305 lines
49 KiB
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305 lines
49 KiB
XML
<document id="DDF17A78D9F165632AB65EF050D7BB64" ID-DOI="10.11646/zootaxa.5536.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239134" ID-ZooBank="94B45D1F-916F-4CE7-B022-C3D941E0CF9E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732799161048" checkinUser="plazi" docAuthor="Mikhaljova, Elena V., Kazarin, Vitaly M. & Marusik, Yury M." docDate="2024" docId="03B51125FF80E30DFCF6FF037DF9D6AF" docLanguage="en" docName="zootaxa.5536.3.8.pdf" docOrigin="Zootaxa 5536 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Skleroprotopus securifer Mikhaljova, Kazarin & Marusik, 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="488" masterDocId="FF8C695DFF83E308FC61FF94792BD656" masterDocTitle="The millipede genus Skleroprotopus Attems, 1901 (Diplopoda, Julida, Mongoliulidae) in China, with description of a new species" masterLastPageNumber="491" masterPageNumber="483" pageNumber="486" updateTime="1732920472400" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="9D3F93166FF33777FA19329B14A7EB7A">
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<mods:title id="AB52602C857D76C93056E3C103E2BE5B">The millipede genus Skleroprotopus Attems, 1901 (Diplopoda, Julida, Mongoliulidae) in China, with description of a new species</mods:title>
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<mods:name id="2AD87D0D64406B99F7C3A2079329B0FE" type="personal">
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<mods:roleTerm id="92895EE3BE75D3C2D6E00851CBD0A4EB">Author</mods:roleTerm>
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<mods:namePart id="5D3EAF22DFA4C43B4B62684FD796947A">Mikhaljova, Elena V.</mods:namePart>
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<mods:nameIdentifier id="8C3E57857170D453B152762D5F15BD7D" type="ORCID">0000-0001-7132-8677</mods:nameIdentifier>
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<mods:affiliation id="474909900C675BE3D1D83122A981122D">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
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<mods:nameIdentifier id="7C3B4A6E3FA783FD4C4E65C4A74EA90C" type="email">mikhaljova@biosoil.ru</mods:nameIdentifier>
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<mods:name id="0D519BBA138ED706A818524257BB9720" type="personal">
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<mods:roleTerm id="A43ACF56A0F85592C6F5D798210A09F3">Author</mods:roleTerm>
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<mods:namePart id="1BFB3E67EAE51FBB04CBCC0378FA3089">Kazarin, Vitaly M.</mods:namePart>
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<mods:nameIdentifier id="58BBF8DB567F1B12557D90E3BCF7DC5E" type="ORCID">0009-0001-3771-4782</mods:nameIdentifier>
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<mods:affiliation id="197A2A0596D7621C32129BFF1C74D70C">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
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<mods:nameIdentifier id="71E75D1CF2A4091E45F9D926CE752E5E" type="email">kazarin@biosoil.ru</mods:nameIdentifier>
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<mods:name id="730F2C8770742B5812B4EF80E9840655" type="personal">
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<mods:roleTerm id="9B3B05731BFEEE87350B950B3B94CE75">Author</mods:roleTerm>
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<mods:namePart id="D8A5272D0EF65F2BF5D1FDD9868A821C">Marusik, Yury M.</mods:namePart>
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<mods:affiliation id="B7E29757531C1A8D748BB9F751AA2167">Institute for Biological Problems of the North, Far Eastern Branch of the Russian Academy of Sciences, Magadan 685000, Russia. & Altai State University, Barnaul, RF- 656049, Russia.</mods:affiliation>
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<mods:typeOfResource id="5101D404291B9C7F5E50BC0DC72B3903">text</mods:typeOfResource>
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<mods:title id="D88F22F1D6BE43FCF61CE54FB874FF63">Zootaxa</mods:title>
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<mods:part id="964A9E96BB6C011FB2E8075063B6297A">
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<mods:date id="4EBA50369846A7D802D17A440B6A8B3D">2024</mods:date>
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<mods:number id="0A6FAEBE43F305C8EEAAAF7A766E4052">2024-11-05</mods:number>
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<mods:number id="C849A5102CBF5466F8EFF38E8546D5D9">5536</mods:number>
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<mods:url id="2B5E8DF42509F24346A0998ECCABECA6">http://dx.doi.org/10.11646/zootaxa.5536.3.8</mods:url>
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<mods:classification id="3EDCDADF6F3A790A9F89E8C89C9AFD53">journal article</mods:classification>
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<mods:identifier id="F1104BF11C2022198D69C502031F0A04" type="DOI">10.11646/zootaxa.5536.3.8</mods:identifier>
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<mods:identifier id="7B6CA03AF9515717FC436C9E675E7A82" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B51125FF80E30DFCF6FF037DF9D6AF" ID-DOI="http://doi.org/10.5281/zenodo.14248045" ID-Zenodo-Dep="14248045" LSID="urn:lsid:plazi:treatment:03B51125FF80E30DFCF6FF037DF9D6AF" httpUri="http://treatment.plazi.org/id/03B51125FF80E30DFCF6FF037DF9D6AF" lastPageId="5" lastPageNumber="488" pageId="3" pageNumber="486">
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<subSubSection id="C306F3B8FF80E30BFCF6FF037BEBD6E7" box="[151,704,151,178]" pageId="3" pageNumber="486" type="nomenclature">
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<paragraph id="8BA3A033FF80E30BFCF6FF037BEBD6E7" blockId="3.[151,704,151,213]" box="[151,704,151,178]" pageId="3" pageNumber="486">
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<heading id="D0EB175FFF80E30BFCF6FF037BEBD6E7" bold="true" box="[151,704,151,178]" fontSize="11" level="1" pageId="3" pageNumber="486" reason="1">
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<emphasis id="B9687C21FF80E30BFCF6FF037BEBD6E7" bold="true" box="[151,704,151,178]" pageId="3" pageNumber="486">
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<taxonomicName id="4C1CDBB0FF80E30BFCF6FF0378E8D6E7" authorityName="Mikhaljova & Kazarin & Marusik" authorityYear="2024" box="[151,451,151,178]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="securifer" status="sp. nov.">
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<emphasis id="B9687C21FF80E30BFCF6FF0378E8D6E7" bold="true" box="[151,451,151,178]" italics="true" pageId="3" pageNumber="486">Skleroprotopus securifer</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25BC15AFF80E30BFDABFF037B0CD6E7" box="[458,551,151,177]" pageId="3" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
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Mikhaljova
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306F3B8FF80E30BFCF6FF2F782CD683" box="[151,263,187,213]" pageId="3" pageNumber="486" type="description">
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<paragraph id="8BA3A033FF80E30BFCF6FF2F782CD683" blockId="3.[151,704,151,213]" box="[151,263,187,213]" pageId="3" pageNumber="486">
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<figureCitation id="1327BCB6FF80E30BFCF6FF2F782CD683" box="[151,263,187,213]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–15" captionStart-2="FIGURES 16–19" captionStart-3="FIGURES 20–22" captionStartId-0="2.[151,264,1914,1938]" captionStartId-1="4.[151,264,1663,1687]" captionStartId-2="5.[151,264,1511,1535]" captionStartId-3="6.[151,264,1883,1907]" captionTargetBox-0="[152,1435,309,1891]" captionTargetBox-1="[151,1435,345,1640]" captionTargetBox-2="[151,1435,273,1488]" captionTargetBox-3="[151,1435,1094,1859]" captionTargetId-0="figure-87@2.[152,1435,309,1891]" captionTargetId-1="figure-99@4.[151,1435,345,1640]" captionTargetId-2="figure-64@5.[151,1435,273,1488]" captionTargetId-3="figure-255@6.[151,1435,1094,1859]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." captionText-1="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." captionText-2="FIGURES 16–19. Skleroprotopus securifer sp. nov., male holotype (FSCB). 16. Anterior gonopod, lateral view. 17. Anterior gonopod, mesal view. 18. Second and third telopoditemeres of anterior gonopod, lateral view. 19. Distal part of flagellum. Abbreviations: b, poleaxe-like outgrowth; e, third telopoditomere; f, flagellum; s, basal setae; t, short setae. Scales: 20 μm (Figs 18, 19), 100 μm (Figs 16, 17)." captionText-3="FIGURES 20–22. Skleroprotopus securifer sp. nov., male holotype (FSCB). 20. Posterior gonopod, mesal view, slightly turned back. 21. Posterior gonopod, plus tip of flagellum, lateral view. 22. Distal part of posterior gonopod, lateral view. Abbreviations: a, anterior branch; f, flagellum apex; p, posterior part; r, telopodite remnant. Scales: 20 μm (Fig. 22), 100 μm (Fig. 20), in mm (Fig. 21)." figureDoi-0="http://doi.org/10.5281/zenodo.14239136" figureDoi-1="http://doi.org/10.5281/zenodo.14239138" figureDoi-2="http://doi.org/10.5281/zenodo.14239140" figureDoi-3="http://doi.org/10.5281/zenodo.14239142" httpUri-0="https://zenodo.org/record/14239136/files/figure.png" httpUri-1="https://zenodo.org/record/14239138/files/figure.png" httpUri-2="https://zenodo.org/record/14239140/files/figure.png" httpUri-3="https://zenodo.org/record/14239142/files/figure.png" pageId="3" pageNumber="486">Figs 1–22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306F3B8FF80E30BFCF6FE977DD4D717" pageId="3" pageNumber="486" type="materials_examined">
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<paragraph id="8BA3A033FF80E30BFCF6FE977DD4D717" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
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<emphasis id="B9687C21FF80E30BFCF6FE9778A3D74B" bold="true" box="[151,392,259,285]" pageId="3" pageNumber="486">Material examined.</emphasis>
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<materialsCitation id="3B74AA6EFF80E30BFDF6FE9778D9D717" collectingDate="2005-09-19" collectionCode="FSCB" collectorName="Y. M. Marusik" country="China" location="Xiaowutai Mt." pageId="3" pageNumber="486" specimenCount="1" specimenCount-male="1" stateProvince="Hebei" typeStatus="holotype">
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<typeStatus id="54A71E91FF80E30BFDF6FE977B2FD74B" box="[407,516,259,285]" pageId="3" pageNumber="486" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D1A6BBAFF80E30BFE77FE977B42D74B" box="[534,617,259,285]" count="1" pageId="3" pageNumber="486" type="male">1 male</specimenCount>
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(
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<collectionCode id="ED0D38F6FF80E30BFEE1FE977BE1D74B" box="[640,714,259,285]" pageId="3" pageNumber="486">FSCB</collectionCode>
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),
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<collectingCountry id="F30BE0A3FF80E30BFE84FE977A00D74B" box="[741,811,259,285]" name="China" pageId="3" pageNumber="486">China</collectingCountry>
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,
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<collectingRegion id="49D86ED1FF80E30BFF5FFE977AE6D748" box="[830,973,259,286]" country="China" name="Hebei" pageId="3" pageNumber="486">Hebei Prov.</collectingRegion>
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,
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<location id="8EC3F6E8FF80E30BFFBEFE977DA6D748" LSID="urn:lsid:plazi:treatment:03B51125FF80E30DFCF6FF037DF9D6AF:8EC3F6E8FF80E30BFFBEFE977DA6D748" box="[991,1165,259,286]" country="China" name="Xiaowutai Mt." pageId="3" pageNumber="486" stateProvince="Hebei">Xiaowutai Mt.</location>
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, 40.025˚N, 115.303˚E,
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<date id="FFA286F3FF80E30BFCF6FEB37833D714" box="[151,280,295,322]" pageId="3" pageNumber="486" value="2005-09-19">
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<collectingDate id="EFE67F1BFF80E30BFCF6FEB37833D714" box="[151,280,295,322]" pageId="3" pageNumber="486" value="2005-09-19">19.09.2005</collectingDate>
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</date>
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, leg.
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<collectorName id="26E9C5E5FF80E30BFD33FEBC78D9D717" box="[338,498,295,322]" pageId="3" pageNumber="486">Y.M. Marusik</collectorName>
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</materialsCitation>
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;
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<materialsCitation id="3B74AA6EFF80E30BFD9CFEBC7DD0D717" box="[509,1275,295,322]" collectingDate="2005-09-19" collectionCode="FSCB" collectorName="Y. M. Marusik" country="China" location="Xiaowutai Mt." pageId="3" pageNumber="486" specimenCount="10" specimenCount-female="2" specimenCount-juvenile="8" stateProvince="Hebei" typeStatus="paratype">
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<typeStatus id="54A71E91FF80E30BFD9CFEBC7B45D714" box="[509,622,296,322]" pageId="3" pageNumber="486" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D1A6BBAFF80E30BFE18FEB37BC3D717" box="[633,744,295,321]" count="2" pageId="3" pageNumber="486" type="female">2 females</specimenCount>
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,
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<specimenCount id="9D1A6BBAFF80E30BFE92FEB37A46D717" box="[755,877,295,321]" count="8" pageId="3" pageNumber="486" type="juvenile">8 juveniles</specimenCount>
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(
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<collectionCode id="ED0D38F6FF80E30BFF1DFEB37AEDD717" box="[892,966,295,321]" pageId="3" pageNumber="486">FSCB</collectionCode>
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), same data as for holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306F3B8FF80E30BFCA6FEDF7C60D7DC" pageId="3" pageNumber="486" type="diagnosis">
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<paragraph id="8BA3A033FF80E30BFCA6FEDF7C60D7DC" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
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<emphasis id="B9687C21FF80E30BFCA6FEDF7869D733" bold="true" box="[199,322,331,357]" pageId="3" pageNumber="486">Diagnosis.</emphasis>
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Differs from congeners mainly by the flattened, poleaxe-like outgrowth in the middle of the caudal surface of the anterior gonopod coxa in combination with very short and stout male coxites of the legpair 7.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306F3B8FF80E30BFCA6FE077C23D0E8" pageId="3" pageNumber="486" type="description">
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<paragraph id="8BA3A033FF80E30BFCA6FE077DDBD44F" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
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<emphasis id="B9687C21FF80E30BFCA6FE077872D7FB" bold="true" box="[199,345,403,429]" pageId="3" pageNumber="486">Description.</emphasis>
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<emphasis id="B9687C21FF80E30BFD3FFE0778B4D7FB" box="[350,415,403,429]" italics="true" pageId="3" pageNumber="486">Male.</emphasis>
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Length in alcohol 44.0–45.0 mm, midbody vertical diameter 1.6–1.7 mm, with 60(–1) rings, excluding telson. Coloration (in alcohol) transversely striated from dark brown prozonites and beige metazonites. Anterior part of head light brown, vertex and occiput marbled dark brown. Eyes black, antennae brown. Collum with two transversely-oval, large, marbled light brown spots. Legs light brown. Telson light brown.
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</paragraph>
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<paragraph id="8BA3A033FF80E30BFCA6FDB77867D543" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
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Head smooth, epicranial setae absent, 4 supralabral setae; 22 labral setae. Eye patches subtriangle, composed of at least 50 ommatidia. Antennae medium-sized, rather slender and clavate. A small group of sensilla basiconica at distodorsal surface of each of antennomeres 5, 6 and 7, sensilla basiconica gradually decrease to tiny ones on antennomere 7 (
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<figureCitation id="1327BCB6FF80E30BFD12FD1B78F7D4FF" box="[371,476,655,681]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1914,1938]" captionTargetBox="[152,1435,309,1891]" captionTargetId="figure-87@2.[152,1435,309,1891]" captionTargetPageId="2" captionText="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." figureDoi="http://doi.org/10.5281/zenodo.14239136" httpUri="https://zenodo.org/record/14239136/files/figure.png" pageId="3" pageNumber="486">Figs 1, 2</figureCitation>
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). Mandibular stipites enlarged, ventrally incised, forming 2 nearly equal, subconical lobes. Gnathochilarium with lamellae linguales each of which with 6 setae arranged longitudinally. Promentum elongated longitudinally, expanded basally, swollen anteriorly. Collum laterally with distinct striae reaching to anterior margin.
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</paragraph>
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<paragraph id="8BA3A033FF80E30BFCA6FC8B7B9CD5BB" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">Body slightly compressed laterally, this becoming increasingly pronounced towards telson. Prozona with dorsal shallow striae located diagonally to the body axis and lateral very shallow transverse striae. Metazona with regular, longitudinal striae reaching hind margin and disappearing above the ozopore level; setae on hind margins absent. Ozopores small, set far behind pro-metazonital suture. Epiproct broadly rounded and slightly protruding caudad, marginally with two setae. Preanal ring not pubescent, but with two marginal setae ventrally. Anal valves and subanal scale each with two marginal setae only.</paragraph>
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<paragraph id="8BA3A033FF80E30BFCA6FC6C7D2ED367" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
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Legs long and slender, without modifications; claws long, weakly curved, at base with a short (in pregonopodal legs) (
|
||
<figureCitation id="1327BCB6FF80E30BFCBDFB8F7835D260" box="[220,286,1051,1078]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1914,1938]" captionTargetBox="[152,1435,309,1891]" captionTargetId="figure-87@2.[152,1435,309,1891]" captionTargetPageId="2" captionText="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." figureDoi="http://doi.org/10.5281/zenodo.14239136" httpUri="https://zenodo.org/record/14239136/files/figure.png" pageId="3" pageNumber="486">Fig. 5</figureCitation>
|
||
) and relatively short (in postgonopodal legs; about half the length of the claw) setiform accessory claw ventrally (
|
||
<figureCitation id="1327BCB6FF80E30BFD6DFBAB785CD20F" box="[268,375,1087,1113]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1914,1938]" captionTargetBox="[152,1435,309,1891]" captionTargetId="figure-87@2.[152,1435,309,1891]" captionTargetPageId="2" captionText="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." figureDoi="http://doi.org/10.5281/zenodo.14239136" httpUri="https://zenodo.org/record/14239136/files/figure.png" pageId="3" pageNumber="486">Figs 3, 7</figureCitation>
|
||
). Leg pair 1 enlarged, with 4-segmented telopodites, terminal segment with thin short setae distomesally (
|
||
<figureCitation id="1327BCB6FF80E30BFD52FBF778B3D22B" box="[307,408,1123,1149]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1914,1938]" captionTargetBox="[152,1435,309,1891]" captionTargetId="figure-87@2.[152,1435,309,1891]" captionTargetPageId="2" captionText="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." figureDoi="http://doi.org/10.5281/zenodo.14239136" httpUri="https://zenodo.org/record/14239136/files/figure.png" pageId="3" pageNumber="486">Figs 4, 6</figureCitation>
|
||
), 3rd telopoditomere longest, dorsoventrally strongly flattened, with a front surface covered with short setae; telopoditomeres 1 and 2 with longer setae frontally, coxa with two long setae apically and three similar setae frontally. Penis elongated, with 4 subapical setae on the front surface (
|
||
<figureCitation id="1327BCB6FF80E30BF851FB3F7D5CD290" box="[1072,1143,1195,1222]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1914,1938]" captionTargetBox="[152,1435,309,1891]" captionTargetId="figure-87@2.[152,1435,309,1891]" captionTargetPageId="2" captionText="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." figureDoi="http://doi.org/10.5281/zenodo.14239136" httpUri="https://zenodo.org/record/14239136/files/figure.png" pageId="3" pageNumber="486">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="1327BCB6FF80E30BF8E3FB3F7D97D293" box="[1154,1212,1195,1221]" captionStart="FIGURES 8–15" captionStartId="4.[151,264,1663,1687]" captionTargetBox="[151,1435,345,1640]" captionTargetId="figure-99@4.[151,1435,345,1640]" captionTargetPageId="4" captionText="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." figureDoi="http://doi.org/10.5281/zenodo.14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="3" pageNumber="486">9, 10</figureCitation>
|
||
). Coxae 2 each with one distal seta anteriorly and posteriorly (
|
||
<figureCitation id="1327BCB6FF80E30BFE0AFB5B7B9FD2BC" box="[619,692,1231,1258]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1914,1938]" captionTargetBox="[152,1435,309,1891]" captionTargetId="figure-87@2.[152,1435,309,1891]" captionTargetPageId="2" captionText="FIGURES 1–7. Skleroprotopus securifer sp. nov., male holotype (FSCB). 1. Distal part of antenna. 2. Antenna apex. 3. Midbody leg. 4. Leg 1, anterior view. 5. Leg 2 and penis, posterior view. 6. Terminal segment of leg 1, anterior view. 7. Claw of mid-body leg. Scales: 10 μm (Fig. 7), 20 μm (Figs 2, 6), 100 μm (Figs 1, 3, 4, 5)." figureDoi="http://doi.org/10.5281/zenodo.14239136" httpUri="https://zenodo.org/record/14239136/files/figure.png" pageId="3" pageNumber="486">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="1327BCB6FF80E30BFEA0FB5B7BCBD2BF" box="[705,736,1231,1257]" captionStart="FIGURES 8–15" captionStartId="4.[151,264,1663,1687]" captionTargetBox="[151,1435,345,1640]" captionTargetId="figure-99@4.[151,1435,345,1640]" captionTargetPageId="4" captionText="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." figureDoi="http://doi.org/10.5281/zenodo.14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="3" pageNumber="486">13</figureCitation>
|
||
). Leg pair 7 with coxal processes as short knobs bearing setae both mesally and apically (
|
||
<figureCitation id="1327BCB6FF80E30BFDADFB677B39D35B" box="[460,530,1267,1293]" captionStart="FIGURES 8–15" captionStartId="4.[151,264,1663,1687]" captionTargetBox="[151,1435,345,1640]" captionTargetId="figure-99@4.[151,1435,345,1640]" captionTargetPageId="4" captionText="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." figureDoi="http://doi.org/10.5281/zenodo.14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="3" pageNumber="486">Fig. 8</figureCitation>
|
||
). Telopodites 7 variable even within same male
|
||
<typeStatus id="54A71E91FF80E30BF85DFB677D89D35B" box="[1084,1186,1267,1293]" pageId="3" pageNumber="486" type="holotype">holotype</typeStatus>
|
||
: telopodite on the left 3-segmented (
|
||
<figureCitation id="1327BCB6FF80E30BFD54FA83788CD367" box="[309,423,1303,1329]" captionStart="FIGURES 8–15" captionStartId="4.[151,264,1663,1687]" captionTargetBox="[151,1435,345,1640]" captionTargetId="figure-99@4.[151,1435,345,1640]" captionTargetPageId="4" captionText="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." figureDoi="http://doi.org/10.5281/zenodo.14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="3" pageNumber="486">Figs 8, 11</figureCitation>
|
||
), but telopodite on the right 2-segmented (
|
||
<figureCitation id="1327BCB6FF80E30BFFE5FA837AD3D367" box="[900,1016,1303,1329]" captionStart="FIGURES 8–15" captionStartId="4.[151,264,1663,1687]" captionTargetBox="[151,1435,345,1640]" captionTargetId="figure-99@4.[151,1435,345,1640]" captionTargetPageId="4" captionText="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." figureDoi="http://doi.org/10.5281/zenodo.14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="3" pageNumber="486">Figs 8, 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA3A033FF80E30BFCA6FAAF7BBAD007" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
|
||
Gonopods protruding. Anterior gonopod with 3-segmented telopodite (telopoditomere 3 (
|
||
<emphasis id="B9687C21FF80E30BF8B8FAAF7DCED303" bold="true" box="[1241,1253,1339,1365]" pageId="3" pageNumber="486">e</emphasis>
|
||
,
|
||
<figureCitation id="1327BCB6FF80E30BF899FAAF7C66D303" box="[1272,1357,1339,1365]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1511,1535]" captionTargetBox="[151,1435,273,1488]" captionTargetId="figure-64@5.[151,1435,273,1488]" captionTargetPageId="5" captionText="FIGURES 16–19. Skleroprotopus securifer sp. nov., male holotype (FSCB). 16. Anterior gonopod, lateral view. 17. Anterior gonopod, mesal view. 18. Second and third telopoditemeres of anterior gonopod, lateral view. 19. Distal part of flagellum. Abbreviations: b, poleaxe-like outgrowth; e, third telopoditomere; f, flagellum; s, basal setae; t, short setae. Scales: 20 μm (Figs 18, 19), 100 μm (Figs 16, 17)." figureDoi="http://doi.org/10.5281/zenodo.14239140" httpUri="https://zenodo.org/record/14239140/files/figure.png" pageId="3" pageNumber="486">Fig. 18</figureCitation>
|
||
) tiny), bearing setae of different length apically and mesally on first telopoditomere, subapically on second telopoditomere and apically on third one (
|
||
<figureCitation id="1327BCB6FF80E30BFDD8FA177B48D3CB" box="[441,611,1411,1437]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1511,1535]" captionTargetBox="[151,1435,273,1488]" captionTargetId="figure-64@5.[151,1435,273,1488]" captionTargetPageId="5" captionText="FIGURES 16–19. Skleroprotopus securifer sp. nov., male holotype (FSCB). 16. Anterior gonopod, lateral view. 17. Anterior gonopod, mesal view. 18. Second and third telopoditemeres of anterior gonopod, lateral view. 19. Distal part of flagellum. Abbreviations: b, poleaxe-like outgrowth; e, third telopoditomere; f, flagellum; s, basal setae; t, short setae. Scales: 20 μm (Figs 18, 19), 100 μm (Figs 16, 17)." figureDoi="http://doi.org/10.5281/zenodo.14239140" httpUri="https://zenodo.org/record/14239140/files/figure.png" pageId="3" pageNumber="486">Figs 16, 17, 18</figureCitation>
|
||
). Coxal process of the anterior gonopods long, with large, broad, flattened poleaxe-like outgrowth (
|
||
<emphasis id="B9687C21FF80E30BFDCDFA337897D397" bold="true" box="[428,444,1447,1473]" pageId="3" pageNumber="486">b</emphasis>
|
||
), rounded apex with protrusion, 2 basal setae (
|
||
<emphasis id="B9687C21FF80E30BFFA8FA337AFFD397" bold="true" box="[969,980,1447,1473]" pageId="3" pageNumber="486">s</emphasis>
|
||
) mesally and a longitudinal row of short setae (
|
||
<emphasis id="B9687C21FF80E30BFC81FA5F79C2D3B3" bold="true" box="[224,233,1483,1509]" pageId="3" pageNumber="486">t</emphasis>
|
||
) anterolaterally (
|
||
<figureCitation id="1327BCB6FF80E30BFDCDFA5F7B05D3B3" box="[428,558,1483,1509]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1511,1535]" captionTargetBox="[151,1435,273,1488]" captionTargetId="figure-64@5.[151,1435,273,1488]" captionTargetPageId="5" captionText="FIGURES 16–19. Skleroprotopus securifer sp. nov., male holotype (FSCB). 16. Anterior gonopod, lateral view. 17. Anterior gonopod, mesal view. 18. Second and third telopoditemeres of anterior gonopod, lateral view. 19. Distal part of flagellum. Abbreviations: b, poleaxe-like outgrowth; e, third telopoditomere; f, flagellum; s, basal setae; t, short setae. Scales: 20 μm (Figs 18, 19), 100 μm (Figs 16, 17)." figureDoi="http://doi.org/10.5281/zenodo.14239140" httpUri="https://zenodo.org/record/14239140/files/figure.png" pageId="3" pageNumber="486">Figs 16, 17</figureCitation>
|
||
,
|
||
<figureCitation id="1327BCB6FF80E30BFE5AFA5F7B71D3B3" box="[571,602,1483,1509]" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1883,1907]" captionTargetBox="[151,1435,1094,1859]" captionTargetId="figure-255@6.[151,1435,1094,1859]" captionTargetPageId="6" captionText="FIGURES 20–22. Skleroprotopus securifer sp. nov., male holotype (FSCB). 20. Posterior gonopod, mesal view, slightly turned back. 21. Posterior gonopod, plus tip of flagellum, lateral view. 22. Distal part of posterior gonopod, lateral view. Abbreviations: a, anterior branch; f, flagellum apex; p, posterior part; r, telopodite remnant. Scales: 20 μm (Fig. 22), 100 μm (Fig. 20), in mm (Fig. 21)." figureDoi="http://doi.org/10.5281/zenodo.14239142" httpUri="https://zenodo.org/record/14239142/files/figure.png" pageId="3" pageNumber="486">21</figureCitation>
|
||
). Parabasal relatively long flagellum (
|
||
<emphasis id="B9687C21FF80E30BF868FA5F7D39D3B3" bold="true" box="[1033,1042,1483,1509]" pageId="3" pageNumber="486">f</emphasis>
|
||
) with thin spines distally and nonbifurcated apex (
|
||
<figureCitation id="1327BCB6FF80E30BFD32FA7B78D6D05F" box="[339,509,1519,1545]" captionStart="FIGURES 16–19" captionStartId="5.[151,264,1511,1535]" captionTargetBox="[151,1435,273,1488]" captionTargetId="figure-64@5.[151,1435,273,1488]" captionTargetPageId="5" captionText="FIGURES 16–19. Skleroprotopus securifer sp. nov., male holotype (FSCB). 16. Anterior gonopod, lateral view. 17. Anterior gonopod, mesal view. 18. Second and third telopoditemeres of anterior gonopod, lateral view. 19. Distal part of flagellum. Abbreviations: b, poleaxe-like outgrowth; e, third telopoditomere; f, flagellum; s, basal setae; t, short setae. Scales: 20 μm (Figs 18, 19), 100 μm (Figs 16, 17)." figureDoi="http://doi.org/10.5281/zenodo.14239140" httpUri="https://zenodo.org/record/14239140/files/figure.png" pageId="3" pageNumber="486">Figs 16, 17, 19</figureCitation>
|
||
,
|
||
<figureCitation id="1327BCB6FF80E30BFE68FA7B7B03D05F" box="[521,552,1519,1545]" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1883,1907]" captionTargetBox="[151,1435,1094,1859]" captionTargetId="figure-255@6.[151,1435,1094,1859]" captionTargetPageId="6" captionText="FIGURES 20–22. Skleroprotopus securifer sp. nov., male holotype (FSCB). 20. Posterior gonopod, mesal view, slightly turned back. 21. Posterior gonopod, plus tip of flagellum, lateral view. 22. Distal part of posterior gonopod, lateral view. Abbreviations: a, anterior branch; f, flagellum apex; p, posterior part; r, telopodite remnant. Scales: 20 μm (Fig. 22), 100 μm (Fig. 20), in mm (Fig. 21)." figureDoi="http://doi.org/10.5281/zenodo.14239142" httpUri="https://zenodo.org/record/14239142/files/figure.png" pageId="3" pageNumber="486">21</figureCitation>
|
||
). Posterior gonopod with long, slender anterior branch (
|
||
<emphasis id="B9687C21FF80E30BF8FAFA7B7D82D05F" bold="true" box="[1179,1193,1519,1545]" pageId="3" pageNumber="486">a</emphasis>
|
||
) pointed apically and posterior part (posterior branch) (
|
||
<emphasis id="B9687C21FF80E30BFE6FF9877B35D07B" bold="true" box="[526,542,1555,1581]" pageId="3" pageNumber="486">p</emphasis>
|
||
) bearing long setae in its inner surface and external edge, 1-segmented remnant of telopodite (
|
||
<emphasis id="B9687C21FF80E30BFD57F9A37869D007" bold="true" box="[310,322,1591,1617]" pageId="3" pageNumber="486">r</emphasis>
|
||
) at the base (
|
||
<figureCitation id="1327BCB6FF80E30BFDB9F9A37BAFD007" box="[472,644,1591,1617]" captionStart="FIGURES 20–22" captionStartId="6.[151,264,1883,1907]" captionTargetBox="[151,1435,1094,1859]" captionTargetId="figure-255@6.[151,1435,1094,1859]" captionTargetPageId="6" captionText="FIGURES 20–22. Skleroprotopus securifer sp. nov., male holotype (FSCB). 20. Posterior gonopod, mesal view, slightly turned back. 21. Posterior gonopod, plus tip of flagellum, lateral view. 22. Distal part of posterior gonopod, lateral view. Abbreviations: a, anterior branch; f, flagellum apex; p, posterior part; r, telopodite remnant. Scales: 20 μm (Fig. 22), 100 μm (Fig. 20), in mm (Fig. 21)." figureDoi="http://doi.org/10.5281/zenodo.14239142" httpUri="https://zenodo.org/record/14239142/files/figure.png" pageId="3" pageNumber="486">Figs 20, 21, 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA3A033FF80E30BFCA6F9CF7C23D0E8" blockId="3.[151,1437,259,2014]" pageId="3" pageNumber="486">
|
||
<emphasis id="B9687C21FF80E30BFCA6F9CF7830D023" box="[199,283,1627,1653]" italics="true" pageId="3" pageNumber="486">Female</emphasis>
|
||
. Length in alcohol 53.0–59.0 mm, midbody vertical diameter 2.4–2.5 mm, with 73(–1), 66(–1) rings, excluding telson. Ventral edge of the mandibular stipites with small knob medially. Leg pair 1 somewhat thicker and shorter, but leg pair 2 only somewhat shorter as compared the following legs. Vulva as in
|
||
<figureCitation id="1327BCB6FF80E30BF8E5F9377C2FD0E8" box="[1156,1284,1699,1726]" captionStart="FIGURES 8–15" captionStartId="4.[151,264,1663,1687]" captionTargetBox="[151,1435,345,1640]" captionTargetId="figure-99@4.[151,1435,345,1640]" captionTargetPageId="4" captionText="FIGURES 8–15. Skleroprotopus securifer sp. nov., male holotype and female paratype (FSCB). 8. Legpair 7, posterior view. 9. Penis apex, anterior view. 10. Penis, posterior view. 11. Telopodite of leg 7 on the left, posterior view. 12. Telopodite of leg 7 on the right, posterior view. 13. Distal part of coxa 2, anterior view. 14. Right vulva, lateral view. 15. Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15." figureDoi="http://doi.org/10.5281/zenodo.14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="3" pageNumber="486">Figs 14, 15</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306F3B8FF80E30DFCA6F9537AE2D683" lastPageId="5" lastPageNumber="488" pageId="3" pageNumber="486" type="discussion">
|
||
<paragraph id="8BA3A033FF80E30CFCA6F9537C4CD683" blockId="3.[151,1437,259,2014]" lastBlockId="4.[151,1437,151,321]" lastPageId="4" lastPageNumber="487" pageId="3" pageNumber="486">
|
||
<emphasis id="B9687C21FF80E30BFCA6F9537810D0B7" bold="true" box="[199,315,1735,1761]" pageId="3" pageNumber="486">Remarks.</emphasis>
|
||
In some
|
||
<taxonomicName id="4C1CDBB0FF80E30BFDC9F9537B80D0B7" box="[424,683,1735,1761]" pageId="3" pageNumber="486">
|
||
<emphasis id="B9687C21FF80E30BFDC9F9537B7AD0B7" box="[424,593,1735,1761]" italics="true" pageId="3" pageNumber="486">Skleroprotopus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="4C1CDBB0FF80E30BFE8FF9537A59D0B7" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[750,882,1735,1761]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="coreanus">
|
||
<emphasis id="B9687C21FF80E30BFE8FF9537A59D0B7" box="[750,882,1735,1761]" italics="true" pageId="3" pageNumber="486">S. coreanus</emphasis>
|
||
</taxonomicName>
|
||
from the south of the Russian Far East), a large number of heterogeneous forms of males with various combinations of the main distinguishing characters were noted; these forms are young individuals belonging to different postembryonic stages of development (
|
||
<bibRefCitation id="EF8DDDC2FF80E30BF978F89B79FFD11B" author="Mikhaljova, E. V." pageId="3" pageNumber="486" pagination="501 - 529" refId="ref4571" refString="Mikhaljova, E. V. (2019) Identities of the millipede genera Skleroprotopus Attems, 1901 and Ansiulus Takakuwa, 1940 (Diplopoda: Julida: Mongoliulidae), with emphasis on the postembryonic development of Skleroprotopus coreanus (Pocock, 1895). Zootaxa, 4551 (5), 501 - 529. https: // doi. org / 10.11646 / zootaxa. 4551.5.1" type="journal article" year="2019">Mikhaljova 2019</bibRefCitation>
|
||
). The male specimen of
|
||
<taxonomicName id="4C1CDBB0FF80E30BFD84F8A77BD2D11B" authorityName="Mikhaljova & Kazarin & Marusik" authorityYear="2024" box="[485,761,1843,1869]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="securifer" status="sp. nov.">
|
||
<emphasis id="B9687C21FF80E30BFD84F8A77BD2D11B" box="[485,761,1843,1869]" italics="true" pageId="3" pageNumber="486">Skleroprotopus securifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9687C21FF80E30BFF61F8A77A72D11B" bold="true" box="[768,857,1843,1869]" pageId="3" pageNumber="486">
|
||
<taxonomicNameLabel id="A25BC15AFF80E30BFF61F8A77A72D11B" box="[768,857,1843,1869]" pageId="3" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
described above is a fully-formed and mature adult male. The criteria for this interpretation are as follows: penis is conical bearing several setae apically; leg pair 1 is large, thick, with 4-segmented telopodites; leg pair 7 is modified, with 2–3-segmented telopodites and a short knobby setose coxal processes; anterior gonopods have 3-segmented setose telopodites (third lateral telopoditomere is tiny), flat coxal processes with large, broad, flattened poleaxe-like outgrowth, relatively long flagella, bearing small spinules apically; posterior gonopods with long, slender pointed anterior branches and posterior parts bearing long setae in them inner surfaces and external edges; posterior gonopods with remnants of telopodites at base.
|
||
</paragraph>
|
||
<paragraph id="8BA3A033FF87E30CFCA6FF4B79DBD717" blockId="4.[151,1437,151,321]" pageId="4" pageNumber="487">
|
||
As for the variation in the number of segments of the telopodite 7, similar variations were also noted in other species (
|
||
<bibRefCitation id="EF8DDDC2FF87E30CFC98FE977B3BD74B" author="Murakami, Y. & Paik, K. Y." box="[249,528,259,285]" pageId="4" pageNumber="487" pagination="363 - 384" refId="ref4859" refString="Murakami, Y. & Paik, K. Y. (1968) Results of the speleological survey in South Korea 1966. XI. Cave-dwelling myriapods from the southern part of Korea. Bulletin of the National Science Museum, 11 (4), 363 - 384." type="journal article" year="1968">Murakami & Paik 1968</bibRefCitation>
|
||
; Mikhaljova & Korsόs 2003), thus, this feature is unreliable for distinguishing species.
|
||
</paragraph>
|
||
<caption id="DF63F0BBFF87E30CFCF6F9EB7DE5D155" ID-DOI="http://doi.org/10.5281/zenodo.14239138" ID-Zenodo-Dep="14239138" httpUri="https://zenodo.org/record/14239138/files/figure.png" pageId="4" pageNumber="487" startId="4.[151,264,1663,1687]" targetBox="[151,1435,345,1640]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BA3A033FF87E30CFCF6F9EB7DE5D155" blockId="4.[151,1437,1663,1796]" pageId="4" pageNumber="487">
|
||
<emphasis id="B9687C21FF87E30CFCF6F9EB7863D0C1" bold="true" box="[151,328,1663,1687]" pageId="4" pageNumber="487">FIGURES 8–15.</emphasis>
|
||
<taxonomicName id="4C1CDBB0FF87E30CFD2EF9EB7B62D0C1" authorityName="Mikhaljova & Kazarin & Marusik" authorityYear="2024" box="[335,585,1663,1687]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="securifer" status="sp. nov.">
|
||
<emphasis id="B9687C21FF87E30CFD2EF9EB7B62D0C1" box="[335,585,1663,1687]" italics="true" pageId="4" pageNumber="487">Skleroprotopus securifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9687C21FF87E30CFE31F9EB7B8AD0C1" bold="true" box="[592,673,1663,1687]" pageId="4" pageNumber="487">
|
||
<taxonomicNameLabel id="A25BC15AFF87E30CFE31F9EB7B8AD0C1" box="[592,673,1663,1687]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype and female paratype (
|
||
<emphasis id="B9687C21FF87E30CF844F9EB7D4DD0C1" bold="true" box="[1061,1126,1663,1687]" pageId="4" pageNumber="487">FSCB</emphasis>
|
||
).
|
||
<emphasis id="B9687C21FF87E30CF81AF9EB7DA5D0C1" bold="true" box="[1147,1166,1663,1687]" pageId="4" pageNumber="487">8.</emphasis>
|
||
Legpair 7, posterior view.
|
||
<emphasis id="B9687C21FF87E30CFCF6F9377981D0ED" bold="true" box="[151,170,1699,1723]" pageId="4" pageNumber="487">9.</emphasis>
|
||
Penis apex, anterior view.
|
||
<emphasis id="B9687C21FF87E30CFDDAF93778F1D0ED" bold="true" box="[443,474,1699,1723]" pageId="4" pageNumber="487">10.</emphasis>
|
||
Penis, posterior view.
|
||
<emphasis id="B9687C21FF87E30CFEA1F9377BF5D0ED" bold="true" box="[704,734,1699,1723]" pageId="4" pageNumber="487">11.</emphasis>
|
||
Telopodite of leg 7 on the left, posterior view.
|
||
<emphasis id="B9687C21FF87E30CF8D9F9377DF3D0ED" bold="true" box="[1208,1240,1699,1723]" pageId="4" pageNumber="487">12.</emphasis>
|
||
Telopodite of leg 7 on the right, posterior view.
|
||
<emphasis id="B9687C21FF87E30CFDD6F95378FCD089" bold="true" box="[439,471,1735,1759]" pageId="4" pageNumber="487">13.</emphasis>
|
||
Distal part of coxa 2, anterior view.
|
||
<emphasis id="B9687C21FF87E30CFF2FF9537A45D089" bold="true" box="[846,878,1735,1759]" pageId="4" pageNumber="487">14.</emphasis>
|
||
Right vulva, lateral view.
|
||
<emphasis id="B9687C21FF87E30CF81AF9537DB0D089" bold="true" box="[1147,1179,1735,1759]" pageId="4" pageNumber="487">15.</emphasis>
|
||
Right vulva apex, lateral view. Scales: 10 μm (Figs 9, 11, 12), 20 μm (Fig. 10), 100 μm (Figs 8, 13); without scales: Figs 14 and 15.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA3A033FF87E30DFCA6F8DF7AE2D683" blockId="4.[151,1437,1866,2037]" lastBlockId="5.[151,1436,151,249]" lastPageId="5" lastPageNumber="488" pageId="4" pageNumber="487">
|
||
<taxonomicName id="4C1CDBB0FF87E30CFCA6F8DF78F5D133" authorityName="Mikhaljova & Kazarin & Marusik" authorityYear="2024" box="[199,478,1867,1893]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="securifer" status="sp. nov.">
|
||
<emphasis id="B9687C21FF87E30CFCA6F8DF78F5D133" box="[199,478,1867,1893]" italics="true" pageId="4" pageNumber="487">Skleroprotopus securifer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9687C21FF87E30CFD89F8DF7B6ED133" bold="true" box="[488,581,1867,1893]" pageId="4" pageNumber="487">
|
||
<taxonomicNameLabel id="A25BC15AFF87E30CFD89F8DF7B6ED133" box="[488,581,1867,1893]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C1CDBB0FF87E30CFE82F8DF7DDFD133" authority="Zhang, 1985" authorityName="Zhang" authorityYear="1985" box="[739,1268,1866,1893]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="membranipedalis">
|
||
<emphasis id="B9687C21FF87E30CFE82F8DF7D73D133" box="[739,1112,1867,1893]" italics="true" pageId="4" pageNumber="487">Skleroprotopus membranipedalis</emphasis>
|
||
<bibRefCitation id="EF8DDDC2FF87E30CF802F8DF7DDFD133" author="Zhang, C. Z." box="[1123,1268,1866,1893]" pageId="4" pageNumber="487" pagination="154 - 156" refId="ref5421" refString="Zhang, C. Z. (1985) A new species of millipeds of the genus Skleroprotopus in Stone Buddha Cave, Fangshang County, Beijing. In: Karst geomorphology and speleology. Science Press, Beijing, pp. 154 - 156. [In Chinese]" type="book chapter" year="1985">
|
||
Zhang,
|
||
<quantity id="4CE40DD6FF87E30CF8DDF8DE7DDFD133" box="[1212,1268,1866,1893]" metricMagnitude="1" metricUnit="m" metricValue="5.0419" pageId="4" pageNumber="487" unit="in" value="1985.0">1985</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in the general configuration of gonopods, but differs from it mainly by the short, knob-like coxal processes of the leg-pair 7 (vs. elongated in
|
||
<taxonomicName id="4C1CDBB0FF87E30CFD4DF8077B27D1FB" authorityName="Zhang" authorityYear="1985" box="[300,524,1939,1965]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="membranipedalis">
|
||
<emphasis id="B9687C21FF87E30CFD4DF8077B27D1FB" box="[300,524,1939,1965]" italics="true" pageId="4" pageNumber="487">S. membranipedalis</emphasis>
|
||
</taxonomicName>
|
||
), by the middle location of the poleaxe-like outgrowth (
|
||
<emphasis id="B9687C21FF87E30CF8F7F8077D8DD1FB" bold="true" box="[1174,1190,1939,1965]" pageId="4" pageNumber="487">b</emphasis>
|
||
) in the middle of the caudal surface of the anterior gonopod coxa (vs. subbasal in
|
||
<taxonomicName id="4C1CDBB0FF87E30CFF33F8237D19D187" authorityName="Zhang" authorityYear="1985" box="[850,1074,1975,2001]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="membranipedalis">
|
||
<emphasis id="B9687C21FF87E30CFF33F8237D19D187" box="[850,1074,1975,2001]" italics="true" pageId="4" pageNumber="487">S. membranipedalis</emphasis>
|
||
</taxonomicName>
|
||
), by the penis with 4 subapical setae on the front surface (vs. 2 apical setae laterally in
|
||
<taxonomicName id="4C1CDBB0FF87E30CFF68F84F7ACCD1A3" authorityName="Zhang" authorityYear="1985" box="[777,999,2011,2037]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="membranipedalis">
|
||
<emphasis id="B9687C21FF87E30CFF68F84F7ACCD1A3" box="[777,999,2011,2037]" italics="true" pageId="4" pageNumber="487">S. membranipedalis</emphasis>
|
||
</taxonomicName>
|
||
), by the darker coloration: dark brown prozonites and beige metazonites (vs. light brown-beige coloration, head whitish in
|
||
<taxonomicName id="4C1CDBB0FF86E30DF85DFF037C31D6E7" authorityName="Zhang" authorityYear="1985" box="[1084,1306,151,177]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="membranipedalis">
|
||
<emphasis id="B9687C21FF86E30DF85DFF037C31D6E7" box="[1084,1306,151,177]" italics="true" pageId="5" pageNumber="488">S. membranipedalis</emphasis>
|
||
</taxonomicName>
|
||
), as well as by the terrestrial habitat (vs.
|
||
<taxonomicName id="4C1CDBB0FF86E30DFDB8FF2F7B9CD683" authorityName="Zhang" authorityYear="1985" box="[473,695,187,213]" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="membranipedalis">
|
||
<emphasis id="B9687C21FF86E30DFDB8FF2F7B9CD683" box="[473,695,187,213]" italics="true" pageId="5" pageNumber="488">S. membranipedalis</emphasis>
|
||
</taxonomicName>
|
||
was collected in caves).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306F3B8FF86E30DFCA6FF4B7DF9D6AF" box="[199,1234,223,249]" pageId="5" pageNumber="488" type="etymology">
|
||
<paragraph id="8BA3A033FF86E30DFCA6FF4B7DF9D6AF" blockId="5.[151,1436,151,249]" box="[199,1234,223,249]" pageId="5" pageNumber="488">
|
||
<emphasis id="B9687C21FF86E30DFCA6FF4B7865D6AF" bold="true" box="[199,334,223,249]" pageId="5" pageNumber="488">Etymology.</emphasis>
|
||
The specific epithet refers to the axe-like posterior process of anterior gonopod.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |