treatments-xml/data/03/C2/87/03C287DEFFDE931BFF47FAA800EDFC15.xml

<document id="CC0D004C94AD9C597AF9C99128D16FF1" ID-DOI="10.11646/zootaxa.3669.4.5" ID-GBIF-Dataset="105e3c13-a88f-4aa7-a8b3-8a74ee8c244e" ID-ISSN="1175-5326" ID-Zenodo-Dep="249423" ID-ZooBank="DC8922B5-F317-46EC-92E0-7AED66A0C714" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460632703143" checkinUser="plazi" docAuthor="Aguila, Rayner Núñez, Matos- Maraví, Pável F. &amp; Wahlberg, Niklas" docDate="2013" docId="03C287DEFFDE931BFF47FAA800EDFC15" docLanguage="en" docName="zt03669p521.pdf" docOrigin="Zootaxa 3669 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calisto dissimulatum Nunez, sp. n." docType="treatment" docVersion="8" lastPageNumber="513" masterDocId="FFFBFFA6FFDD9311FFD0FF99022AFFFA" masterDocTitle="New Calisto species from Cuba, with insights on the relationships of Cuban and Bahamian taxa (Lepidoptera, Nymphalidae, Satyrinae)" masterLastPageNumber="521" masterPageNumber="503" pageNumber="506" updateTime="1698334641043" updateUser="plazi">
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<mods:title id="C99D949DCC53B171F2265B2D392D85BB">New Calisto species from Cuba, with insights on the relationships of Cuban and Bahamian taxa (Lepidoptera, Nymphalidae, Satyrinae)</mods:title>
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<mods:namePart id="4AF73D60BE63AF348EEDCC39642F18F8">Aguila, Rayner Núñez</mods:namePart>
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<mods:namePart id="9C5724C6652EA0C3AAEFD4E9C5AD948C">Matos- Maraví, Pável F.</mods:namePart>
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<treatment id="03C287DEFFDE931BFF47FAA800EDFC15" ID-DOI="http://doi.org/10.5281/zenodo.6161726" ID-GBIF-Taxon="119570019" ID-Zenodo-Dep="6161726" LSID="urn:lsid:plazi:treatment:03C287DEFFDE931BFF47FAA800EDFC15" httpUri="http://treatment.plazi.org/id/03C287DEFFDE931BFF47FAA800EDFC15" lastPageId="10" lastPageNumber="513" pageId="3" pageNumber="506">
<subSubSection id="C3716543FFDE9312FF47FAA80046FA96" pageId="3" pageNumber="506" type="nomenclature">
<paragraph id="8BD436C8FFDE9312FF47FAA80017FAB1" blockId="3.[151,620,1329,1388]" box="[151,573,1329,1355]" pageId="3" pageNumber="506">
<heading id="D09C81A4FFDE9312FF47FAA80017FAB1" bold="true" box="[151,573,1329,1355]" fontSize="11" level="1" pageId="3" pageNumber="506" reason="1">
<emphasis id="B91FEADAFFDE9312FF47FAA80017FAB1" bold="true" box="[151,573,1329,1355]" pageId="3" pageNumber="506">
<taxonomicName id="4C6B4D4BFFDE9312FF47FAA803C4FAB1" authority="Nunez" authorityName="Nunez" box="[151,494,1329,1355]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFDE9312FF47FAA803BCFAB1" bold="true" box="[151,406,1329,1355]" italics="true" pageId="3" pageNumber="506">Calisto dissimulatum</emphasis>
Núñez
</taxonomicName>
, 
<taxonomicNameLabel id="A22C57A1FFDE9312FE2BFAA80017FAB1" box="[507,573,1329,1355]" pageId="3" pageNumber="506" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD436C8FFDE9312FF47FACD0046FA96" blockId="3.[151,620,1329,1388]" box="[151,620,1364,1388]" pageId="3" pageNumber="506">
<figureCitation id="13502A4DFFDE9312FF47FACD0334FA96" box="[151,286,1364,1388]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,269,1848,1872]" captionTargetBox="[253,1332,211,1820]" captionTargetId="figure@5.[247,1337,194,1831]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Types of Calisto from Cuba. Calisto torrei sp. n.: 1 — 3 holotype upper side, 2 — 3 holotype under side, from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 10 / V / 2012, DNA voucher RN 01 – 01 (M 065), 3 — Ƥ paratype upper side, 4 — Ƥ paratype under side, from Cienfuegos, Pico San Juan, 1140 m, 12 / V / 2012, DNA voucher RN 01 – 03 (M 067). Calisto dissimulatum sp. n.: 5 — 3 holotype upper side, 6 — 3 holotype under side, from Holguín, La Zoilita Station, 400 m, 20 / II / 2010, DNA voucher PM 07 – 20 (M 037), 7 — Ƥ paratype upper side, 8 — Ƥ paratype under side, same data as holotype except 3 – 10 / IV / 2012, DNA voucher RN 02 – 01 (M 073)." httpUri="https://zenodo.org/record/249424/files/figure.png" pageId="3" pageNumber="506">Figures 5–8</figureCitation>
, 
<figureCitation id="13502A4DFFDE9312FEFAFACD0358FA96" box="[298,370,1364,1388]" captionStart="FIGURES 17 – 22" captionStartId="7.[151,269,1537,1561]" captionTargetBox="[226,1359,194,1515]" captionTargetId="figure@7.[226,1360,194,1516]" captionTargetPageId="7" captionText="FIGURES 17 – 22. Live adults of Cuban Calisto. Calisto torrei sp. n.: 17 — from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 18 — from Sancti Spiritus, Topes de Collante, La Represa, 800 m. Calisto dissimulatum sp. n.: 19, 20 — from Holguín, alrededores de La Zoilita, 400 m. Calisto aquilum sp. n.: 21 — from Cienfuegos, road to Pico San Juan, 22 — from Cienfuegos, Pico San Juan." httpUri="https://zenodo.org/record/249426/files/figure.png" pageId="3" pageNumber="506">19–20</figureCitation>
, 
<figureCitation id="13502A4DFFDE9312FEAEFACD03DBFA96" box="[382,497,1364,1388]" captionStart="FIGURES 23 – 32" captionStartId="8.[151,269,1842,1866]" captionTargetBox="[185,1400,189,1841]" captionTargetId="figure@8.[181,1425,162,1872]" captionTargetPageId="8" captionText="FIGURES 23 – 32. Shape and location of androconial patch, and male and female genitalia of Cuban Calisto. Androconial patch: 23 — Calisto torrei sp. n., 24 — Calisto dissimulatum sp. n., 25 — Calisto aquilum sp. n. Male genitalia, from top to bottom main body in lateral view, aedeagus in lateral view, and aedeagus in dorsal view (scale bar 0.5 mm): 26 — Calisto torrei sp. n., 27 — Calisto dissimulatum sp. n., 28 — Calisto aquilum sp. n. Female genitalia, ventral view (scale bar 0.5 mm): 29 — Calisto torrei sp. n., 30 — Calisto dissimulatum sp. n., 31 — Calisto aquilum sp. n. 32 — Calisto aquilum occidentalis ssp. n." httpUri="https://zenodo.org/record/249427/files/figure.png" pageId="3" pageNumber="506">24, 27, 30</figureCitation>
, 
<figureCitation id="13502A4DFFDE9312FE2CFACD0030FA96" box="[508,538,1364,1388]" captionStart="FIGURES 33 – 36" captionStartId="9.[151,269,1822,1846]" captionTargetBox="[326,1258,201,1784]" captionTargetId="figure@9.[326,1259,194,1785]" captionTargetPageId="9" captionText="FIGURES 33 – 36. Geographical distribution of new species of Cuban Calisto. 33 — Map of Cuba showing areas where the new taxa were found. 34 — Middle rectangle in figure 33, localities where Calisto torrei sp. n. (black circles) and Calisto aquilum sp. n. (white stars) have been found in the Guamuhaya massif. 35 — Rigth rectangle in figure 33, localities where Calisto dissimulatum sp. n. has been found in the Sierra Maestra and Nipe – Sagua – Baracoa massifs. 36 — Left rectangle in figure 33, localities where Calistoaquilum occidentalis ssp. n. has been found in the middle portion of the Guaniguanico massif." httpUri="https://zenodo.org/record/249428/files/figure.png" pageId="3" pageNumber="506">35</figureCitation>
, 
<figureCitation id="13502A4DFFDE9312FDF6FACD0046FA96" box="[550,620,1364,1388]" captionStart-0="FIGURES 51 – 54" captionStart-1="FIGURES 55 – 67" captionStartId-0="13.[151,269,1205,1229]" captionStartId-1="14.[151,269,1926,1950]" captionTargetBox-0="[155,1432,194,1183]" captionTargetBox-1="[207,1379,741,1905]" captionTargetId-0="figure@13.[155,1432,194,1184]" captionTargetId-1="figure@14.[207,1379,741,1905]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 51 – 54. Habitats, host plant, and eggs of Calisto dissimulatum sp. n. 51 — Pine forest near La Zoilita, Sierra Cristal, Nipe – Sagua – Baracoa massif. 52 — Charrascal (scrub forest) near La Zoilita. 53, 54 — Ichnanthus mayariensis (Poaceae), host plant with eggs (arrows) at the underside of leaves." captionText-1="FIGURES 55 – 67. Imatures stages of Calisto dissimulatum sp. n. 55 — First instar. 56 — Second instar. 57 — Third instar. 58 — Fourth instar. Fifth instar: 59 — lateral view, 60 — dorsal view, 61 — head capsule. Pupa showing variation in spot pattern: 62 – 64 ventral view, 65, 66 — lateral view, 67 — dorsal view." httpUri-0="https://zenodo.org/record/249431/files/figure.png" httpUri-1="https://zenodo.org/record/249432/files/figure.png" pageId="3" pageNumber="506">51–67</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFDE9312FF47FA0F0767F8C3" pageId="3" pageNumber="506" type="materials_examined">
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<emphasis id="B91FEADAFFDE9312FF47FA0F03F9FA54" bold="true" box="[151,467,1429,1454]" pageId="3" pageNumber="506">
<typeStatus id="54D0886AFFDE9312FF47FA0F02F8FA54" box="[151,210,1430,1454]" pageId="3" pageNumber="506">Type</typeStatus>
material. Holotype—
</emphasis>
3, 
<collectingCountry id="F37C7658FFDE9312FE25FA0C006FFA57" box="[501,581,1429,1453]" name="Cuba" pageId="3" pageNumber="506">CUBA</collectingCountry>
, Holguín, (“Estación La Zoilita” in the original label) La Zoilita Station, 
<quantity id="4C939B2DFFDE9312FAA2FA0C0287FA2B" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="506" unit="m" value="400.0">400 m</quantity>
<date id="FFD51008FFDE9312FF7DFA20030CFA2B" box="[173,294,1465,1489]" pageId="3" pageNumber="506" value="2010-02-20">20/II/2010</date>
, 
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, R. Núñez, DNA voucher PM07–20 (M037). Deposited in CZACC. 
<emphasis id="B91FEADAFFDE9312FF47FA470301FA0C" bold="true" box="[151,299,1502,1526]" pageId="3" pageNumber="506">Paratypes—</emphasis>
16 3, 9 Ƥ: same data as 
<typeStatus id="54D0886AFFDE9312FD94FA44008CFA0F" box="[580,678,1501,1525]" pageId="3" pageNumber="506" type="holotype">holotype</typeStatus>
except 
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, DNA vouchers RN01–08 (M072) &amp; RN02– 0 1 (M073), genitalia 3 &amp; Ƥ in glycerine (6 3, 3 Ƥ); same data as preceding except ex ova, emerged 
<date id="FFD51008FFDE9312FB2FF99807B6F9E3" box="[1279,1436,1537,1561]" pageId="3" pageNumber="506" value="2012-06-06" valueMax="2012-06-16" valueMin="2012-06-06">6–16/VI/2012</date>
(8 3, 5 Ƥ); road to (“camino de” in the original label) La Zoilita, 
<date id="FFD51008FFDE9312FCA4F9BC01FEF9C7" box="[884,980,1573,1597]" bridgedPair="/" pageId="3" pageNumber="506" value="1986-09">IX/1986</date>
, 
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<geoCoordinate id="EE5F500FFFDE9312FB93F9BC068FF9C7" box="[1091,1189,1573,1597]" direction="west" orientation="longitude" pageId="3" pageNumber="506" precision="925" value="-75.48333">75°29'W</geoCoordinate>
, R. Rodríguez (2 3, 1 Ƥ).Deposited in CZACC.
</paragraph>
<paragraph id="8BD436C8FFDE9312FF17F9F40767F8C3" blockId="3.[151,1437,1429,2034]" pageId="3" pageNumber="506">
<emphasis id="B91FEADAFFDE9312FF17F9F40394F97C" bold="true" box="[199,446,1645,1670]" pageId="3" pageNumber="506">Additional material.</emphasis>
9 3, 3 Ƥ: Santiago de 
<collectingCountry id="F37C7658FFDE9312FD31F9F40135F97F" box="[737,799,1645,1669]" name="Cuba" pageId="3" pageNumber="506">Cuba</collectingCountry>
, Baire, road to (“camino a” in the original label) Las Manuelas, 
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, 
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<geoCoordinate id="EE5F500FFFDE9312FE2AF90800A9F953" box="[506,643,1681,1705]" direction="west" orientation="longitude" pageId="3" pageNumber="506" precision="15" value="-76.36444">76°21'52&quot;W</geoCoordinate>
, 
<date id="FFD51008FFDE9312FD45F9080132F953" box="[661,792,1681,1705]" pageId="3" pageNumber="506" value="1952-09-23">23/IX/1952</date>
, S.L. de La Torre (1 3); Loma del Gato, 
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<geoCoordinate id="EE5F500FFFDE9312FF47F92C0308F937" box="[151,290,1717,1741]" direction="west" orientation="longitude" pageId="3" pageNumber="506" precision="15" value="-76.03778">76°02'16&quot;W</geoCoordinate>
, 
<date id="FFD51008FFDE9312FEFDF92C0389F937" box="[301,419,1717,1741]" pageId="3" pageNumber="506" value="1951-09-06">6/IX/1951</date>
, S.L. de La Torre, genitalia 
<quantity id="4C939B2DFFDE9312FD31F92F013EF937" box="[737,788,1717,1742]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="506" unit="in" value="3.0">3 in</quantity>
glycerine (4 3, 2 Ƥ); same data as preceding except 
<date id="FFD51008FFDE9312FAA2F92C0331F90B" pageId="3" pageNumber="506" value="1952-06-17" valueMax="1952-06-20" valueMin="1952-06-17">17– 20/VI/1952</date>
, F. de Zayas &amp; P. Alayo, genitalia in glycerine, prep. wings RNA187, prep. legs RNA222 (3 3); in CZACC. Guantánamo, Piedra La Vela, 
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, 
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, 
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, R. Núñez (1 3); Guantánamo, Piedra La Vela, El Mulo, 
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, 
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, 
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<geoCoordinate id="EE5F500FFFDE9312FCD1F8B801A1F8C3" box="[769,907,1825,1849]" direction="west" orientation="longitude" pageId="3" pageNumber="506" precision="15" value="-74.90889">74°54'32&quot;W</geoCoordinate>
, R. Núñez (1 3). Deposited in CZACC.
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFDE9312FF17F8DF072AF87B" pageId="3" pageNumber="506" type="etymology">
<paragraph id="8BD436C8FFDE9312FF17F8DF072AF87B" blockId="3.[151,1437,1429,2034]" pageId="3" pageNumber="506">
<emphasis id="B91FEADAFFDE9312FF17F8DF0364F8A4" bold="true" box="[199,334,1862,1886]" pageId="3" pageNumber="506">Etymology.</emphasis>
The species name is derived from the Latin 
<emphasis id="B91FEADAFFDE9312FC90F8DF0181F8A4" box="[832,939,1862,1886]" italics="true" pageId="3" pageNumber="506">dissimulo</emphasis>
(dissimulate, feign), in reference to its great resemblance with 
<taxonomicName id="4C6B4D4BFFDE9312FEB3F8F000BBF87B" authority="Torre 1973" authorityName="Torre" authorityYear="1973" box="[355,657,1897,1921]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFDE9312FEB3F8F00025F87B" box="[355,527,1897,1921]" italics="true" pageId="3" pageNumber="506">Calisto brochei</emphasis>
Torre 1973
</taxonomicName>
and in a less extent with 
<taxonomicName id="4C6B4D4BFFDE9312FC7DF8F006D6F87B" authority="Bates 1935" authorityName="Bates" authorityYear="1935" box="[941,1276,1897,1921]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFDE9312FC7DF8F0065FF87B" box="[941,1141,1897,1921]" italics="true" pageId="3" pageNumber="506">Calisto smintheus</emphasis>
Bates 1935
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFDE9315FF17F814034AFE35" lastPageId="4" lastPageNumber="507" pageId="3" pageNumber="506" type="diagnosis">
<paragraph id="8BD436C8FFDE9315FF17F814034AFE35" blockId="3.[151,1437,1429,2034]" lastBlockId="4.[151,1437,151,1955]" lastPageId="4" lastPageNumber="507" pageId="3" pageNumber="506">
<emphasis id="B91FEADAFFDE9312FF17F814036BF85C" bold="true" box="[199,321,1933,1958]" pageId="3" pageNumber="506">Diagnosis.</emphasis>
<taxonomicName id="4C6B4D4BFFDE9312FE98F817001EF85C" box="[328,564,1934,1958]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFDE9312FE98F817001EF85C" box="[328,564,1934,1958]" italics="true" pageId="3" pageNumber="506">Calisto dissimulatum</emphasis>
</taxonomicName>
is superficially most similar to 
<taxonomicName id="4C6B4D4BFFDE9312FC46F81706CFF85F" authority="Bates 1935" authorityName="Bates" authorityYear="1935" box="[918,1253,1933,1958]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFDE9312FC46F8170674F85C" box="[918,1118,1934,1958]" italics="true" pageId="3" pageNumber="506">Calisto smintheus</emphasis>
Bates 1935
</taxonomicName>
, 
<taxonomicName id="4C6B4D4BFFDE9312FB20F8170339F833" authority="Torre 1973" authorityName="Torre" authorityYear="1973" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFDE9312FB20F81707B6F85C" box="[1264,1436,1934,1958]" italics="true" pageId="3" pageNumber="506">Calisto brochei</emphasis>
Torre 1973
</taxonomicName>
and 
<taxonomicName id="4C6B4D4BFFDE9312FE9CF828002BF833" box="[332,513,1969,1993]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="aquilum">
<emphasis id="B91FEADAFFDE9312FE9CF828002BF833" box="[332,513,1969,1993]" italics="true" pageId="3" pageNumber="506">Calisto aquilum</emphasis>
</taxonomicName>
described below. From the first two, it differs by having an enlarged white dot at M2–M3, a lesser area covered by red scales on underside of forewing cell, and a smaller average size: 3 FWL=17.2± 
<quantity id="4C939B2DFFD99315FEF2FF0E0356FF55" box="[290,380,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="507" unit="mm" value="0.7">0.7 mm</quantity>
, N=16, Ƥ FWL=18.6± 
<quantity id="4C939B2DFFD99315FD51FF0E00F0FF55" box="[641,730,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="507" unit="mm" value="0.3">0.3 mm</quantity>
, N=7 
<emphasis id="B91FEADAFFD99315FCF2FF000143FF4A" box="[802,873,153,176]" italics="true" pageId="4" pageNumber="507">versus</emphasis>
3 FWL=21.9± 
<quantity id="4C939B2DFFD99315FBCCFF0E065EFF55" box="[1052,1140,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="507" unit="mm" value="1.6">1.6 mm</quantity>
, N=50, Ƥ FWL=22.5± 
<quantity id="4C939B2DFFD99315FAABFF0E02EFFF2E" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="507" unit="mm" value="1.4">1.4 mm</quantity>
, N=23 of 
<taxonomicName id="4C6B4D4BFFD99315FEE9FF2403E0FF2E" box="[313,458,188,212]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD99315FEE9FF2403E0FF2E" box="[313,458,188,212]" italics="true" pageId="4" pageNumber="507">C. smintheus</emphasis>
</taxonomicName>
and 3 FWL=20.9± 
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, N=9, Ƥ FWL=21.5± 
<quantity id="4C939B2DFFD99315FC29FF250678FF2E" box="[1017,1106,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="507" unit="mm" value="0.4">0.4 mm</quantity>
, N=5 of 
<taxonomicName id="4C6B4D4BFFD99315FB68FF240704FF2E" box="[1208,1326,188,212]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD99315FB68FF240704FF2E" box="[1208,1326,188,212]" italics="true" pageId="4" pageNumber="507">C. brochei</emphasis>
</taxonomicName>
. From 
<taxonomicName id="4C6B4D4BFFD99315FA52FF24032DFF02" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD99315FA52FF24032DFF02" italics="true" pageId="4" pageNumber="507">C. smintheus</emphasis>
</taxonomicName>
also differs by its less bright overall ventral wing pattern and the absence of the strong reddish splashing at basal two thirds at underside of hindwing. From 
<taxonomicName id="4C6B4D4BFFD99315FCDDFE9C01A2FEE6" box="[781,904,260,284]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD99315FCDDFE9C01A2FEE6" box="[781,904,260,284]" italics="true" pageId="4" pageNumber="507">C. brochei</emphasis>
</taxonomicName>
also lacks the waved outer margin of male androconial patch. Internally, the male genitalia of 
<taxonomicName id="4C6B4D4BFFD99315FD3BFEB1018EFEBA" box="[747,932,296,320]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD99315FD3BFEB1018EFEBA" box="[747,932,296,320]" italics="true" pageId="4" pageNumber="507">C. dissimulatum</emphasis>
</taxonomicName>
possess an uncus proportionally smaller in relation to tegument; the latter is tapered from the mid-point onwards, unlike 
<taxonomicName id="4C6B4D4BFFD99315FC28FED406A2FE9E" box="[1016,1160,332,356]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD99315FC28FED406A2FE9E" box="[1016,1160,332,356]" italics="true" pageId="4" pageNumber="507">C. smintheus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6B4D4BFFD99315FB6EFED40719FE9E" box="[1214,1331,332,356]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD99315FB6EFED40719FE9E" box="[1214,1331,332,356]" italics="true" pageId="4" pageNumber="507">C. brochei</emphasis>
</taxonomicName>
, where it tapers starting from the base The digitiform projections of valvae are straight as in 
<taxonomicName id="4C6B4D4BFFD99315FBADFEE90738FE72" box="[1149,1298,368,392]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD99315FBADFEE90738FE72" box="[1149,1298,368,392]" italics="true" pageId="4" pageNumber="507">C. smintheus</emphasis>
</taxonomicName>
but shorter whereas they are slightly curved in 
<taxonomicName id="4C6B4D4BFFD99315FDE7FE0C0085FE56" box="[567,687,404,428]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD99315FDE7FE0C0085FE56" box="[567,687,404,428]" italics="true" pageId="4" pageNumber="507">C. brochei</emphasis>
</taxonomicName>
. Differences from 
<taxonomicName id="4C6B4D4BFFD99315FC5DFE0C0627FE56" box="[909,1037,404,428]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="aquilum">
<emphasis id="B91FEADAFFD99315FC5DFE0C0627FE56" box="[909,1037,404,428]" italics="true" pageId="4" pageNumber="507">C. aquilum</emphasis>
</taxonomicName>
will be treated below in the latter species diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFD99315FF17FE420347FBE8" pageId="4" pageNumber="507" type="description">
<paragraph id="8BD436C8FFD99315FF17FE420347FBE8" blockId="4.[151,1437,151,1955]" pageId="4" pageNumber="507">
<emphasis id="B91FEADAFFD99315FF17FE420373FE0E" bold="true" box="[199,345,475,500]" pageId="4" pageNumber="507">Description.</emphasis>
Forewing length 3 
<quantity id="4C939B2DFFD99315FD81FE4500D6FE0E" box="[593,764,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" metricValueMax="1.81" metricValueMin="1.63" pageId="4" pageNumber="507" unit="mm" value="17.2" valueMax="18.1" valueMin="16.3">16.3–18.1 mm</quantity>
, Ƥ 
<quantity id="4C939B2DFFD99315FCFFFE4501F0FE0E" box="[815,986,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="1.845" metricValueMax="1.88" metricValueMin="1.81" pageId="4" pageNumber="507" unit="mm" value="18.45" valueMax="18.8" valueMin="18.1">18.1–18.8 mm</quantity>
. Upper surface of wings dark brown, uniform in both sexes; male androconial patch darker than background, almost black, anterior margin behind posterior margin of cell, outer margin parallel to wing margin (
<figureCitation id="13502A4DFFD99315FC8CFDBD01CCFDC6" box="[860,998,548,572]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,269,1848,1872]" captionTargetBox="[253,1332,211,1820]" captionTargetId="figure@5.[247,1337,194,1831]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Types of Calisto from Cuba. Calisto torrei sp. n.: 1 — 3 holotype upper side, 2 — 3 holotype under side, from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 10 / V / 2012, DNA voucher RN 01 – 01 (M 065), 3 — Ƥ paratype upper side, 4 — Ƥ paratype under side, from Cienfuegos, Pico San Juan, 1140 m, 12 / V / 2012, DNA voucher RN 01 – 03 (M 067). Calisto dissimulatum sp. n.: 5 — 3 holotype upper side, 6 — 3 holotype under side, from Holguín, La Zoilita Station, 400 m, 20 / II / 2010, DNA voucher PM 07 – 20 (M 037), 7 — Ƥ paratype upper side, 8 — Ƥ paratype under side, same data as holotype except 3 – 10 / IV / 2012, DNA voucher RN 02 – 01 (M 073)." httpUri="https://zenodo.org/record/249424/files/figure.png" pageId="4" pageNumber="507">Figures 5, 7</figureCitation>
, 
<figureCitation id="13502A4DFFD99315FC22FDBD0638FDC6" box="[1010,1042,548,572]" captionStart="FIGURES 23 – 32" captionStartId="8.[151,269,1842,1866]" captionTargetBox="[185,1400,189,1841]" captionTargetId="figure@8.[181,1425,162,1872]" captionTargetPageId="8" captionText="FIGURES 23 – 32. Shape and location of androconial patch, and male and female genitalia of Cuban Calisto. Androconial patch: 23 — Calisto torrei sp. n., 24 — Calisto dissimulatum sp. n., 25 — Calisto aquilum sp. n. Male genitalia, from top to bottom main body in lateral view, aedeagus in lateral view, and aedeagus in dorsal view (scale bar 0.5 mm): 26 — Calisto torrei sp. n., 27 — Calisto dissimulatum sp. n., 28 — Calisto aquilum sp. n. Female genitalia, ventral view (scale bar 0.5 mm): 29 — Calisto torrei sp. n., 30 — Calisto dissimulatum sp. n., 31 — Calisto aquilum sp. n. 32 — Calisto aquilum occidentalis ssp. n." httpUri="https://zenodo.org/record/249427/files/figure.png" pageId="4" pageNumber="507">24</figureCitation>
). Under surface brown with darker lines, background slightly paler at outer side of post discal line; forewing ocellus with two bluish white iridescent pupils, post discal line externally edged with pale yellow scaling at ocellus area; about two fifths of cell extension red colored, both base and apex brown colored as surrounding background; hindwing background mixed with pale yellow and ochre scales basal to pos dical line which is externally edge by dense pale yellow scaling, heavier around ocellus; post discal area splashed with pale yellow and lilac scales and four white dots at vein interspaces Rs–M1, M1–M2, M2–M3 y M3–Cu1, dot at M2–M3 distinctly larger than M3–Cu1 one and slightly larger than remaining ones; apex of both wings densely covered by ochre scaling around subterminal lines (
<figureCitation id="13502A4DFFD99315FB30FCBB0741FCC0" box="[1248,1387,802,826]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,269,1848,1872]" captionTargetBox="[253,1332,211,1820]" captionTargetId="figure@5.[247,1337,194,1831]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Types of Calisto from Cuba. Calisto torrei sp. n.: 1 — 3 holotype upper side, 2 — 3 holotype under side, from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 10 / V / 2012, DNA voucher RN 01 – 01 (M 065), 3 — Ƥ paratype upper side, 4 — Ƥ paratype under side, from Cienfuegos, Pico San Juan, 1140 m, 12 / V / 2012, DNA voucher RN 01 – 03 (M 067). Calisto dissimulatum sp. n.: 5 — 3 holotype upper side, 6 — 3 holotype under side, from Holguín, La Zoilita Station, 400 m, 20 / II / 2010, DNA voucher PM 07 – 20 (M 037), 7 — Ƥ paratype upper side, 8 — Ƥ paratype under side, same data as holotype except 3 – 10 / IV / 2012, DNA voucher RN 02 – 01 (M 073)." httpUri="https://zenodo.org/record/249424/files/figure.png" pageId="4" pageNumber="507">Figures 6, 8</figureCitation>
, 
<figureCitation id="13502A4DFFD99315FAA9FCBB029DFCA5" captionStart="FIGURES 17 – 22" captionStartId="7.[151,269,1537,1561]" captionTargetBox="[226,1359,194,1515]" captionTargetId="figure@7.[226,1360,194,1516]" captionTargetPageId="7" captionText="FIGURES 17 – 22. Live adults of Cuban Calisto. Calisto torrei sp. n.: 17 — from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 18 — from Sancti Spiritus, Topes de Collante, La Represa, 800 m. Calisto dissimulatum sp. n.: 19, 20 — from Holguín, alrededores de La Zoilita, 400 m. Calisto aquilum sp. n.: 21 — from Cienfuegos, road to Pico San Juan, 22 — from Cienfuegos, Pico San Juan." httpUri="https://zenodo.org/record/249426/files/figure.png" pageId="4" pageNumber="507">19, 20</figureCitation>
). Male genitalia with tegumen about two thirds the length of uncus, slightly concave at the middle and rounded at posterior half; uncus slightly arched, basal half about the same height then gradually tapering toward apex; digitiform projection of valvae, including ventral margin, straight and moderately short; aedeagus sinuated with distinct right curves at both basal and apical halves and an additional smaller curve at the middle (
<figureCitation id="13502A4DFFD99315FACAFC2B07BAFC30" box="[1306,1424,946,970]" captionStart="FIGURES 23 – 32" captionStartId="8.[151,269,1842,1866]" captionTargetBox="[185,1400,189,1841]" captionTargetId="figure@8.[181,1425,162,1872]" captionTargetPageId="8" captionText="FIGURES 23 – 32. Shape and location of androconial patch, and male and female genitalia of Cuban Calisto. Androconial patch: 23 — Calisto torrei sp. n., 24 — Calisto dissimulatum sp. n., 25 — Calisto aquilum sp. n. Male genitalia, from top to bottom main body in lateral view, aedeagus in lateral view, and aedeagus in dorsal view (scale bar 0.5 mm): 26 — Calisto torrei sp. n., 27 — Calisto dissimulatum sp. n., 28 — Calisto aquilum sp. n. Female genitalia, ventral view (scale bar 0.5 mm): 29 — Calisto torrei sp. n., 30 — Calisto dissimulatum sp. n., 31 — Calisto aquilum sp. n. 32 — Calisto aquilum occidentalis ssp. n." httpUri="https://zenodo.org/record/249427/files/figure.png" pageId="4" pageNumber="507">Figure 27</figureCitation>
). Female genitalia with dorsal crown tall, height about 0.33x width; corpus bursae about the same length of ductus bursae (
<figureCitation id="13502A4DFFD99315FF3EFC63034AFBE8" box="[238,352,1018,1042]" captionStart="FIGURES 23 – 32" captionStartId="8.[151,269,1842,1866]" captionTargetBox="[185,1400,189,1841]" captionTargetId="figure@8.[181,1425,162,1872]" captionTargetPageId="8" captionText="FIGURES 23 – 32. Shape and location of androconial patch, and male and female genitalia of Cuban Calisto. Androconial patch: 23 — Calisto torrei sp. n., 24 — Calisto dissimulatum sp. n., 25 — Calisto aquilum sp. n. Male genitalia, from top to bottom main body in lateral view, aedeagus in lateral view, and aedeagus in dorsal view (scale bar 0.5 mm): 26 — Calisto torrei sp. n., 27 — Calisto dissimulatum sp. n., 28 — Calisto aquilum sp. n. Female genitalia, ventral view (scale bar 0.5 mm): 29 — Calisto torrei sp. n., 30 — Calisto dissimulatum sp. n., 31 — Calisto aquilum sp. n. 32 — Calisto aquilum occidentalis ssp. n." httpUri="https://zenodo.org/record/249427/files/figure.png" pageId="4" pageNumber="507">Figure 30</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFD99315FF17FB87069AFBA0" pageId="4" pageNumber="507" type="distribution">
<paragraph id="8BD436C8FFD99315FF17FB87069AFBA0" blockId="4.[151,1437,151,1955]" pageId="4" pageNumber="507">
<emphasis id="B91FEADAFFD99315FF17FB87034AFBCD" bold="true" box="[199,352,1054,1079]" pageId="4" pageNumber="507">Distribution.</emphasis>
<taxonomicName id="4C6B4D4BFFD99315FEBAFB860073FBCD" box="[362,601,1055,1079]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD99315FEBAFB860073FBCD" box="[362,601,1055,1079]" italics="true" pageId="4" pageNumber="507">Calisto dissimulatum</emphasis>
</taxonomicName>
is known from few localities of the western halves of the Nipe–Sagua– Baracoa and Sierra Maestra, the two major mountain ranges of eastern 
<collectingCountry id="F37C7658FFD99315FC63FBDB01C4FBA0" box="[947,1006,1090,1114]" name="Cuba" pageId="4" pageNumber="507">Cuba</collectingCountry>
(
<figureCitation id="13502A4DFFD99315FC2DFBDB0689FBA0" box="[1021,1187,1090,1114]" captionStart="FIGURES 33 – 36" captionStartId="9.[151,269,1822,1846]" captionTargetBox="[326,1258,201,1784]" captionTargetId="figure@9.[326,1259,194,1785]" captionTargetPageId="9" captionText="FIGURES 33 – 36. Geographical distribution of new species of Cuban Calisto. 33 — Map of Cuba showing areas where the new taxa were found. 34 — Middle rectangle in figure 33, localities where Calisto torrei sp. n. (black circles) and Calisto aquilum sp. n. (white stars) have been found in the Guamuhaya massif. 35 — Rigth rectangle in figure 33, localities where Calisto dissimulatum sp. n. has been found in the Sierra Maestra and Nipe – Sagua – Baracoa massifs. 36 — Left rectangle in figure 33, localities where Calistoaquilum occidentalis ssp. n. has been found in the middle portion of the Guaniguanico massif." httpUri="https://zenodo.org/record/249428/files/figure.png" pageId="4" pageNumber="507">Figures 33, 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFD9931BFF17FBFF03A0FF2D" lastPageId="10" lastPageNumber="513" pageId="4" pageNumber="507" type="biology_ecology">
<paragraph id="8BD436C8FFD99315FF17FBFF01ABFB58" blockId="4.[151,1437,151,1955]" pageId="4" pageNumber="507">
<emphasis id="B91FEADAFFD99315FF17FBFF030DFB85" bold="true" box="[199,295,1126,1151]" pageId="4" pageNumber="507">Biology.</emphasis>
Collecting sites are at low elevations (
<quantity id="4C939B2DFFD99315FD30FBFE014FFB85" box="[736,869,1127,1151]" metricMagnitude="2" metricUnit="m" metricValue="6.25" metricValueMax="8.5" metricValueMin="4.0" pageId="4" pageNumber="507" unit="m" value="625.0" valueMax="850.0" valueMin="400.0">400–850 m</quantity>
). Habitats include rainforest, broad leaf evergreen forest, pine forest, and charrascales (scrub forest) (
<figureCitation id="13502A4DFFD99315FD1DFB13015EFB58" box="[717,884,1162,1186]" captionStart="FIGURES 51 – 54" captionStartId="13.[151,269,1205,1229]" captionTargetBox="[155,1432,194,1183]" captionTargetId="figure@13.[155,1432,194,1184]" captionTargetPageId="13" captionText="FIGURES 51 – 54. Habitats, host plant, and eggs of Calisto dissimulatum sp. n. 51 — Pine forest near La Zoilita, Sierra Cristal, Nipe – Sagua – Baracoa massif. 52 — Charrascal (scrub forest) near La Zoilita. 53, 54 — Ichnanthus mayariensis (Poaceae), host plant with eggs (arrows) at the underside of leaves." httpUri="https://zenodo.org/record/249431/files/figure.png" pageId="4" pageNumber="507">Figures 51, 52</figureCitation>
).
</paragraph>
<paragraph id="8BD436C8FFD99315FF17FB28063CFAA6" blockId="4.[151,1437,151,1955]" pageId="4" pageNumber="507">
On 
<date id="FFD51008FFD99315FF21FB2803A1FB33" box="[241,395,1198,1225]" pageId="4" pageNumber="507" value="2012-04-05">April 5th 2012</date>
a female was observed at 12:30 pm ovipositing twice on a small grass species. A three egg cluster and one single egg were laid glued to the underside of host leaves (Figures 53,54). Host plant was classified as 
<taxonomicName id="4C6B4D4BFFD99315FF67FB60007DFAE8" authority="Hitchc. Eggs" authorityName="Hitchc. Eggs" box="[183,599,1273,1298]" class="Liliopsida" family="Poaceae" genus="Ichnanthus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="507" phylum="Tracheophyta" rank="species" species="mayarensis">
<emphasis id="B91FEADAFFD99315FF67FB600391FAEB" box="[183,443,1273,1297]" italics="true" pageId="4" pageNumber="507">Ichnanthus mayarensis</emphasis>
Hitchc. Eggs
</taxonomicName>
were pale yellow when laid and after 24 hours turned to beige with tiny orange brown spots. 11 additional eggs were obtained on 6– 
<date id="FFD51008FFD99315FCA2FAB90154FAC2" box="[882,894,1312,1336]" pageId="4" pageNumber="507">8</date>
th April from two females confined to plastic containers. All eggs (n=15) hatched after 6–8 days. Larvae ate the entire corion.
</paragraph>
<paragraph id="8BD436C8FFD99315FF17FAF106E2FA32" blockId="4.[151,1437,151,1955]" pageId="4" pageNumber="507">
First instar (
<figureCitation id="13502A4DFFD99315FE81FAF103EAFA7A" box="[337,448,1384,1408]" captionStart="FIGURES 55 – 67" captionStartId="14.[151,269,1926,1950]" captionTargetBox="[207,1379,741,1905]" captionTargetId="figure@14.[207,1379,741,1905]" captionTargetPageId="14" captionText="FIGURES 55 – 67. Imatures stages of Calisto dissimulatum sp. n. 55 — First instar. 56 — Second instar. 57 — Third instar. 58 — Fourth instar. Fifth instar: 59 — lateral view, 60 — dorsal view, 61 — head capsule. Pupa showing variation in spot pattern: 62 – 64 ventral view, 65, 66 — lateral view, 67 — dorsal view." httpUri="https://zenodo.org/record/249432/files/figure.png" pageId="4" pageNumber="507">Figure 55</figureCitation>
)–Head width 
<quantity id="4C939B2DFFD99315FDB4FAF10121FA7A" box="[612,779,1384,1408]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="6.8" metricValueMin="6.2" pageId="4" pageNumber="507" unit="mm" value="0.65" valueMax="0.68" valueMin="0.62">0.62–0.68 mm</quantity>
, height 
<quantity id="4C939B2DFFD99315FCB7FAF10625FA7A" box="[871,1039,1384,1408]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" pageId="4" pageNumber="507" unit="mm" value="0.61" valueMax="0.64" valueMin="0.58">0.58–0.64 mm</quantity>
. Initial length 
<quantity id="4C939B2DFFD99315FB65FAF10776FA7A" box="[1205,1372,1384,1408]" metricMagnitude="-3" metricUnit="m" metricValue="2.895" metricValueMax="3.17" metricValueMin="2.62" pageId="4" pageNumber="507" unit="mm" value="2.895" valueMax="3.17" valueMin="2.62">2.62–3.17 mm</quantity>
, final length 
<quantity id="4C939B2DFFD99315FF37FA1503BAFA5E" box="[231,400,1420,1444]" metricMagnitude="-3" metricUnit="m" metricValue="4.045" metricValueMax="4.39" metricValueMin="3.7" pageId="4" pageNumber="507" unit="mm" value="4.045" valueMax="4.39" valueMin="3.7">3.70–4.39 mm</quantity>
. Head dark brown, almost black; body pale beige with 7 thin reddish brown longitudinal lines. After feeding on grass background turned pale green. First molt occurred after 9–12 days.
</paragraph>
<paragraph id="8BD436C8FFD99315FF17FA4D0139F858" blockId="4.[151,1437,151,1955]" pageId="4" pageNumber="507">
Second to fifth instars (
<figureCitation id="13502A4DFFD99315FE00FA4D005AFA16" box="[464,624,1492,1516]" captionStart="FIGURES 55 – 67" captionStartId="14.[151,269,1926,1950]" captionTargetBox="[207,1379,741,1905]" captionTargetId="figure@14.[207,1379,741,1905]" captionTargetPageId="14" captionText="FIGURES 55 – 67. Imatures stages of Calisto dissimulatum sp. n. 55 — First instar. 56 — Second instar. 57 — Third instar. 58 — Fourth instar. Fifth instar: 59 — lateral view, 60 — dorsal view, 61 — head capsule. Pupa showing variation in spot pattern: 62 – 64 ventral view, 65, 66 — lateral view, 67 — dorsal view." httpUri="https://zenodo.org/record/249432/files/figure.png" pageId="4" pageNumber="507">Figures 56–61</figureCitation>
)—After first molt the coloration of larva changed to a new pattern that persisted during subsequent instars with the only variation being a gradual darkening toward final instar. Head width, height, final length and duration of second to fifth instars were: 0.78–0.90, 0.78–0.90, 
<quantity id="4C939B2DFFD99315FB69F9850769F9CE" box="[1209,1347,1564,1588]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.6" metricValueMin="5.7" pageId="4" pageNumber="507" unit="mm" value="6.15" valueMax="6.6" valueMin="5.7">5.7–6.6 mm</quantity>
, and 7– 10 days at the 2nd (n=15), l.16–1.32, 1.18–1.30, 
<quantity id="4C939B2DFFD99315FD61F9DA0111F9A1" box="[689,827,1603,1627]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.7" metricValueMin="8.5" pageId="4" pageNumber="507" unit="mm" value="9.1" valueMax="9.7" valueMin="8.5">8.5–9.7 mm</quantity>
, and 6–8 days at the 3rd (n=12), 1.63–1.78, 1.73–1.85, 
<quantity id="4C939B2DFFD99315FF47F9F30314F978" box="[151,318,1642,1666]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.48" metricValueMin="1.34" pageId="4" pageNumber="507" unit="mm" value="14.1" valueMax="14.8" valueMin="13.4">13.4–14.8 mm</quantity>
, and 7–11 days at the 4th (n=12), and 2.24–2.40, 2.48–2.68, 
<quantity id="4C939B2DFFD99315FC3CF9F30640F978" box="[1004,1130,1642,1666]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="4" pageNumber="507" unit="mm" value="22.5" valueMax="23.0" valueMin="22.0">22–23 mm</quantity>
, and 10–15 days at the 5th, respectively. Head background pale grayish brown heavily speckle in dark brown and with the following dark brown, almost black, marks: lateral vertical lines from stemmata to horns; X shaped mark on epicranium; a complete transverse band across the frontoclypeus. Body background stramineous with an increasing ochre tint toward final instar; lines brown, almost black at final instar; dorsal line with dots on sides at the beginning and end of each segment; subdorsal lines in zig zag with angles closest to dorsal line at the middle of each segment; suprastigmatal line diffuse, present as discontinuous speckling, forming dots over spiracles; spiracle dark brown surrounded by a contrasting white oval shaped area; stigmatal line waved, convex between spiracles; infrastigmatal line straight; sides of last abdominal segment tails white.
</paragraph>
<caption id="DF146640FFD89314FF47F8A101B4F815" httpUri="https://zenodo.org/record/249424/files/figure.png" pageId="5" pageNumber="508" targetBox="[253,1332,211,1820]" targetPageId="5">
<paragraph id="8BD436C8FFD89314FF47F8A101B4F815" blockId="5.[151,1436,1848,2031]" pageId="5" pageNumber="508">
<emphasis id="B91FEADAFFD89314FF47F8A10368F8AA" bold="true" box="[151,322,1848,1872]" pageId="5" pageNumber="508">FIGURES 1–8.</emphasis>
Types of 
<taxonomicName id="4C6B4D4BFFD89314FE7CF8A103D3F8B5" box="[428,505,1848,1871]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEADAFFD89314FE7CF8A103D3F8B5" box="[428,505,1848,1871]" italics="true" pageId="5" pageNumber="508">Calisto</emphasis>
</taxonomicName>
from Cuba. 
<taxonomicName id="4C6B4D4BFFD89314FDAFF8A10125F8B5" box="[639,783,1848,1871]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="torrei" status="sp. nov.">
<emphasis id="B91FEADAFFD89314FDAFF8A10125F8B5" box="[639,783,1848,1871]" italics="true" pageId="5" pageNumber="508">Calisto torrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD89314FCC5F8A1017AF8AA" box="[789,848,1848,1872]" pageId="5" pageNumber="508" rank="species">
<emphasis id="B91FEADAFFD89314FCC5F8A1017AF8AA" bold="true" box="[789,848,1848,1872]" pageId="5" pageNumber="508">sp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFD89314FC8FF8A101ADF8AA" bold="true" box="[863,903,1848,1872]" pageId="5" pageNumber="508">1—</emphasis>
3 holotype upper side, 
<emphasis id="B91FEADAFFD89314FB51F8A10682F8AA" bold="true" box="[1153,1192,1848,1872]" pageId="5" pageNumber="508">2—</emphasis>
3 holotype under side, from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 10/V/2012, DNA voucher RN01–01 (M065), 
<emphasis id="B91FEADAFFD89314FB2CF8C1070EF88A" bold="true" box="[1276,1316,1880,1904]" pageId="5" pageNumber="508">3—</emphasis>
Ƥ paratype upper side, 
<emphasis id="B91FEADAFFD89314FEC8F8E1036BF86A" bold="true" box="[280,321,1912,1936]" pageId="5" pageNumber="508">4—</emphasis>
Ƥ paratype under side, from Cienfuegos, Pico San Juan, 1140m, 12/V/2012, DNA voucher RN01–03 (M067). 
<taxonomicName id="4C6B4D4BFFD89314FF28F80103F6F855" box="[248,476,1944,1967]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFD89314FF28F80103F6F855" box="[248,476,1944,1967]" italics="true" pageId="5" pageNumber="508">Calisto dissimulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD89314FE35F8010009F84A" box="[485,547,1944,1968]" pageId="5" pageNumber="508" rank="species">
<emphasis id="B91FEADAFFD89314FE35F8010009F84A" bold="true" box="[485,547,1944,1968]" pageId="5" pageNumber="508">sp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFD89314FDE4F8010071F84A" bold="true" box="[564,603,1944,1968]" pageId="5" pageNumber="508">5—</emphasis>
3 holotype upper side, 
<emphasis id="B91FEADAFFD89314FCB0F80101A2F84A" bold="true" box="[864,904,1944,1968]" pageId="5" pageNumber="508">6—</emphasis>
3 holotype under side, from Holguín, La Zoilita Station, 400 m, 20/II/2010, DNA voucher PM07–20 (M037), 
<emphasis id="B91FEADAFFD89314FCF6F8210165F82A" bold="true" box="[806,847,1976,2000]" pageId="5" pageNumber="508">7—</emphasis>
Ƥ paratype upper side, 
<emphasis id="B91FEADAFFD89314FB96F8210644F82A" bold="true" box="[1094,1134,1976,2000]" pageId="5" pageNumber="508">8—</emphasis>
Ƥ paratype under side, same data as holotype except 3–10/IV/2012, DNA voucher RN02–01 (M073).
</paragraph>
</caption>
<caption id="DF146640FFDB9317FF47F8970734F83F" httpUri="https://zenodo.org/record/249425/files/figure.png" pageId="6" pageNumber="509" targetBox="[298,1290,201,1777]" targetPageId="6">
<paragraph id="8BD436C8FFDB9317FF47F8970734F83F" blockId="6.[151,1436,1806,1989]" pageId="6" pageNumber="509">
<emphasis id="B91FEADAFFDB9317FF47F897037AF8DC" bold="true" box="[151,336,1806,1830]" pageId="6" pageNumber="509">FIGURES 9–16.</emphasis>
Types of 
<taxonomicName id="4C6B4D4BFFDB9317FE69F897002CF8DF" box="[441,518,1806,1829]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEADAFFDB9317FE69F897002CF8DF" box="[441,518,1806,1829]" italics="true" pageId="6" pageNumber="509">Calisto</emphasis>
</taxonomicName>
from Cuba. 
<taxonomicName id="4C6B4D4BFFDB9317FD5DF8970112F8DF" box="[653,824,1806,1829]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="509" phylum="Arthropoda" rank="species" species="aquilum" status="sp. nov.">
<emphasis id="B91FEADAFFDB9317FD5DF8970112F8DF" box="[653,824,1806,1829]" italics="true" pageId="6" pageNumber="509">Calisto aquilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFDB9317FCEFF8970150F8DC" box="[831,890,1806,1830]" pageId="6" pageNumber="509" rank="species">
<emphasis id="B91FEADAFFDB9317FCEFF8970150F8DC" bold="true" box="[831,890,1806,1830]" pageId="6" pageNumber="509">sp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFDB9317FC59F897019BF8DC" bold="true" box="[905,945,1806,1830]" pageId="6" pageNumber="509">9—</emphasis>
3 holotype upper side, 
<emphasis id="B91FEADAFFDB9317FB7BF89706CBF8DC" bold="true" box="[1195,1249,1806,1830]" pageId="6" pageNumber="509">10—</emphasis>
3 holotype under side, from Sancti Spiritus, Trinidad, ladera norte de Pico Potrerillo, 750 
<emphasis id="B91FEADAFFDB9317FC7DF8B70190F8BC" bold="true" box="[941,954,1838,1862]" pageId="6" pageNumber="509">–</emphasis>
850 m, 11/V/2012, DNA voucher RN01–02 (M070), 
<emphasis id="B91FEADAFFDB9317FF28F8D70300F89C" bold="true" box="[248,298,1870,1894]" pageId="6" pageNumber="509">11—</emphasis>
Ƥ paratype upper side, 
<emphasis id="B91FEADAFFDB9317FDFDF8D70049F89C" bold="true" box="[557,611,1870,1894]" pageId="6" pageNumber="509">12—</emphasis>
Ƥ paratype under side, same data as holotype. 
<taxonomicName id="4C6B4D4BFFDB9317FBB5F8D707B6F89F" box="[1125,1436,1870,1893]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="509" phylum="Arthropoda" rank="subSpecies" species="aquilum" status="subsp. nov." subSpecies="occidentalis">
<emphasis id="B91FEADAFFDB9317FBB5F8D707B6F89F" box="[1125,1436,1870,1893]" italics="true" pageId="6" pageNumber="509">Calisto aquilum occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFDB9317FF47F8F702F6F87C" box="[151,220,1902,1926]" pageId="6" pageNumber="509" rank="subSpecies">
<emphasis id="B91FEADAFFDB9317FF47F8F702F6F87C" bold="true" box="[151,220,1902,1926]" pageId="6" pageNumber="509">ssp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFDB9317FF3AF8F70335F87C" bold="true" box="[234,287,1902,1926]" pageId="6" pageNumber="509">13—</emphasis>
Ƥ holotype upper side, 
<emphasis id="B91FEADAFFDB9317FDC7F8F70067F87C" bold="true" box="[535,589,1902,1926]" pageId="6" pageNumber="509">14—</emphasis>
Ƥ holotype under side, from Pinar del Río, base sur de Pan de Azúcar, 75 m, 25/ IX/2011, DNA voucher PM20–02 (M056), 
<emphasis id="B91FEADAFFDB9317FDBBF817008AF85C" bold="true" box="[619,672,1934,1958]" pageId="6" pageNumber="509">15—</emphasis>
Ƥ paratype upper side, 
<emphasis id="B91FEADAFFDB9317FC4BF81701FAF85C" bold="true" box="[923,976,1934,1958]" pageId="6" pageNumber="509">16—</emphasis>
Ƥ paratype under side, from Pinar del Río, Sendero Las Maravillas (8 km SW of Viñales village), 200 m, 24/IX/2011, DNA voucher PM20–03 (M057).
</paragraph>
</caption>
<caption id="DF146640FFDA9316FF47F9980000F982" httpUri="https://zenodo.org/record/249426/files/figure.png" pageId="7" pageNumber="510" targetBox="[226,1359,194,1515]" targetPageId="7">
<paragraph id="8BD436C8FFDA9316FF47F9980000F982" blockId="7.[151,1436,1537,1657]" pageId="7" pageNumber="510">
<emphasis id="B91FEADAFFDA9316FF47F9980373F9E3" bold="true" box="[151,345,1537,1561]" pageId="7" pageNumber="510">FIGURES 17–22</emphasis>
. Live adults of Cuban 
<taxonomicName id="4C6B4D4BFFDA9316FD8CF99B0083F9E3" box="[604,681,1538,1561]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEADAFFDA9316FD8CF99B0083F9E3" box="[604,681,1538,1561]" italics="true" pageId="7" pageNumber="510">Calisto</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C6B4D4BFFDA9316FD69F99B0166F9E3" box="[697,844,1538,1561]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="510" phylum="Arthropoda" rank="species" species="torrei" status="sp. nov.">
<emphasis id="B91FEADAFFDA9316FD69F99B0166F9E3" box="[697,844,1538,1561]" italics="true" pageId="7" pageNumber="510">Calisto torrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFDA9316FC85F99801BEF9E3" box="[853,916,1537,1561]" pageId="7" pageNumber="510" rank="species">
<emphasis id="B91FEADAFFDA9316FC85F99801BEF9E3" bold="true" box="[853,916,1537,1561]" pageId="7" pageNumber="510">sp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFDA9316FC75F99801F0F9E3" bold="true" box="[933,986,1537,1561]" pageId="7" pageNumber="510">17—</emphasis>
from Sancti Spiritus, Banao, subida a La Sabina, 450 m, 
<emphasis id="B91FEADAFFDA9316FE90F9B8035FF9C3" bold="true" box="[320,373,1569,1593]" pageId="7" pageNumber="510">18—</emphasis>
from Sancti Spiritus, Topes de Collante, La Represa, 800 m. 
<taxonomicName id="4C6B4D4BFFDA9316FBD7F9BB06C0F9C3" box="[1031,1258,1570,1593]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="510" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFDA9316FBD7F9BB06C0F9C3" box="[1031,1258,1570,1593]" italics="true" pageId="7" pageNumber="510">Calisto dissimulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFDA9316FB22F9B80704F9C3" box="[1266,1326,1569,1593]" pageId="7" pageNumber="510" rank="species">
<emphasis id="B91FEADAFFDA9316FB22F9B80704F9C3" bold="true" box="[1266,1326,1569,1593]" pageId="7" pageNumber="510">sp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFDA9316FAEDF9B8077DF9C3" bold="true" box="[1341,1367,1569,1593]" pageId="7" pageNumber="510">19</emphasis>
, 
<emphasis id="B91FEADAFFDA9316FAB6F9B807B1F9C3" bold="true" box="[1382,1435,1569,1593]" pageId="7" pageNumber="510">20—</emphasis>
from Holguín, alrededores de La Zoilita, 400m. 
<taxonomicName id="4C6B4D4BFFDA9316FD77F9DB017EF9A3" box="[679,852,1602,1625]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="510" phylum="Arthropoda" rank="species" species="aquilum" status="sp. nov.">
<emphasis id="B91FEADAFFDA9316FD77F9DB017EF9A3" box="[679,852,1602,1625]" italics="true" pageId="7" pageNumber="510">Calisto aquilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFDA9316FC8DF9D801B0F9A3" box="[861,922,1601,1625]" pageId="7" pageNumber="510" rank="species">
<emphasis id="B91FEADAFFDA9316FC8DF9D801B0F9A3" bold="true" box="[861,922,1601,1625]" pageId="7" pageNumber="510">sp. n.</emphasis>
</taxonomicNameLabel>
: 
<emphasis id="B91FEADAFFDA9316FC7BF9D801CBF9A3" bold="true" box="[939,993,1601,1625]" pageId="7" pageNumber="510">21—</emphasis>
from Cienfuegos, road to Pico San Juan, 
<emphasis id="B91FEADAFFDA9316FF47F9F802E6F983" bold="true" box="[151,204,1633,1657]" pageId="7" pageNumber="510">22—</emphasis>
from Cienfuegos, Pico San Juan.
</paragraph>
</caption>
<paragraph id="8BD436C8FFDA9316FF17F93C0758F947" blockId="7.[151,1436,1701,2013]" box="[199,1394,1701,1726]" pageId="7" pageNumber="510">The prepupal stage lasts three days, two in a straight position, followed by one day in J position until molt.</paragraph>
<paragraph id="8BD436C8FFDA9316FF17F953005FF827" blockId="7.[151,1436,1701,2013]" pageId="7" pageNumber="510">
Pupa (
<figureCitation id="13502A4DFFDA9316FEC0F950039FF91B" box="[272,437,1737,1761]" captionStart="FIGURES 55 – 67" captionStartId="14.[151,269,1926,1950]" captionTargetBox="[207,1379,741,1905]" captionTargetId="figure@14.[207,1379,741,1905]" captionTargetPageId="14" captionText="FIGURES 55 – 67. Imatures stages of Calisto dissimulatum sp. n. 55 — First instar. 56 — Second instar. 57 — Third instar. 58 — Fourth instar. Fifth instar: 59 — lateral view, 60 — dorsal view, 61 — head capsule. Pupa showing variation in spot pattern: 62 – 64 ventral view, 65, 66 — lateral view, 67 — dorsal view." httpUri="https://zenodo.org/record/249432/files/figure.png" pageId="7" pageNumber="510">Figures 62–67</figureCitation>
)–Length 
<quantity id="4C939B2DFFDA9316FDF7F9500089F91B" box="[551,675,1737,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="7" pageNumber="510" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
, maximum width 4.0– 
<quantity id="4C939B2DFFDA9316FC79F950062BF91B" box="[937,1025,1737,1761]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="510" unit="mm" value="4.5">4.5 mm</quantity>
(n=12). Background pale brownish gray with a dark brown spot pattern whose intensity varies between individuals; however constant spots were traced on the available series: a pair of ventral black dots on eyes and on apex of metathoracic tibiae, and first two tarsal segments; a pair of orange dots at the base of labial palpi; a row of tiny dark brown dots on veins near wing outer margin; a waved dark spot at the middle of the wing sheaths, a dark dot near costa at apical third, and a discontinuous series of dark spots on dorsal margin; abdomen with a dark brown line on sides and transverse dark band on dorsum, heavier on the reduced crests of segments 1 to 6; last abdominal segment long, stout, cremaster area enlarged, broad and slightly flattened.
</paragraph>
<caption id="DF146640FFD59319FF47F8AB0061F810" httpUri="https://zenodo.org/record/249427/files/figure.png" pageId="8" pageNumber="511" targetBox="[185,1400,189,1841]" targetPageId="8">
<paragraph id="8BD436C8FFD59319FF47F8AB0061F810" blockId="8.[151,1436,1842,2026]" pageId="8" pageNumber="511">
<emphasis id="B91FEADAFFD59319FF47F8AB034CF8B0" bold="true" box="[151,358,1842,1866]" pageId="8" pageNumber="511">FIGURES 23–32.</emphasis>
Shape and location of androconial patch, and male and female genitalia of Cuban 
<taxonomicName id="4C6B4D4BFFD59319FA98F8AA07BFF8B0" box="[1352,1429,1843,1866]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEADAFFD59319FA98F8AA07BFF8B0" box="[1352,1429,1843,1866]" italics="true" pageId="8" pageNumber="511">Calisto</emphasis>
</taxonomicName>
. Androconial patch: 
<emphasis id="B91FEADAFFD59319FEA7F8CB0386F890" bold="true" box="[375,428,1874,1898]" pageId="8" pageNumber="511">23—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FE7CF8CA0068F890" box="[428,578,1875,1898]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="torrei" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FE7CF8CA0068F890" box="[428,578,1875,1898]" italics="true" pageId="8" pageNumber="511">Calisto torrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FD9EF8CB00BAF890" box="[590,656,1874,1898]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FD9EF8CB00BAF890" bold="true" box="[590,656,1874,1898]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B91FEADAFFD59319FD74F8CB00F3F890" bold="true" box="[676,729,1874,1898]" pageId="8" pageNumber="511">24—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FD09F8CA01EBF890" box="[729,961,1875,1898]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FD09F8CA01EBF890" box="[729,961,1875,1898]" italics="true" pageId="8" pageNumber="511">Calisto dissimulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FC1DF8CB0625F890" box="[973,1039,1874,1898]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FC1DF8CB0625F890" bold="true" box="[973,1039,1874,1898]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B91FEADAFFD59319FBF3F8CB0672F890" bold="true" box="[1059,1112,1874,1898]" pageId="8" pageNumber="511">25—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FB88F8CA0723F890" box="[1112,1289,1875,1898]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="aquilum" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FB88F8CA0723F890" box="[1112,1289,1875,1898]" italics="true" pageId="8" pageNumber="511">Calisto aquilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FAC6F8CB0772F890" box="[1302,1368,1874,1898]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FAC6F8CB0772F890" bold="true" box="[1302,1368,1874,1898]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
Male genitalia, from top to bottom main body in lateral view, aedeagus in lateral view, and aedeagus in dorsal view (scale bar 0.5 mm): 
<emphasis id="B91FEADAFFD59319FED8F80B0317F850" bold="true" box="[264,317,1938,1962]" pageId="8" pageNumber="511">26—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FEEDF80A03FBF850" box="[317,465,1939,1962]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="torrei" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FEEDF80A03FBF850" box="[317,465,1939,1962]" italics="true" pageId="8" pageNumber="511">Calisto torrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FE0CF80B0031F850" box="[476,539,1938,1962]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FE0CF80B0031F850" bold="true" box="[476,539,1938,1962]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B91FEADAFFD59319FDFDF80B0048F850" bold="true" box="[557,610,1938,1962]" pageId="8" pageNumber="511">27—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FDB3F80A0163F850" box="[611,841,1939,1962]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FDB3F80A0163F850" box="[611,841,1939,1962]" italics="true" pageId="8" pageNumber="511">Calisto dissimulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FC84F80B01BEF850" box="[852,916,1938,1962]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FC84F80B01BEF850" bold="true" box="[852,916,1938,1962]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B91FEADAFFD59319FC75F80B01F0F850" bold="true" box="[933,986,1938,1962]" pageId="8" pageNumber="511">28—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FC0BF80A06A3F850" box="[987,1161,1939,1962]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="aquilum" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FC0BF80A06A3F850" box="[987,1161,1939,1962]" italics="true" pageId="8" pageNumber="511">Calisto aquilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FB45F80B06FFF850" box="[1173,1237,1938,1962]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FB45F80B06FFF850" bold="true" box="[1173,1237,1938,1962]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
Female genitalia, ventral view (scale bar 0.5 mm): 
<emphasis id="B91FEADAFFD59319FE2AF82B0005F830" bold="true" box="[506,559,1970,1994]" pageId="8" pageNumber="511">29—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FDFFF82A0095F830" box="[559,703,1971,1994]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="torrei" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FDFFF82A0095F830" box="[559,703,1971,1994]" italics="true" pageId="8" pageNumber="511">Calisto torrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FD16F82B0128F830" box="[710,770,1970,1994]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FD16F82B0128F830" bold="true" box="[710,770,1970,1994]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B91FEADAFFD59319FCC0F82B016FF830" bold="true" box="[784,837,1970,1994]" pageId="8" pageNumber="511">30—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FC95F82A060DF830" box="[837,1063,1971,1994]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FC95F82A060DF830" box="[837,1063,1971,1994]" italics="true" pageId="8" pageNumber="511">Calisto dissimulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FBFEF82B0640F830" box="[1070,1130,1970,1994]" pageId="8" pageNumber="511" rank="species">
<emphasis id="B91FEADAFFD59319FBFEF82B0640F830" bold="true" box="[1070,1130,1970,1994]" pageId="8" pageNumber="511">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<emphasis id="B91FEADAFFD59319FBA8F82B0686F830" bold="true" box="[1144,1196,1970,1994]" pageId="8" pageNumber="511">31—</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FB7DF82A0772F830" box="[1197,1368,1971,1994]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="species" species="aquilum" status="sp. nov.">
<emphasis id="B91FEADAFFD59319FB7DF82A0772F830" box="[1197,1368,1971,1994]" italics="true" pageId="8" pageNumber="511">Calisto aquilum</emphasis>
</taxonomicName>
<emphasis id="B91FEADAFFD59319FAB0F82B02E6F810" bold="true" pageId="8" pageNumber="511">
<taxonomicNameLabel id="A22C57A1FFD59319FAB0F82B07B1F830" box="[1376,1435,1970,1994]" pageId="8" pageNumber="511" rank="species">sp. n.</taxonomicNameLabel>
32—
</emphasis>
<taxonomicName id="4C6B4D4BFFD59319FF1CF84A03D5F810" box="[204,511,2003,2026]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="511" phylum="Arthropoda" rank="subSpecies" species="aquilum" status="subsp. nov." subSpecies="occidentalis">
<emphasis id="B91FEADAFFD59319FF1CF84A03D5F810" box="[204,511,2003,2026]" italics="true" pageId="8" pageNumber="511">Calisto aquilum occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD59319FDD5F84B0061F810" box="[517,587,2002,2026]" pageId="8" pageNumber="511" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF146640FFD49318FF47F8870032F82F" httpUri="https://zenodo.org/record/249428/files/figure.png" pageId="9" pageNumber="512" targetBox="[326,1258,201,1784]" targetPageId="9">
<paragraph id="8BD436C8FFD49318FF47F8870032F82F" blockId="9.[151,1437,1822,2005]" pageId="9" pageNumber="512">
<emphasis id="B91FEADAFFD49318FF47F8870375F8CC" bold="true" box="[151,351,1822,1846]" pageId="9" pageNumber="512">FIGURES 33–36.</emphasis>
Geographical distribution of new species of Cuban 
<taxonomicName id="4C6B4D4BFFD49318FC43F88601CAF8CC" box="[915,992,1823,1846]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEADAFFD49318FC43F88601CAF8CC" box="[915,992,1823,1846]" italics="true" pageId="9" pageNumber="512">Calisto</emphasis>
</taxonomicName>
. 
<emphasis id="B91FEADAFFD49318FC3EF8870609F8CC" bold="true" box="[1006,1059,1822,1846]" pageId="9" pageNumber="512">33—</emphasis>
Map of Cuba showing areas where the new taxa were found. 
<emphasis id="B91FEADAFFD49318FE66F8A703C1F8AC" bold="true" box="[438,491,1854,1878]" pageId="9" pageNumber="512">34—</emphasis>
Middle rectangle in figure 33, localities where 
<taxonomicName id="4C6B4D4BFFD49318FC20F8A606A8F8AC" box="[1008,1154,1855,1878]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="512" phylum="Arthropoda" rank="species" species="torrei" status="sp. nov.">
<emphasis id="B91FEADAFFD49318FC20F8A606A8F8AC" box="[1008,1154,1855,1878]" italics="true" pageId="9" pageNumber="512">Calisto torrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD49318FB5BF8A706E3F8AC" box="[1163,1225,1854,1878]" pageId="9" pageNumber="512" rank="species">
<emphasis id="B91FEADAFFD49318FB5BF8A706E3F8AC" bold="true" box="[1163,1225,1854,1878]" pageId="9" pageNumber="512">sp. n.</emphasis>
</taxonomicNameLabel>
(black circles) and 
<taxonomicName id="4C6B4D4BFFD49318FF47F8C6036CF88C" box="[151,326,1887,1910]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="512" phylum="Arthropoda" rank="species" species="aquilum" status="sp. nov.">
<emphasis id="B91FEADAFFD49318FF47F8C6036CF88C" box="[151,326,1887,1910]" italics="true" pageId="9" pageNumber="512">Calisto aquilum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD49318FE82F8C703B8F88C" box="[338,402,1886,1910]" pageId="9" pageNumber="512" rank="species">
<emphasis id="B91FEADAFFD49318FE82F8C703B8F88C" bold="true" box="[338,402,1886,1910]" pageId="9" pageNumber="512">sp. n.</emphasis>
</taxonomicNameLabel>
(white stars) have been found in the Guamuhaya massif. 
<emphasis id="B91FEADAFFD49318FBF7F8C70676F88C" bold="true" box="[1063,1116,1886,1910]" pageId="9" pageNumber="512">35—</emphasis>
Rigth rectangle in figure 33, localities where 
<taxonomicName id="4C6B4D4BFFD49318FE9BF8E6001AF86C" box="[331,560,1919,1942]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="512" phylum="Arthropoda" rank="species" species="dissimulatum" status="sp. nov.">
<emphasis id="B91FEADAFFD49318FE9BF8E6001AF86C" box="[331,560,1919,1942]" italics="true" pageId="9" pageNumber="512">Calisto dissimulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57A1FFD49318FDEAF8E70052F86C" box="[570,632,1918,1942]" pageId="9" pageNumber="512" rank="species">
<emphasis id="B91FEADAFFD49318FDEAF8E70052F86C" bold="true" box="[570,632,1918,1942]" pageId="9" pageNumber="512">sp. n.</emphasis>
</taxonomicNameLabel>
has been found in the Sierra Maestra and Nipe–Sagua–Baracoa massifs. 
<emphasis id="B91FEADAFFD49318FF47F80702E6F84C" bold="true" box="[151,204,1950,1974]" pageId="9" pageNumber="512">36—</emphasis>
Left rectangle in figure 33, localities where 
<emphasis id="B91FEADAFFD49318FD6FF80601DAF84C" box="[703,1008,1951,1974]" italics="true" pageId="9" pageNumber="512">Calistoaquilum occidentalis</emphasis>
<taxonomicNameLabel id="A22C57A1FFD49318FC2BF807066FF84C" box="[1019,1093,1950,1974]" pageId="9" pageNumber="512" rank="subSpecies">
<emphasis id="B91FEADAFFD49318FC2BF807066FF84C" bold="true" box="[1019,1093,1950,1974]" pageId="9" pageNumber="512">ssp. n.</emphasis>
</taxonomicNameLabel>
has been found in the middle portion of the Guaniguanico massif.
</paragraph>
</caption>
<paragraph id="8BD436C8FFD7931BFF17FF0303A0FF2D" blockId="10.[151,1437,151,1007]" pageId="10" pageNumber="513">
The adults emerged between 9:30 am and 4:00 pm from 
<date id="FFD51008FFD7931BFC97FF0201ECFF5F" box="[839,966,151,178]" pageId="10" pageNumber="513">June 6–10th</date>
after 9–12 days (n=12). Total developing time was 61–71 days.
</paragraph>
</subSubSection>
<subSubSection id="C3716543FFD7931BFF17FF7A00EDFC15" pageId="10" pageNumber="513" type="discussion">
<paragraph id="8BD436C8FFD7931BFF17FF7A0196FE55" blockId="10.[151,1437,151,1007]" pageId="10" pageNumber="513">
<emphasis id="B91FEADAFFD7931BFF17FF7A0311FF01" bold="true" box="[199,315,227,251]" pageId="10" pageNumber="513">Remarks.</emphasis>
<taxonomicName id="4C6B4D4BFFD7931BFE91FF7A0004FF01" box="[321,558,227,251]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFE91FF7A0004FF01" box="[321,558,227,251]" italics="true" pageId="10" pageNumber="513">Calisto dissimulatum</emphasis>
</taxonomicName>
was another “orphan” lineage found by Núñez et al (2012). It was represented by the specimen coded as PM07-20. At that time the single available specimen led us to confuse it with 
<taxonomicName id="4C6B4D4BFFD7931BFAF6FE9107B6FEE5" box="[1318,1436,263,287]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD7931BFAF6FE9107B6FEE5" box="[1318,1436,263,287]" italics="true" pageId="10" pageNumber="513">C. brochei</emphasis>
</taxonomicName>
which exhibits a similar wing pattern. More field work showed that this taxon seems to replace 
<taxonomicName id="4C6B4D4BFFD7931BFB0EFEB2077FFEB9" box="[1246,1365,299,323]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD7931BFB0EFEB2077FFEB9" box="[1246,1365,299,323]" italics="true" pageId="10" pageNumber="513">C. brochei</emphasis>
</taxonomicName>
at the west end of Nipe–Sagua–Baracoa mountain range. The collection of more individuals, including immature stages, and the sequencing of more specimens allow its present description. It was confused with the similar 
<taxonomicName id="4C6B4D4BFFD7931BFA9BFEEA0304FE55" authority="Torre" authorityName="Torre" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="brochei">
<emphasis id="B91FEADAFFD7931BFA9BFEEA02C1FE55" italics="true" pageId="10" pageNumber="513">Calisto brochei</emphasis>
Torre
</taxonomicName>
by Fontenla &amp; Rodríguez (1990) and Núñez 
<emphasis id="B91FEADAFFD7931BFCFBFE010149FE55" box="[811,867,407,431]" italics="true" pageId="10" pageNumber="513">et al.</emphasis>
(2012).
</paragraph>
<paragraph id="8BD436C8FFD7931BFF17FE2201B6FD50" blockId="10.[151,1437,151,1007]" pageId="10" pageNumber="513">
DNA analysis showed large genetic distances of 
<taxonomicName id="4C6B4D4BFFD7931BFCD4FE220197FE29" box="[772,957,443,467]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFCD4FE220197FE29" box="[772,957,443,467]" italics="true" pageId="10" pageNumber="513">C. dissimulatum</emphasis>
</taxonomicName>
from all its Cuban relatives. The closest taxon is the following new species described herein that inhabits the central and western Cuban mountain ranges, with distance values of 4.3 and 4.7% respectively. Another close relative in the Sierra Maestra, 
<taxonomicName id="4C6B4D4BFFD7931BFB38FD9A0751FDE1" box="[1256,1403,515,539]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD7931BFB38FD9A0751FDE1" box="[1256,1403,515,539]" italics="true" pageId="10" pageNumber="513">C. smintheus</emphasis>
</taxonomicName>
(
<specimenCount id="9D6DFD41FFD7931BFA5DFD9B0325FDC5" pageId="10" pageNumber="513" type="generic">4 specimens</specimenCount>
, 3 localities, with 0.19% intraspecific variation), is separated by 6.2%. From sympatric 
<taxonomicName id="4C6B4D4BFFD7931BFAD4FDB10751FDC5" box="[1284,1403,551,575]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="bruneri">
<emphasis id="B91FEADAFFD7931BFAD4FDB10751FDC5" box="[1284,1403,551,575]" italics="true" pageId="10" pageNumber="513">C. bruneri</emphasis>
</taxonomicName>
(
<specimenCount id="9D6DFD41FFD7931BFA5DFDBE0325FD98" pageId="10" pageNumber="513" type="generic">4 specimens</specimenCount>
, 3 localities, with 0.37% intraspecific variation), 
<taxonomicName id="4C6B4D4BFFD7931BFC88FDD201C2FD99" box="[856,1000,587,611]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="herophile">
<emphasis id="B91FEADAFFD7931BFC88FDD201C2FD99" box="[856,1000,587,611]" italics="true" pageId="10" pageNumber="513">C. herophile</emphasis>
</taxonomicName>
(
<specimenCount id="9D6DFD41FFD7931BFC2CFDD306B7FD98" box="[1020,1181,586,610]" pageId="10" pageNumber="513" type="generic">10 specimens</specimenCount>
, 7 localities, 0.8% of intraspecific variation), and 
<taxonomicName id="4C6B4D4BFFD7931BFE0BFDE9007AFD7D" box="[475,592,623,647]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="occulta">
<emphasis id="B91FEADAFFD7931BFE0BFDE9007AFD7D" box="[475,592,623,647]" italics="true" pageId="10" pageNumber="513">C. occulta</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C6B4D4BFFD7931BFDB1FDE90130FD7D" box="[609,794,623,647]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFDB1FDE90130FD7D" box="[609,794,623,647]" italics="true" pageId="10" pageNumber="513">C. dissimulatum</emphasis>
</taxonomicName>
differs by 8.3, 7.9 and 7.7%, respectively. Intraspecific variation within 
<taxonomicName id="4C6B4D4BFFD7931BFE80FD0A002FFD51" box="[336,517,659,683]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFE80FD0A002FFD51" box="[336,517,659,683]" italics="true" pageId="10" pageNumber="513">C. dissimulatum</emphasis>
</taxonomicName>
was 0.1% (
<specimenCount id="9D6DFD41FFD7931BFD5AFD0B013CFD50" box="[650,790,658,682]" pageId="10" pageNumber="513" type="generic">3 specimens</specimenCount>
, 1 locality).
</paragraph>
<paragraph id="8BD436C8FFD7931BFF17FD2103DBFCC0" blockId="10.[151,1437,151,1007]" pageId="10" pageNumber="513">
Recent specimens from localities other than La Zoilita at Sierra Cristal were not available for DNA extraction. Since most known Cuban taxa, excepting 
<taxonomicName id="4C6B4D4BFFD7931BFDBCFD4200DCFD09" box="[620,758,731,755]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="herophile">
<emphasis id="B91FEADAFFD7931BFDBCFD4200DCFD09" box="[620,758,731,755]" italics="true" pageId="10" pageNumber="513">C. herophile</emphasis>
</taxonomicName>
and the next new species, are restricted to a single mountain range the assignment of other populations to 
<taxonomicName id="4C6B4D4BFFD7931BFD49FC99017AFCED" box="[665,848,767,791]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFD49FC99017AFCED" box="[665,848,767,791]" italics="true" pageId="10" pageNumber="513">C. dissimulatum</emphasis>
</taxonomicName>
must be confirmed, mainly for those inhabiting the western half of Sierra Maestra.
</paragraph>
<paragraph id="8BD436C8FFD7931BFF17FCDE00EDFC15" blockId="10.[151,1437,151,1007]" pageId="10" pageNumber="513">
The immature stages of 
<taxonomicName id="4C6B4D4BFFD7931BFE09FCD100BAFCA5" box="[473,656,839,863]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFE09FCD100BAFCA5" box="[473,656,839,863]" italics="true" pageId="10" pageNumber="513">C. dissimulatum</emphasis>
</taxonomicName>
also showed some diagnostic features. Last larval instar is similar to that 
<taxonomicName id="4C6B4D4BFFD7931BFF18FCF20372FC79" box="[200,344,875,899]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD7931BFF18FCF20372FC79" box="[200,344,875,899]" italics="true" pageId="10" pageNumber="513">C. smintheus</emphasis>
</taxonomicName>
but all markings are larger resulting in an overall darker pattern. The majorities of head markings present at 
<taxonomicName id="4C6B4D4BFFD7931BFEDCFC0903E8FC5D" box="[268,450,911,935]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="dissimulatum">
<emphasis id="B91FEADAFFD7931BFEDCFC0903E8FC5D" box="[268,450,911,935]" italics="true" pageId="10" pageNumber="513">C. dissimulatum</emphasis>
</taxonomicName>
last instar are reduced or absent in 
<taxonomicName id="4C6B4D4BFFD7931BFC87FC0901C2FC5D" box="[855,1000,911,935]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD7931BFC87FC0901C2FC5D" box="[855,1000,911,935]" italics="true" pageId="10" pageNumber="513">C. smintheus</emphasis>
</taxonomicName>
including the vertical lateral lines, the epicraneum X mark, and transverse band of frons. Among all other known taxa, the heavily spotted pattern of pupa is only shared by the closely related 
<taxonomicName id="4C6B4D4BFFD7931BFDFFFC4100EAFC15" box="[559,704,983,1007]" class="Insecta" family="Nymphalidae" genus="Calisto" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="513" phylum="Arthropoda" rank="species" species="smintheus">
<emphasis id="B91FEADAFFD7931BFDFFFC4100EAFC15" box="[559,704,983,1007]" italics="true" pageId="10" pageNumber="513">C. smintheus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>