA64FE20" box="[1320,1436,367,392]" pageId="23" pageNumber="258">M.A. Jäch</collectorName>
leg. (
<collectionCode id="ED041E94FF8CFF84FF0710D7FED4FE04" box="[206,300,404,428]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="258">NHMW</collectionCode>
,
<collectionCode id="ED041E94FF8CFF84FEFE10D7FE60FE04" box="[311,408,404,428]" pageId="23" pageNumber="258">GWCW</collectionCode>
)
</materialsCitation>
; 1♂, Galle, Unawatuna,
<date id="FFABA091FF8CFF84FD7C10D7FCD7FE05" box="[693,815,404,429]" pageId="23" pageNumber="258" value="2002-11-30">30.xi.2002</date>
, M. Janalík leg. (NMPC).
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8CFF84FF0E10F4FB49FD65" pageId="23" pageNumber="258" type="discussion">
<paragraph id="8BAA8651FF8CFF84FF0E10F4FB49FD65" blockId="23.[151,1436,151,861]" pageId="23" pageNumber="258">
<emphasis id="B9615A43FF8CFF84FF0E10F4FDE1FE78" bold="true" box="[199,537,439,464]" pageId="23" pageNumber="258">Comments on classification.</emphasis>
<bibRefCitation id="EF84FBA0FF8CFF84FDEB10F4FD2BFE78" box="[546,723,439,464]" pageId="23" pageNumber="258" refId="ref16474" refString="Vazirani, T. G. (1969) Two new species and notes on other species of aquatic Coleoptera from Ceylon. Spolia Zeylanica, 31, 399 - 403." type="journal article">Vazirani (1969)</bibRefCitation>
described
<taxonomicName id="4C15FDD2FF8CFF84FC9810FBFC12FE78" authorityName="Vazirani" authorityYear="1969" box="[849,1002,439,464]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8CFF84FC9810FBFC12FE78" box="[849,1002,439,464]" italics="true" pageId="23" pageNumber="258">C. ceylonicus</emphasis>
</taxonomicName>
based on a single female collected in 'pond in Museum' in Colombo city. The author mentioned its similarity to
<taxonomicName id="4C15FDD2FF8CFF84FC1E109EFB88FE5D" authorityName="Regimbart" authorityYear="1880" box="[983,1136,476,501]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF8CFF84FC1E109EFB88FE5D" box="[983,1136,476,501]" italics="true" pageId="23" pageNumber="258">C. tenebrosus</emphasis>
</taxonomicName>
and differentiated the new species by the more subparallel habitus, rufo-ferruginous coloration and by the elytral striae terminating apically at the same level. We had no possibility to see the holotype, but have studied three males identified as
<taxonomicName id="4C15FDD2FF8CFF84FACD1366FA64FD95" authorityName="Vazirani" authorityYear="1969" box="[1284,1436,548,573]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8CFF84FACD1366FA64FD95" box="[1284,1436,548,573]" italics="true" pageId="23" pageNumber="258">C. ceylonicus</emphasis>
</taxonomicName>
by Vazirani and deposited in ZMUC (
<bibRefCitation id="EF84FBA0FF8CFF84FD811304FC96FDC8" box="[584,878,583,608]" pageId="23" pageNumber="258" refId="ref15798" refString="Holmen, M. &amp; Vazirani, T. G. (1990) Notes on the genera Neptosternus Sharp and Copelatus Erichson from Sri Lanka and India with the description of new species (Coleoptera: Dytiscidae). Koleopterologische Rundschau, 60, 19 - 32." type="journal article">Holmen &amp; Vazirani 1990</bibRefCitation>
). They are all teneral, which explains their more subparallel habitus and rufo-ferruginous coloration. Otherwise, they agree well with typical specimens of
<taxonomicName id="4C15FDD2FF8CFF84FA4B132EFEF7FD00" authorityName="Regimbart" authorityYear="1880" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF8CFF84FA4B132EFEF7FD00" italics="true" pageId="23" pageNumber="258">C. tenebrosus</emphasis>
</taxonomicName>
from Sri Lanka and other countries across its area of distribution; the length of the dorsal elytral striae is variable even within specimens in one population and it is not usable for species delimitation.
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8CFF8BFF0E1394FE51FECD" lastPageId="24" lastPageNumber="259" pageId="23" pageNumber="258" type="reference_group">
<paragraph id="8BAA8651FF8CFF8BFF0E1394FE51FECD" blockId="23.[151,1436,151,861]" lastBlockId="24.[151,1437,151,464]" lastPageId="24" lastPageNumber="259" pageId="23" pageNumber="258">
<taxonomicName id="4C15FDD2FF8CFF84FF0E1394FE46FD58" ID-CoL="8L46Y" authorityName="Vazirani" authorityYear="1970" box="[199,446,727,752]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B9615A43FF8CFF84FF0E1394FE46FD58" box="[199,446,727,752]" italics="true" pageId="23" pageNumber="258">Copelatus assamensis</emphasis>
</taxonomicName>
was described based on a single male specimen. It was compared with
<taxonomicName id="4C15FDD2FF8CFF84FB2C139BFA64FD58" ID-CoL="Y336" authorityName="Regimbart" authorityYear="1899" box="[1253,1436,727,752]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="B9615A43FF8CFF84FB2C139BFA64FD58" box="[1253,1436,727,752]" italics="true" pageId="23" pageNumber="258">C. andamanicus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C15FDD2FF8CFF84FF7113BEFEDDFCBD" ID-CoL="Y3DC" authorityName="Sharp" authorityYear="1882" box="[184,293,764,789]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="oblitus">
<emphasis id="B9615A43FF8CFF84FF7113BEFEDDFCBD" box="[184,293,764,789]" italics="true" pageId="23" pageNumber="258">C. oblitus</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C15FDD2FF8CFF84FEAF13BEFD6CFCBD" ID-CoL="Y3BS" authority="Guignot, 1954" authorityName="Guignot" authorityYear="1954" box="[358,660,764,789]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="B9615A43FF8CFF84FEAF13BEFE1EFCBD" box="[358,486,764,789]" italics="true" pageId="23" pageNumber="258">C. malaisei</emphasis>
Guignot, 1954
</taxonomicName>
, but actually it was keyed with
<taxonomicName id="4C15FDD2FF8CFF84FC3313BEFB6BFCBD" ID-CoL="Y3GY" authorityName="Regimbart" authorityYear="1880" box="[1018,1171,764,789]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF8CFF84FC3313BEFB6BFCBD" box="[1018,1171,764,789]" italics="true" pageId="23" pageNumber="258">C. tenebrosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C15FDD2FF8CFF84FB0213BEFA9AFCBD" ID-CoL="8L474" authorityName="Vazirani" authorityYear="1969" box="[1227,1378,764,789]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF8CFF84FB0213BEFA9AFCBD" box="[1227,1378,764,789]" italics="true" pageId="23" pageNumber="258">C. ceylonicus</emphasis>
</taxonomicName>
with only one difference mentioned: dorsal elytral striae equally abridged apically (i.e. terminated at the same level). As mentioned above, the length of the elytral striae is variable, and moreover the same character was also mentioned in description of
<taxonomicName id="4C15FDD2FF83FF8BFE9311DBFE0AFF18" ID-CoL="8L474" authorityName="Vazirani" authorityYear="1969" box="[346,498,151,176]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF83FF8BFE9311DBFE0AFF18" box="[346,498,151,176]" italics="true" pageId="24" pageNumber="259">C. ceylonicus</emphasis>
</taxonomicName>
. We had no possibility to study the
<typeStatus id="54AE38F3FF83FF8BFC4611D4FC0CFF18" box="[911,1012,151,176]" pageId="24" pageNumber="259" type="holotype">holotype</typeStatus>
, however all characters mentioned in the original description, especially the testaceous surface of the pronotum and the elytra, body size
<quantity id="4CED2BB4FF83FF8BFADB11FFFA92FF7C" box="[1298,1386,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="24" pageNumber="259" unit="mm" value="4.9">4.9 mm</quantity>
and sketchy drawing of median lobe fits with extensive material of
<taxonomicName id="4C15FDD2FF83FF8BFCAB11A3FC03FF50" ID-CoL="Y3GY" authorityName="Regimbart" authorityYear="1880" box="[866,1019,223,248]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF83FF8BFCAB11A3FC03FF50" box="[866,1019,223,248]" italics="true" pageId="24" pageNumber="259">C. tenebrosus</emphasis>
</taxonomicName>
from
<collectingRegion id="49D148B3FF83FF8BFB8811A3FB75FF50" box="[1089,1165,224,248]" country="India" name="Assam" pageId="24" pageNumber="259">Assam</collectingRegion>
and neighbouring areas in our hands. Therefore, we have no doubts about its identity and establish the following new synonymies:
<taxonomicName id="4C15FDD2FF83FF8BFF5E1064FDA8FEE8" ID-CoL="Y3GY" authority="Regimbart, 1880" authorityName="Regimbart" authorityYear="1880" box="[151,592,295,320]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF83FF8BFF5E1064FE71FEE8" box="[151,393,295,320]" italics="true" pageId="24" pageNumber="259">Copelatus tenebrosus</emphasis>
<bibRefCitation id="EF84FBA0FF83FF8BFE5B1064FDA8FEE8" box="[402,592,295,320]" pageId="24" pageNumber="259" refId="ref16151" refString="Regimbart, M. (1880) The new Dytiscidae and Gyrinidae collected during the recent scientific Sumatra-expedition. Notes from the Leyden Museum, 2, 209 - 216." type="journal article">Régimbart, 1880</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C15FDD2FF83FF8BFDBB1064FBE8FEE8" ID-CoL="8L474" authority="Vazirani, 1969" authorityName="Vazirani" authorityYear="1969" box="[626,1040,295,320]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="species" species="ceylonicus">
<emphasis id="B9615A43FF83FF8BFDBB1064FC9AFEE8" box="[626,866,295,320]" italics="true" pageId="24" pageNumber="259">Copelatus ceylonicus</emphasis>
<bibRefCitation id="EF84FBA0FF83FF8BFCA21064FBE8FEE8" box="[875,1040,295,320]" pageId="24" pageNumber="259" refId="ref16474" refString="Vazirani, T. G. (1969) Two new species and notes on other species of aquatic Coleoptera from Ceylon. Spolia Zeylanica, 31, 399 - 403." type="journal article">Vazirani, 1969</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A252E738FF83FF8BFBD01064FB7AFEE8" box="[1049,1154,295,320]" pageId="24" pageNumber="259">
<emphasis id="B9615A43FF83FF8BFBD01064FB7AFEE8" bold="true" box="[1049,1154,295,320]" pageId="24" pageNumber="259">syn. nov.</emphasis>
</taxonomicNameLabel>
=
<taxonomicName id="4C15FDD2FF83FF8BFB6D1064FEC2FECC" ID-CoL="8L46Y" authority="Vazirani, 1970" authorityName="Vazirani" authorityYear="1970" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B9615A43FF83FF8BFB6D1064FA64FEE8" box="[1188,1436,295,320]" italics="true" pageId="24" pageNumber="259">Copelatus assamensis</emphasis>
<bibRefCitation id="EF84FBA0FF83FF8BFF5E100FFEC2FECC" box="[151,314,332,357]" pageId="24" pageNumber="259" refId="ref16506" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4, 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article">Vazirani, 1970</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A252E738FF83FF8BFE88100FFE51FECD" box="[321,425,332,357]" pageId="24" pageNumber="259">
<emphasis id="B9615A43FF83FF8BFE88100FFE51FECD" bold="true" box="[321,425,332,357]" pageId="24" pageNumber="259">syn. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF6AD6D9FF8CFF84FF5E16DCFB5CF87D" httpUri="https://zenodo.org/record/1458572/files/figure.png" pageId="23" pageNumber="258" startId="23.[151,263,1951,1974]" targetBox="[192,1412,971,1918]" targetPageId="23">
<paragraph id="8BAA8651FF8CFF84FF5E16DCFB5CF87D" blockId="23.[151,1436,1951,2005]" pageId="23" pageNumber="258">
<emphasis id="B9615A43FF8CFF84FF5E16DCFEAEF81D" bold="true" box="[151,342,1951,1974]" pageId="23" pageNumber="258">FIGURES 49–51.</emphasis>
<taxonomicName id="4C15FDD2FF8CFF84FEA916E3FDCCF81E" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[352,564,1952,1974]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8CFF84FEA916E3FDCCF81E" box="[352,564,1952,1974]" italics="true" pageId="23" pageNumber="258">Copelatus sociennus</emphasis>
</taxonomicName>
. 49. male habitus (Nepal); 50. median lobe in lateral view (redrawn from original description of
<taxonomicName id="4C15FDD2FF8CFF84FEE01683FE2DF87D" authorityName="Vazirani" authorityYear="1970" box="[297,469,1983,2005]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8CFF84FEE01683FE2DF87D" box="[297,469,1983,2005]" italics="true" pageId="23" pageNumber="258">C. bangalorensis</emphasis>
</taxonomicName>
); 51. same (paratype of
<taxonomicName id="4C15FDD2FF8CFF84FD051683FCB6F87D" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[716,846,1984,2005]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="258" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8CFF84FD051683FCB6F87D" box="[716,846,1984,2005]" italics="true" pageId="23" pageNumber="258">C. sociennus</emphasis>
</taxonomicName>
). Scale bar 0.5 mm (Figs 50–51).
</paragraph>
</caption>
<subSubSection id="C30FD5DAFF83FF8BFF0E102CFD41FE78" pageId="24" pageNumber="259" type="distribution">
<paragraph id="8BAA8651FF83FF8BFF0E102CFD41FE78" blockId="24.[151,1437,151,464]" pageId="24" pageNumber="259">
<emphasis id="B9615A43FF83FF8BFF0E102CFE98FE20" bold="true" box="[199,352,367,392]" pageId="24" pageNumber="259">Distribution.</emphasis>
<taxonomicName id="4C15FDD2FF83FF8BFEA0102CFDA4FE20" authorityName="Regimbart" authorityYear="1880" box="[361,604,367,392]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="B9615A43FF83FF8BFEA0102CFDA4FE20" box="[361,604,367,392]" italics="true" pageId="24" pageNumber="259">Copelatus tenebrosus</emphasis>
</taxonomicName>
is one of the most widespread
<taxonomicName id="4C15FDD2FF83FF8BFC01102CFBC1FE20" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[968,1081,367,392]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="B9615A43FF83FF8BFC01102CFBC1FE20" box="[968,1081,367,392]" italics="true" pageId="24" pageNumber="259">Copelatus</emphasis>
</taxonomicName>
species. It occurs in the entire
<collectingRegion id="49D148B3FF83FF8BFF5E10D7FF0BFE05" box="[151,243,404,429]" country="Democratic Republic of the Congo" name="Orientale" pageId="24" pageNumber="259">Oriental</collectingRegion>
region, in southern
<collectingCountry id="F302C6C1FF83FF8BFE1B10D7FDEDFE05" box="[466,533,404,429]" name="China" pageId="24" pageNumber="259">China</collectingCountry>
and
<collectingCountry id="F302C6C1FF83FF8BFD8210D6FD73FE04" box="[587,651,405,428]" name="Japan" pageId="24" pageNumber="259">Japan</collectingCountry>
in the Palaearctic region, and also reaches
<collectingCountry id="F302C6C1FF83FF8BFBA210D7FB2DFE05" box="[1131,1237,404,429]" name="Australia" pageId="24" pageNumber="259">Australia</collectingCountry>
. Here, we present the first country records from
<collectingCountry id="F302C6C1FF83FF8BFE2E10F4FDC0FE78" box="[487,568,439,464]" name="Bhutan" pageId="24" pageNumber="259">Bhutan</collectingCountry>
and
<collectingCountry id="F302C6C1FF83FF8BFDA610F4FD4EFE78" box="[623,694,439,464]" name="Nepal" pageId="24" pageNumber="259">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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