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<document id="436CE45A2C145F7F13D4475A9B7048B7" ID-CLB-Dataset="46524" ID-DOI="10.11646/zootaxa.3760.3.9" ID-GBIF-Dataset="cf705912-1286-478a-8f89-df37c4093205" ID-ISSN="1175-5326" ID-Zenodo-Dep="231006" ID-ZooBank="12759523-69AB-4374-A78A-B3CDC4428B2A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460379042190" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A. &amp; Makarchenko, Marina A." docDate="2014" docId="03D2713219312E37FF12405DFA35FB2B" docLanguage="en" docName="zt03760p438.pdf" docOrigin="Zootaxa 3760 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrobaenus Fries" docType="key" docVersion="10" lastPageNumber="431" masterDocId="FFEB094A19302E35FF85465EFFA9FFB9" masterDocTitle="On taxonomy of Hydrobaenus Fries, 1830 (Diptera: Chironomidae: Orthocladiinae) from the Russian Far East, with a key to species" masterLastPageNumber="438" masterPageNumber="429" pageNumber="430" updateTime="1698296590671" updateUser="ExternalLinkService">
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<mods:title id="04D9DE4C8544BE379305F26781D4443C">On taxonomy of Hydrobaenus Fries, 1830 (Diptera: Chironomidae: Orthocladiinae) from the Russian Far East, with a key to species</mods:title>
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<mods:namePart id="F02981937B71C3B9226768900E163A4E">Makarchenko, Eugenyi A.</mods:namePart>
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<mods:date id="4D73D8CF176407735C7C2D2D4988FAC5">2014</mods:date>
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<mods:number id="514DAA22E97AD054392FFD21416A4E2F">3760</mods:number>
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<treatment id="03D2713219312E37FF12405DFA35FB2B" ID-DOI="http://doi.org/10.5281/zenodo.5695735" ID-GBIF-Taxon="119413772" ID-Zenodo-Dep="5695735" LSID="urn:lsid:plazi:treatment:03D2713219312E37FF12405DFA35FB2B" httpUri="http://treatment.plazi.org/id/03D2713219312E37FF12405DFA35FB2B" lastPageId="2" lastPageNumber="431" pageId="1" pageNumber="430">
<subSubSection id="C36193AF19312E34FF12405DFF76F9DB" pageId="1" pageNumber="430" type="nomenclature">
<paragraph id="8BC4C02419312E34FF12405DFC01F9A4" blockId="1.[151,936,1538,1565]" box="[151,936,1538,1565]" pageId="1" pageNumber="430">
<heading id="D08C774819312E34FF12405DFC01F9A4" bold="true" box="[151,936,1538,1565]" fontSize="11" level="1" pageId="1" pageNumber="430" reason="1">
<emphasis id="B90F1C3619312E34FF12405DFC01F9A4" bold="true" box="[151,936,1538,1565]" pageId="1" pageNumber="430">
Key to species of
<taxonomicName id="4C7BBBA719312E34FEE8405CFDFEF9A4" ID-CoL="4ZQK" ID-ENA="611358" authority="Fries" authorityName="Fries" box="[365,599,1538,1565]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="B90F1C3619312E34FEE8405CFDB9F9A5" bold="true" box="[365,528,1538,1564]" italics="true" pageId="1" pageNumber="430">Hydrobaenus</emphasis>
Fries
</taxonomicName>
from the Russian Far East
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC4C02419312E34FF124017FF76F9DB" blockId="1.[151,223,1609,1634]" box="[151,223,1609,1634]" pageId="1" pageNumber="430">
<heading id="D08C774819312E34FF124017FF76F9DB" bold="true" box="[151,223,1609,1634]" fontSize="10" level="2" pageId="1" pageNumber="430" reason="2">
<emphasis id="B90F1C3619312E34FF124017FF76F9DB" bold="true" box="[151,223,1609,1634]" pageId="1" pageNumber="430">Males</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36193AF19312E37FF1240CFFA35FB2B" lastPageId="2" lastPageNumber="431" pageId="1" pageNumber="430" type="key">
<keyStep id="308FD88119312E34FF1240CFFA32F97A" pageId="1" pageNumber="430">
<paragraph id="8BC4C02419312E34FF1240CFFA32F91F" blockId="1.[151,1437,1681,2029]" box="[151,1435,1681,1702]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF1240CFFA32F91F" box="[151,1435,1681,1702]" pageId="1" pageNumber="430">1. Anal point reduced and usually in the form of rounded or triangular tubercle or projection............................ 2</keyLead>
</paragraph>
<paragraph id="8BC4C02419312E34FF1240F0FA32F97A" blockId="1.[151,1437,1681,2029]" box="[151,1435,1710,1731]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF1240F0FA32F97A" box="[151,1435,1710,1731]" pageId="1" pageNumber="430">- Anal point well developed.............................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119312E34FF124092FA32F947" pageId="1" pageNumber="430">
<paragraph id="8BC4C02419312E34FF124092FA32F958" blockId="1.[151,1437,1681,2029]" box="[151,1435,1740,1761]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF124092FA32F958" box="[151,1435,1740,1761]" pageId="1" pageNumber="430">2. Gonostylus wide, distally angularly, on the outer edge with small peakedness projection............................ 3</keyLead>
</paragraph>
<paragraph id="8BC4C02419312E34FF1240B7FA32F947" blockId="1.[151,1437,1681,2029]" box="[151,1435,1769,1790]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF1240B7FA32F947" box="[151,1435,1769,1790]" pageId="1" pageNumber="430">- Gonostylus different shapes, without projection on the outer edge............................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119312E34FF124158FA35F8C1" pageId="1" pageNumber="430">
<paragraph id="8BC4C02419312E34FF124158FA35F882" blockId="1.[151,1437,1681,2029]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF124158FA35F882" pageId="1" pageNumber="430">
3. AR 2.50; LR1 0.73–0.74. Dorsocentrals 15–17. Transverse sternapodeme wide in middle part. Tergite IX narrowly triangular. Virga consists of 4 setae. Gonostylus nearest of megaseta with some setae of middle length.....
<taxonomicName id="4C7BBBA719312E34FB124178FA35F882" ID-CoL="3N59Z" authority="Saether, 1976" authorityName="Saether" authorityYear="1976" box="[1175,1436,1829,1851]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="laticaudus">
<emphasis id="B90F1C3619312E34FB124178FAB1F882" box="[1175,1304,1829,1851]" italics="true" pageId="1" pageNumber="430">H. laticaudus</emphasis>
<bibRefCitation id="EFEABDD519312E34FA9B4178FA35F882" author="Saether" box="[1310,1436,1830,1851]" pageId="1" pageNumber="430" refString="Saether, O. A. (1976) Revision of Hydrobaenus, Trissocladius, Zalutschia, Paratrissocladius, and some related genera (Diptera, Chironomidae). Bulletin of the Fisheries Research Board of Canada, 195, 1 - 287." type="journal article" year="1976">Saether, 1976</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419312E34FF12411DFA35F8C1" blockId="1.[151,1437,1681,2029]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF12411DFA35F8C1" pageId="1" pageNumber="430">
- AR 1.27; LR1 0.61.Dorsocentrals 4–5. Transverse sternapodeme narrow in middle part. Tergite IX rounded.Virga consists of 2 setae. Gonostylus nearest of megaseta with 2 strong and long setae........
<taxonomicName id="4C7BBBA719312E34FC50413DFA35F8C1" ID-CoL="BDLCY" authority="Makarchenko et Makarchenko, 2010" authorityName="Makarchenko et Makarchenko" authorityYear="2010" box="[981,1436,1891,1912]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="tiunovi">
<emphasis id="B90F1C3619312E34FC50413DFB9CF8C1" box="[981,1077,1891,1912]" italics="true" pageId="1" pageNumber="430">H. tiunovi</emphasis>
<bibRefCitation id="EFEABDD519312E34FBBE413DFA35F8C1" author="Makarchenko" box="[1083,1436,1891,1912]" pageId="1" pageNumber="430" refString="Makarchenko, E. A. &amp; Makarchenko, M. A. (2010) Hydrobaenus tiunovi sp. n. (Diptera, Chironomidae, Orthocladiinae) from Sokhondinsky State Biosphere Reserve (Zabaikal'sky Territory, Russia). Euroasian Entomological Journal, 9 (3), 411 - 412." type="journal article" year="2010">
Makarchenko
<emphasis id="B90F1C3619312E34FB41413DFB7CF8C1" box="[1220,1237,1891,1912]" italics="true" pageId="1" pageNumber="430">et</emphasis>
Makarchenko, 2010
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119312E34FF1241DEFA3EF869" pageId="1" pageNumber="430">
<paragraph id="8BC4C02419312E34FF1241DEFA35F80A" blockId="1.[151,1437,1681,2029]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF1241DEFA35F80A" pageId="1" pageNumber="430">
4. Gonostylus with tooth distally; inferior volsella double, with large rounded-triangular dorsal and ventral parts which well sep- arated from each other......................................................
<taxonomicName id="4C7BBBA719312E34FBBD41C0FA35F80A" ID-CoL="9HJBB" authority="Sasa et Kawai, 1987" authorityName="Sasa et Kawai" authorityYear="1987" box="[1080,1436,1949,1971]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="biwaquartus">
<emphasis id="B90F1C3619312E34FBBD41C0FB78F80A" box="[1080,1233,1949,1971]" italics="true" pageId="1" pageNumber="430">H. biwaquartus</emphasis>
Sasa
<emphasis id="B90F1C3619312E34FA8F41C0FAB2F80A" box="[1290,1307,1950,1971]" italics="true" pageId="1" pageNumber="430">et</emphasis>
Kawai, 1987
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419312E34FF1241E5FA3EF869" blockId="1.[151,1437,1681,2029]" box="[151,1431,1979,2000]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF1241E5FA3EF869" box="[151,1431,1979,2000]" pageId="1" pageNumber="430">- Gonostylus without tooth distally; inferior volsella different shapes and if consists of two parts, they not clearly separated ... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119312E37FF124186FA35FF5E" lastPageId="2" lastPageNumber="431" pageId="1" pageNumber="430">
<paragraph id="8BC4C02419312E34FF124186FA34F854" blockId="1.[151,1437,1681,2029]" box="[151,1437,2008,2029]" pageId="1" pageNumber="430">
<keyLead id="308A631119312E34FF124186FA34F854" box="[151,1437,2008,2029]" pageId="1" pageNumber="430">5. Gonostylus straight, with large triangular or rounded triangular crista dorsalis in distal part. Virga consists of 4 setae. Acros-</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF6446C9FA35FF5E" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
tichals distally curved and beak-shaped.......................................
<taxonomicName id="4C7BBBA719322E37FBB646C6FA35FF15" ID-CoL="9HWCC" authority="Sasa et Kondo, 1991" authorityName="Sasa et Kondo" authorityYear="1991" box="[1075,1436,151,173]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="kisosecundus">
<emphasis id="B90F1C3619322E37FBB646C6FB67FF14" box="[1075,1230,151,173]" italics="true" pageId="2" pageNumber="431">H. kisosecundus</emphasis>
Sasa
<emphasis id="B90F1C3619322E37FA8246C6FAB1FF14" box="[1287,1304,152,173]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Kondo, 1991
</taxonomicName>
<keyLead id="308A631119322E37FF1246EBFA35FF5E" pageId="2" pageNumber="431">
- Gonostylus slightly curved, distally without crista dorsalis. Virga consists of 2 setae. Acrostichals simple..........................................................................................
<taxonomicName id="4C7BBBA719322E37FBDD468CFA35FF5E" ID-CoL="3N59P" authority="Goetghebuer, 1933" authorityName="Goetghebuer" authorityYear="1933" box="[1112,1436,209,231]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="fusistylus">
<emphasis id="B90F1C3619322E37FBDD468CFB66FF5E" box="[1112,1231,209,231]" italics="true" pageId="2" pageNumber="431">H. fusistylus</emphasis>
(Goetghebuer, 1933)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF1246B1FA32FE9B" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF1246B1FA32FEBD" blockId="2.[151,1437,151,1170]" box="[151,1435,239,260]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1246B1FA32FEBD" box="[151,1435,239,260]" pageId="2" pageNumber="431">6. Gonostylus in apical part on the outer edge with a triangular or rounded-triangular projection......................... 7</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF124753FA32FE9B" blockId="2.[151,1437,151,1170]" box="[151,1435,269,290]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124753FA32FE9B" box="[151,1435,269,290]" pageId="2" pageNumber="431">- Gonostylus in apical part without projection................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF124774FA35FE2E" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF124774FA35FEE5" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124774FA35FEE5" pageId="2" pageNumber="431">
7. Apical projection of gonostylus at angle pointing down. Virga consists of 4 setae. Transverse sternapodema with high rod-like oral projections..............................................................
<taxonomicName id="4C7BBBA719322E37FBE94716FA35FEE5" ID-CoL="3N59J" authority="Holmgren, 1869" authorityName="Holmgren" authorityYear="1869" box="[1132,1436,327,349]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="conformis">
<emphasis id="B90F1C3619322E37FBE94716FB40FEE4" box="[1132,1257,327,349]" italics="true" pageId="2" pageNumber="431">H. conformis</emphasis>
(Holmgren, 1869)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF12473BFA35FE2E" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF12473BFA35FE2E" pageId="2" pageNumber="431">
- Apical projection of gonostylus usually directed forward. Virga absent. Transverse sternapodema with low-triangular oral pro- jections................................................
<taxonomicName id="4C7BBBA719322E37FCF647DCFA35FE2E" authority="Makarchenko et Makarchenko, 2009" authorityName="Makarchenko et Makarchenko" authorityYear="2009" box="[883,1436,385,407]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="pseudoconformis">
<emphasis id="B90F1C3619322E37FCF647DCFB9DFE2E" box="[883,1076,385,407]" italics="true" pageId="2" pageNumber="431">H. pseudoconformis</emphasis>
<bibRefCitation id="EFEABDD519322E37FBBE47DCFA35FE2E" author="Makarchenko" box="[1083,1436,386,407]" pageId="2" pageNumber="431" refString="Makarchenko, E. A., Makarchenko, M. A. &amp; Yavorskaya, N. M. (2009) New findings of chironomids (Diptera, Chironomidae, Orthocladiinae) in Far East and bordering territories. VI. Hydrobaenus Fries. Euroasian Entomological Journal, 8, Supplement 1, 33 - 50. [in Russian]" type="journal article" year="2009">
Makarchenko
<emphasis id="B90F1C3619322E37FB4147DCFB7CFE2E" box="[1220,1237,386,407]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2009
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF1247C1FA32FE6B" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF1247C1FA32FE0D" blockId="2.[151,1437,151,1170]" box="[151,1435,415,436]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1247C1FA32FE0D" box="[151,1435,415,436]" pageId="2" pageNumber="431">8. Gonostylus distally with crista dorsalis.................................................................... 9</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF1247E3FA32FE6B" blockId="2.[151,1437,151,1170]" box="[151,1435,445,466]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1247E3FA32FE6B" box="[151,1435,445,466]" pageId="2" pageNumber="431">- Gonostylus without crista dorsalis....................................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF124784FA32FDFE" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF124784FA32FD93" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124784FA32FD93" pageId="2" pageNumber="431">
9. Crista dorsalis triangular or rounded-triangular, located in apical inner edge and not always clearly visible behind megaseta. Tergite IX with a relatively long (about 33 µm) and narrow anal point. Acrostichals 0–5.................................................................................
<taxonomicName id="4C7BBBA719322E37FCCD444BFA32FD93" authority="Makarchenko, Makarchenko et Yavorskaja, 2009" authorityName="Makarchenko, Makarchenko et Yavorskaja" authorityYear="2009" box="[840,1435,533,554]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="sigaensis">
<emphasis id="B90F1C3619322E37FCCD444BFC14FD93" box="[840,957,533,554]" italics="true" pageId="2" pageNumber="431">H. sigaensis</emphasis>
Makarchenko, Makarchenko
<emphasis id="B90F1C3619322E37FB5B444BFB46FD93" box="[1246,1263,533,554]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Yavorskaja, 2009
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF12446CFA32FDFE" blockId="2.[151,1437,151,1170]" box="[151,1435,562,583]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF12446CFA32FDFE" box="[151,1435,562,583]" pageId="2" pageNumber="431">- Crista dorsalis located on preapical inner edge. Acrostichals 22–24............................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF124411FA32FD26" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF124411FA35FD3B" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124411FA35FD3B" pageId="2" pageNumber="431">
10. Virga absent. Anal point parallel-sided or subparallel-sided, 28–40 µm long (
<figureCitation id="1340DCA119322E37FC464411FB9CFDDD" box="[963,1077,591,612]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,264,1731,1754]" captionTargetBox="[194,1398,195,1707]" captionTargetId="figure@4.[188,1398,195,1707]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Hydrobaenus distinctus (Makarchenko et Makarchenko), male. 1 – 3, hypopygium in dorsal view; 4, anteropronotum and anterior part of scutum from one side; 5, anterior part of antepronotum in dorsal view; 6, 9, gonostylus; 7, wing; 8, transverse sternapodeme, and phallapodemes. Scale bars are as follows: Figs. 1 – 6, 8, 9 — 50 µm; Fig. 8 — 200 µm." httpUri="https://zenodo.org/record/231007/files/figure.png" pageId="2" pageNumber="431">Figs. 1–2, 3</figureCitation>
)................................................................................
<taxonomicName id="4C7BBBA719322E37FCC84433FA96FD3B" ID-CoL="9HJBC" authority="Makarchenko et Makarchenko, 2006" authorityName="Makarchenko et Makarchenko" authorityYear="2006" box="[845,1343,620,642]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="distinctus" status="comb. nov.">
<emphasis id="B90F1C3619322E37FCC84433FC6FFD3B" box="[845,966,620,642]" italics="true" pageId="2" pageNumber="431">H. distinctus</emphasis>
(Makarchenko
<emphasis id="B90F1C3619322E37FBDA4433FBD8FD3B" box="[1119,1137,621,642]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2006)
</taxonomicName>
<taxonomicNameLabel id="A23CA14D19322E37FAC14432FA35FD3B" box="[1348,1436,620,642]" pageId="2" pageNumber="431">
<emphasis id="B90F1C3619322E37FAC14432FA35FD3B" bold="true" box="[1348,1436,620,642]" pageId="2" pageNumber="431">comb. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF1244D4FA32FD26" blockId="2.[151,1437,151,1170]" box="[151,1435,650,671]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1244D4FA32FD26" box="[151,1435,650,671]" pageId="2" pageNumber="431">- Virga present. Anal point triangular, 16–24 µm long........................................................ 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF1244F9FA3FFCAD" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF1244F9FA35FD63" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1244F9FA35FD63" pageId="2" pageNumber="431">
11. Gonostylus in apical quarter naked, crista dorsalis low and long, rounded or rounded-triangular. Dorsal part of inferior volsella rounded-triangular, ventral part long and flat...................
<taxonomicName id="4C7BBBA719322E37FCFB449BFA35FD63" ID-CoL="6MKDY" authority="Makarchenko et Makarchenko, 2006" authorityName="Makarchenko et Makarchenko" authorityYear="2006" box="[894,1436,708,730]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="sikhotealinensis">
<emphasis id="B90F1C3619322E37FCFB449BFB9CFD63" box="[894,1077,708,730]" italics="true" pageId="2" pageNumber="431">H. sikhotealinensis</emphasis>
Makarchenko
<emphasis id="B90F1C3619322E37FB41449BFB7CFD63" box="[1220,1237,709,730]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2006
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF1244BCFA3FFCAD" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1244BCFA3FFCAD" pageId="2" pageNumber="431">
- Gonostylus completely covered with microtrichiae, crista dorsalis short and rounded-triangular. Dorsal part of inferior volsella with finger-like projection, ventral part short and rounded..........
<taxonomicName id="4C7BBBA719322E37FC0E455EFA3FFCAD" ID-CoL="6MKD5" authority="Makarchenko et Makarchenko, 2005" authorityName="Makarchenko et Makarchenko" authorityYear="2005" box="[907,1430,767,789]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="monodentatus">
<emphasis id="B90F1C3619322E37FC0E455EFB86FCAC" box="[907,1071,767,789]" italics="true" pageId="2" pageNumber="431">H. monodentatus</emphasis>
Makarchenko
<emphasis id="B90F1C3619322E37FB3A455EFB79FCAC" box="[1215,1232,768,789]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2005
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF124543FA35FCD5" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF124543FA35FCF6" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124543FA35FCF6" pageId="2" pageNumber="431">
12. Anal point wide and low, with a slightly concave apex. Inferior volsella large and rounded................................................................................... ..
<taxonomicName id="4C7BBBA719322E37FC0C4564FA35FCF6" ID-CoL="6MKD2" authority="Makarchenko et Makarchenko, 2005" authorityName="Makarchenko et Makarchenko" authorityYear="2005" box="[905,1436,825,847]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="septentrionalis">
<emphasis id="B90F1C3619322E37FC0C4564FB9DFCF6" box="[905,1076,825,847]" italics="true" pageId="2" pageNumber="431">H. septentrionalis</emphasis>
Makarchenko
<emphasis id="B90F1C3619322E37FB414564FB7CFCF6" box="[1220,1237,826,847]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2005
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF124509FA35FCD5" blockId="2.[151,1437,151,1170]" box="[151,1436,855,876]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124509FA35FCD5" box="[151,1436,855,876]" pageId="2" pageNumber="431">- Anal point narrow and triangular. Inferior volsella different structure.......................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF12452BFDE1FC7D" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF12452BFA3FFC1E" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF12452BFA3FFC1E" pageId="2" pageNumber="431">
13. Inferior volsella weakly developed and as two low tuberculate projections..................................................................................................
<taxonomicName id="4C7BBBA719322E37FC0645CCFA3FFC1E" authority="Makarchenko et Makarchenko, 2009" authorityName="Makarchenko et Makarchenko" authorityYear="2009" box="[899,1430,913,935]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="parvacaudatus">
<emphasis id="B90F1C3619322E37FC0645CCFB86FC1E" box="[899,1071,913,935]" italics="true" pageId="2" pageNumber="431">H. parvacaudatus</emphasis>
<bibRefCitation id="EFEABDD519322E37FBB045CCFA3FFC1E" author="Makarchenko" box="[1077,1430,914,935]" pageId="2" pageNumber="431" refString="Makarchenko, E. A., Makarchenko, M. A. &amp; Yavorskaya, N. M. (2009) New findings of chironomids (Diptera, Chironomidae, Orthocladiinae) in Far East and bordering territories. VI. Hydrobaenus Fries. Euroasian Entomological Journal, 8, Supplement 1, 33 - 50. [in Russian]" type="journal article" year="2009">
Makarchenko
<emphasis id="B90F1C3619322E37FB3A45CCFB79FC1E" box="[1215,1232,914,935]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2009
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF1245F1FDE1FC7D" blockId="2.[151,1437,151,1170]" box="[151,584,943,964]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF1245F1FDE1FC7D" box="[151,584,943,964]" pageId="2" pageNumber="431">- Inferior volsella well developed ... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF124593FA35FBA5" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF124593FA32FC46" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124593FA32FC46" pageId="2" pageNumber="431">
14. Dorsal part of inferior volsella with finger-like or rounded apical projection (
<figureCitation id="1340DCA119322E37FC474593FB9EFC5B" box="[962,1079,973,994]" captionStart="FIGURES 30 – 33" captionStartId="8.[151,264,1826,1849]" captionTargetBox="[198,1393,747,1805]" captionTargetId="figure@8.[188,1398,738,1805]" captionTargetPageId="8" captionText="FIGURES 30 – 33. Hydrobaenus jacuticus Makarchenko et Makarchenko, male. 30 — hypopygium in dorsal view (South Yakutia); 31 – 32 — inferior volsellae (Amur River basin); 33 — part of hypopyium in dorsal view (Kolyma River basin). Scale bars: Figs. 30 – 31, 33 — 50 µm; Fig. 32 — 20 µm." httpUri="https://zenodo.org/record/231009/files/figure.png" pageId="2" pageNumber="431">Figs. 30–33</figureCitation>
). AR 0.9–1.2................................................................................
<taxonomicName id="4C7BBBA719322E37FC4445B4FA32FC46" ID-CoL="BD3PZ" authority="Makarchenko et Makarchenko, 2011" authorityName="Makarchenko et Makarchenko" authorityYear="2011" box="[961,1435,1002,1023]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="jacuticus">
<emphasis id="B90F1C3619322E37FC4445B4FB9FFC46" box="[961,1078,1002,1023]" italics="true" pageId="2" pageNumber="431">H. jacuticus</emphasis>
<bibRefCitation id="EFEABDD519322E37FBB945B4FA32FC46" author="Makarchenko" box="[1084,1435,1002,1023]" pageId="2" pageNumber="431" refString="Makarchenko, E. A. &amp; Makarchenko, M. A. (2011) New or little-known species of chironomids (Diptera, Chironomidae) from South Yakutia and bordering territory. Euroasian Entomological Journal, 10 (3), 389 - 396. [in Russian]" type="journal article" year="2011">
Makarchenko
<emphasis id="B90F1C3619322E37FB4045B4FB7EFC46" box="[1221,1239,1002,1023]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2011
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF124259FA35FBA5" blockId="2.[151,1437,151,1170]" box="[151,1436,1031,1052]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124259FA35FBA5" box="[151,1436,1031,1052]" pageId="2" pageNumber="431">- Dorsal part of inferior volsella triangular or rounded, without projection. AR&gt; 1.2................................ 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="308FD88119322E37FF12427BFA35FB2B" pageId="2" pageNumber="431">
<paragraph id="8BC4C02419322E37FF12427BFA35FBEE" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF12427BFA35FBEE" pageId="2" pageNumber="431">
15. AR 1.47–1.60. Anal point short (16–20 µm), with subapical pore. Dorsal part of inferior volsella rounded...............................................................................
<taxonomicName id="4C7BBBA719322E37FC5D421CFA35FBEE" authority="Makarchenko et Makarchenko, 2005" authorityName="Makarchenko et Makarchenko" authorityYear="2005" box="[984,1436,1090,1111]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="siricus">
<emphasis id="B90F1C3619322E37FC5D421CFB9CFBEE" box="[984,1077,1090,1111]" italics="true" pageId="2" pageNumber="431">H. siricus</emphasis>
Makarchenko
<emphasis id="B90F1C3619322E37FB41421CFB7CFBEE" box="[1220,1237,1090,1111]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2005
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC4C02419322E37FF124201FA35FB2B" blockId="2.[151,1437,151,1170]" pageId="2" pageNumber="431">
<keyLead id="308A631119322E37FF124201FA35FB2B" pageId="2" pageNumber="431">
- AR 2.0. Anal point more long (28 µm), without subapical pore. Dorsal part of inferior volsella triangular..........................................................................
<taxonomicName id="4C7BBBA719322E37FC0E4223FA35FB2B" authority="Makarchenko et Makarchenko, 2009" authorityName="Makarchenko et Makarchenko" authorityYear="2009" box="[907,1436,1148,1170]" class="Insecta" family="Chironomidae" genus="Hydrobaenus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="maladistinctus">
<emphasis id="B90F1C3619322E37FC0E4223FB9CFB2B" box="[907,1077,1148,1170]" italics="true" pageId="2" pageNumber="431">H. maladistinctus</emphasis>
<bibRefCitation id="EFEABDD519322E37FBBE4223FA35FB2B" author="Makarchenko" box="[1083,1436,1149,1170]" pageId="2" pageNumber="431" refString="Makarchenko, E. A., Makarchenko, M. A. &amp; Yavorskaya, N. M. (2009) New findings of chironomids (Diptera, Chironomidae, Orthocladiinae) in Far East and bordering territories. VI. Hydrobaenus Fries. Euroasian Entomological Journal, 8, Supplement 1, 33 - 50. [in Russian]" type="journal article" year="2009">
Makarchenko
<emphasis id="B90F1C3619322E37FB414223FB7CFB2B" box="[1220,1237,1149,1170]" italics="true" pageId="2" pageNumber="431">et</emphasis>
Makarchenko, 2009
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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