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<document id="2869AA74E67FA90E548FDD444FD54FA3" ID="10.11646/zootaxa.4226.4.1" ID-CLB-Dataset="37251" ID-DOI="10.11646/zootaxa.4226.4.1" ID-GBIF-Dataset="8903776b-ace3-4ff6-aa28-635145d78260" ID-ISSN="1175-5326" ID-Zenodo-Dep="265060" ID-ZooBank="64B19A58-BBB5-4858-833F-F7937C3A351F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1485846904537" checkinUser="plazi" docAuthor="Min, Bum Sik, Seo, Ji Eun, Grischenko, Andrei V. &amp; Gordon, Dennis P." docDate="2017" docId="03C287EA562E4701FF26E5E3FDF7C983" docLanguage="en" docName="zootaxa.4226.4.1.pdf" docOrigin="Zootaxa 4226 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Suhius Min, Seo, Grischenko &amp; Gordon, 2017, n. gen." docType="treatment" docVersion="8" lastPageNumber="464" masterDocId="FFFBFF925623470FFFB1E025FFCCC82E" masterDocTitle="Intertidal Bryozoa from Korea — new additions to the fauna and a new genus of Bitectiporidae (Cheilostomata) from Baengnyeong Island, Yellow Sea" masterLastPageNumber="470" masterPageNumber="451" pageNumber="464" updateTime="1698436510809" updateUser="ExternalLinkService">
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<mods:title id="06D065D1FE3183189D82FC5CA35C7AE0">Intertidal Bryozoa from Korea — new additions to the fauna and a new genus of Bitectiporidae (Cheilostomata) from Baengnyeong Island, Yellow Sea</mods:title>
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<mods:namePart id="F159DBCFC961EDC036EBC7159B7DDDAC">Min, Bum Sik</mods:namePart>
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<mods:namePart id="6F22B6C059BAFB985DCF63C87B2309DE">Seo, Ji Eun</mods:namePart>
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<mods:namePart id="B919FB9AD043BB9E6C2CCAEC5113D78E">Grischenko, Andrei V.</mods:namePart>
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<mods:namePart id="6CA673B2D009EDAAA6E69D39E1572861">Gordon, Dennis P.</mods:namePart>
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<mods:date id="8DF5A6547A7D1FC4BBA5F272465BD719">2017</mods:date>
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<treatment id="03C287EA562E4701FF26E5E3FDF7C983" ID-DOI="http://doi.org/10.5281/zenodo.5220982" ID-GBIF-Taxon="127627956" ID-Zenodo-Dep="5220982" LSID="urn:lsid:plazi:treatment:03C287EA562E4701FF26E5E3FDF7C983" httpUri="http://treatment.plazi.org/id/03C287EA562E4701FF26E5E3FDF7C983" lastPageId="14" lastPageNumber="464" pageId="13" pageNumber="464">
<subSubSection id="C3716577562E4702FF26E5E3FE89CDCE" box="[151,325,1478,1504]" pageId="13" pageNumber="464" type="nomenclature">
<paragraph id="8BD436FC562E4702FF26E5E3FE89CDCE" blockId="13.[151,325,1478,1504]" box="[151,325,1478,1504]" pageId="13" pageNumber="464">
<heading id="D09C8190562E4702FF26E5E3FE89CDCE" bold="true" box="[151,325,1478,1504]" fontSize="11" level="1" pageId="13" pageNumber="464" reason="1">
<emphasis id="B91FEAEE562E4702FF26E5E3FE89CDCE" bold="true" box="[151,325,1478,1504]" pageId="13" pageNumber="464">
<taxonomicName id="4C6B4D7F562E4702FF26E5E3FF20CDCE" ID-CoL="84TY9" box="[151,236,1478,1504]" class="Gymnolaemata" family="Bitectiporidae" genus="Suhius" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="464" phylum="Bryozoa" rank="genus" status="gen. nov.">
<emphasis id="B91FEAEE562E4702FF26E5E3FF20CDCE" bold="true" box="[151,236,1478,1504]" italics="true" pageId="13" pageNumber="464">Suhius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C5795562E4702FF42E5E3FE89CDCE" box="[243,325,1478,1504]" pageId="13" pageNumber="464" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3716577562E4702FF26E628FC18CE0D" box="[151,980,1549,1571]" pageId="13" pageNumber="464" type="materials_examined">
<paragraph id="8BD436FC562E4702FF26E628FC18CE0D" blockId="13.[151,980,1549,1571]" box="[151,980,1549,1571]" pageId="13" pageNumber="464">
<emphasis id="B91FEAEE562E4702FF26E628FEEECE0D" bold="true" box="[151,290,1549,1571]" pageId="13" pageNumber="464">Type species.</emphasis>
<taxonomicName id="4C6B4D7F562E4702FE98E628FCF2CE0D" authority="Liu in Liu, Yin &amp; Ma, 2001" authorityName="Liu in Liu, Yin &amp; Ma" authorityYear="2001" box="[297,830,1549,1571]" class="Gymnolaemata" family="Bitectiporidae" genus="Hippothyris" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="464" phylum="Bryozoa" rank="species" species="rubescentis">
<emphasis id="B91FEAEE562E4702FE98E628FDD0CE0D" box="[297,540,1549,1571]" italics="true" pageId="13" pageNumber="464">Hippothyris rubescentis</emphasis>
Liu
<emphasis id="B91FEAEE562E4702FDFDE62BFDACCE0D" box="[588,608,1550,1571]" italics="true" pageId="13" pageNumber="464">in</emphasis>
<bibRefCitation id="EFFA4B0D562E4702FDD7E628FCF2CE0D" author="Liu" box="[614,830,1549,1571]" pageId="13" pageNumber="464" refString="Liu, X., Yin, X. &amp; Ma, J. (2001) Biology of Marine-Fouling Bryozoans in the Coastal Waters of China. Science Press, Beijing, 860 pp." type="book" year="2001">Liu, Yin &amp; Ma, 2001</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C3716577562E4702FF26E66AFE65CEA3" pageId="13" pageNumber="464" type="etymology">
<paragraph id="8BD436FC562E4702FF26E66AFE65CEA3" blockId="13.[151,1437,1615,2037]" pageId="13" pageNumber="464">
<emphasis id="B91FEAEE562E4702FF26E66AFED2CE46" bold="true" box="[151,286,1615,1640]" pageId="13" pageNumber="464">Etymology.</emphasis>
Honorific for Dr Chang-Hoon Suh, chairman of Woosuk University,
<collectingCountry id="F37C766C562E4702FBE4E675FB50CE49" box="[1109,1180,1616,1639]" name="South Korea" pageId="13" pageNumber="464">Korea</collectingCountry>
, in recognition of his service to the university.
</paragraph>
</subSubSection>
<subSubSection id="C3716577562E4702FF76E6B2FC90CFA6" pageId="13" pageNumber="464" type="diagnosis">
<paragraph id="8BD436FC562E4702FF76E6B2FC90CFA6" blockId="13.[151,1437,1615,2037]" pageId="13" pageNumber="464">
<emphasis id="B91FEAEE562E4702FF76E6B2FE8DCE9E" bold="true" box="[199,321,1687,1712]" pageId="13" pageNumber="464">Diagnosis.</emphasis>
Colony encrusting, unilaminar. Autozooids quincuncial with distinct margins. Orifice hoof-shaped, more or less parallel-sided, with broad, concave poster and stout condyles. No oral spines. Frontal shield nonpseudoporous, hummocky, with many marginal areolar pores. Ooecium acleithral, with fully calcified ectooecium and scattered ectooecial pseudopores; encroaching secondary calcification from neighbouring zooids forming low crests and nodules at fusion points, concealing pseudopores. Avicularia adventitious with complete crossbars, variable in size and disposition, typically lateral-oral in many younger zooids, larger and frontal in older zooids. Multiporous mural septula in lateral walls. Ancestrula tatiform.
</paragraph>
</subSubSection>
<subSubSection id="C3716577562E4701FF76E7B1FDF7C983" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464" type="discussion">
<paragraph id="8BD436FC562E4701FF76E7B1FDF7C983" blockId="13.[151,1437,1615,2037]" lastBlockId="14.[151,1436,151,429]" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464">
<emphasis id="B91FEAEE562E4702FF76E7B1FEF7CF83" bold="true" box="[199,315,1940,1965]" pageId="13" pageNumber="464">Remarks.</emphasis>
The combination of characters in
<taxonomicName id="4C6B4D7F562E4702FD0CE7B1FCC4CF82" box="[701,776,1940,1964]" class="Gymnolaemata" family="Bitectiporidae" genus="Suhius" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="464" phylum="Bryozoa" rank="genus" status="gen. nov.">
<emphasis id="B91FEAEE562E4702FD0CE7B1FCC4CF82" box="[701,776,1940,1964]" italics="true" pageId="13" pageNumber="464">Suhius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C5795562E4702FCBEE7B1FC93CF83" box="[783,863,1940,1965]" pageId="13" pageNumber="464" rank="genus">
<emphasis id="B91FEAEE562E4702FCBEE7B1FC93CF83" bold="true" box="[783,863,1940,1965]" pageId="13" pageNumber="464">n. gen.</emphasis>
</taxonomicNameLabel>
is distinctive. On the one hand, the orifice is hoofshaped as in
<taxonomicName id="4C6B4D7F562E4702FE9AE79DFD80CFFE" authority="Canu, 1917" authorityName="Canu" authorityYear="1917" box="[299,588,1976,2000]" class="Gymnolaemata" family="Hippoporidridae" genus="Hippoporella" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="464" phylum="Bryozoa" rank="genus">
<emphasis id="B91FEAEE562E4702FE9AE79DFE0DCFFE" box="[299,449,1976,2000]" italics="true" pageId="13" pageNumber="464">Hippoporella</emphasis>
<bibRefCitation id="EFFA4B0D562E4702FE7BE79DFD80CFFE" author="Canu" box="[458,588,1976,2000]" pageId="13" pageNumber="464" refString="Canu, F. (1917) Bryozoaires. Revue Critique de Paleozoologie, 21, 29 - 37." type="journal article" year="1917">Canu, 1917</bibRefCitation>
</taxonomicName>
and some other genera of superfamily Celleporoidea, which explains why
<bibRefCitation id="EFFA4B0D562E4702FF26E7F9FEA8CFDB" author="Androsova" box="[151,356,2012,2037]" pageId="13" pageNumber="464" refString="Androsova, E. I. (1959) Bryozoan fauna of the Yellow Sea. Trudy Sovestmoi Kitaisko-Sovietskoi Morskoi Biologicheskoi Expeditsii, 3, 41 - 70, pls 1 - 3. [in Russian]" type="journal article" year="1959">Androsova (1959)</bibRefCitation>
misattributed the
<typeStatus id="54D0885E562E4702FD83E7F8FDAECFDB" box="[562,610,2013,2037]" pageId="13" pageNumber="464">type</typeStatus>
species to
<taxonomicName id="4C6B4D7F562E4702FD51E7F9FB5FCFDB" authority="Smitt, 1868b" authorityName="Smitt" authorityYear="1868" box="[736,1171,2012,2037]" class="Gymnolaemata" family="Hippoporidridae" genus="Hippoporella" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="464" phylum="Bryozoa" rank="species" species="hippopus">
<emphasis id="B91FEAEE562E4702FD51E7F9FC28CFDA" box="[736,996,2012,2036]" italics="true" pageId="13" pageNumber="464">Hippoporella hippopus</emphasis>
(
<bibRefCitation id="EFFA4B0D562E4702FC45E7F9FB47CFDB" author="Smitt" box="[1012,1163,2012,2037]" pageId="13" pageNumber="464" refString="Smitt, F. A. (1868 b) Kritisk forteckning ofver Skandinaviens Hafs-Bryozoer. (IV.). Ofversigt af Kongliga Vetenskaps- Akademiens Forhandlingar, 25 (Supplement), 3 - 230, pls. 24 - 28." type="journal article" year="1868" yearSuffix="b">Smitt, 1868b</bibRefCitation>
)
</taxonomicName>
. On the other hand, the ooecium when first formed is typical of genera of superfamily Smittinoidea, with a smoothly calcified perforated ectooecium. Accordingly,
<bibRefCitation id="EFFA4B0D562D4701FE0FE099FDB8C8FA" box="[446,628,188,213]" pageId="14" pageNumber="465" refString="Liu, X., Yin, X. &amp; Ma, J. (2001) Biology of Marine-Fouling Bryozoans in the Coastal Waters of China. Science Press, Beijing, 860 pp." type="book">
Liu
<emphasis id="B91FEAEE562D4701FE5AE098FDD0C8FA" box="[491,540,188,212]" italics="true" pageId="14" pageNumber="465">et al</emphasis>
. (2001)
</bibRefCitation>
included the
<typeStatus id="54D0885E562D4701FCBAE098FCF7C8FB" box="[779,827,189,213]" pageId="14" pageNumber="465">type</typeStatus>
species in the
<taxonomicName id="4C6B4D7F562D4701FC6EE099FB4CC8FB" box="[991,1152,188,213]" class="Gymnolaemata" family="Bitectiporidae" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="465" phylum="Bryozoa" rank="family">Bitectiporidae</taxonomicName>
, a family of Smittinoidea in which the frontal shields and ooecia resemble those of many
<taxonomicName id="4C6B4D7F562D4701FC28E0FAFBD6C8D6" box="[921,1050,223,248]" class="Gymnolaemata" family="Smittinidae" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="465" phylum="Bryozoa" rank="family">Smittinidae</taxonomicName>
but the orifices are non-lyrulate (
<bibRefCitation id="EFFA4B0D562D4701FF2FE121FE8CC932" author="Gordon" box="[158,320,260,285]" pageId="14" pageNumber="465" refString="Gordon, D. P. (1984) The marine fauna of New Zealand: Bryozoa: Gymnolaemata from the Kermadec Ridge. New Zealand Oceanographic Institute Memoir, 91, 1 - 198." type="journal article" year="1984">Gordon 1984</bibRefCitation>
).
<bibRefCitation id="EFFA4B0D562D4701FEEDE121FDF1C932" box="[348,573,260,285]" pageId="14" pageNumber="465" refString="Liu, X., Yin, X. &amp; Ma, J. (2001) Biology of Marine-Fouling Bryozoans in the Coastal Waters of China. Science Press, Beijing, 860 pp." type="book">
Liu
<emphasis id="B91FEAEE562D4701FE23E120FE1EC932" box="[402,466,260,284]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
s (2001)
</bibRefCitation>
attribution of the species to
<taxonomicName id="4C6B4D7F562D4701FC1DE121FB2DC933" authority="Osburn, 1952" authorityName="Osburn" authorityYear="1952" box="[940,1249,260,285]" class="Gymnolaemata" family="Bitectiporidae" genus="Hippothyris" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="465" phylum="Bryozoa" rank="genus">
<emphasis id="B91FEAEE562D4701FC1DE121FBFEC932" box="[940,1074,260,284]" italics="true" pageId="14" pageNumber="465">Hippothyris</emphasis>
<bibRefCitation id="EFFA4B0D562D4701FBF1E121FB2DC933" author="Osburn" box="[1088,1249,260,285]" pageId="14" pageNumber="465" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America, part 2, Cheilostomata-Ascophora. Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn, 1952</bibRefCitation>
</taxonomicName>
recognized the similarities between the two genera, but the
<typeStatus id="54D0885E562D4701FD35E10DFD78C96E" box="[644,692,296,320]" pageId="14" pageNumber="465">type</typeStatus>
species of
<taxonomicName id="4C6B4D7F562D4701FC81E10DFC7AC96E" box="[816,950,296,320]" class="Gymnolaemata" family="Bitectiporidae" genus="Hippothyris" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="465" phylum="Bryozoa" rank="genus">
<emphasis id="B91FEAEE562D4701FC81E10DFC7AC96E" box="[816,950,296,320]" italics="true" pageId="14" pageNumber="465">Hippothyris</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6B4D7F562D4701FC72E10DFA92C96E" authority="Osburn, 1952" authorityName="Osburn" authorityYear="1952" box="[963,1374,295,320]" class="Gymnolaemata" family="Bitectiporidae" genus="Hippothyris" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="465" phylum="Bryozoa" rank="species" species="emplastra">
<emphasis id="B91FEAEE562D4701FC72E10DFB0CC96E" box="[963,1216,296,320]" italics="true" pageId="14" pageNumber="465">Hippothyris emplastra</emphasis>
<bibRefCitation id="EFFA4B0D562D4701FB77E102FA92C96E" author="Osburn" box="[1222,1374,295,320]" pageId="14" pageNumber="465" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America, part 2, Cheilostomata-Ascophora. Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn, 1952</bibRefCitation>
</taxonomicName>
from deep water off Baja
<collectingRegion id="49AFF81E562D4701FECFE169FE3FC94B" box="[382,499,332,357]" country="United States of America" name="California" pageId="14" pageNumber="465">California</collectingRegion>
, has only a small suboral avicularium, a discrete imperforate area of frontal shield surrounded by numerous pseudopores and an ectooecium with numerous pseudopores that never become wholly concealed by secondary calcification.
</paragraph>
</subSubSection>
</treatment>
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