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<document id="CF3B97586B6ABC4266A1AF51D0D7DBE9" ID-DOI="10.5852/ejt.2020.618" ID-GBIF-Dataset="eb3e0d4d-e623-4c58-ab31-9833367933bd" ID-Zenodo-Dep="3722950" ID-ZooBank="FB1B58E6-7E40-4C16-9DFF-2EA5D43BC0B3" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584790424389" checkinUser="plazi" docAuthor="Vivallo, Felipe" docDate="2020" docId="03C287D4873CAA2C4448FC5CDE28A86E" docLanguage="en" docName="ejt-618_vivallo.pdf.imf" docOrigin="European Journal of Taxonomy 618" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Centris escomeli Cockerell 1926" docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="FFFBFFAC871BAA04461DFFF2DB60AA25" masterDocTitle="The bees of the genus Centris Fabricius, 1804 described by Theodore Dru Alison Cockerell (Hymenoptera: Apidae)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="40" updateTime="1698730621050" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="74CB4521463C3DE4977B2440ED598C47">The bees of the genus Centris Fabricius, 1804 described by Theodore Dru Alison Cockerell (Hymenoptera: Apidae)</mods:title>
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<mods:title id="88EBE8F1E75147E85E164A3B9967BFB2">European Journal of Taxonomy</mods:title>
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<mods:date id="0F34CF2B756E52095FEC48CAD26AA711">2020</mods:date>
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<treatment id="03C287D4873CAA2C4448FC5CDE28A86E" ID-DOI="http://doi.org/10.5281/zenodo.3730325" ID-GBIF-Taxon="162658477" ID-Zenodo-Dep="3730325" LSID="urn:lsid:plazi:treatment:03C287D4873CAA2C4448FC5CDE28A86E" httpUri="http://treatment.plazi.org/id/03C287D4873CAA2C4448FC5CDE28A86E" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection id="C3716549873CAA234448FC5CD8BFA9ED" box="[597,991,942,969]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8BD436C2873CAA234448FC5CD8BFA9ED" blockId="39.[597,991,942,1007]" box="[597,991,942,969]" pageId="39" pageNumber="40">
<heading id="D09C81AE873CAA234448FC5CD8BFA9ED" box="[597,991,942,969]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="40" reason="2">
<taxonomicName id="4C6B4D41873CAA234448FC5CD8BFA9ED" ID-CoL="S97W" authority="Cockerell, 1926" authorityName="Cockerell" authorityYear="1926" box="[597,991,942,969]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="escomeli">
<emphasis id="B91FEAD0873CAA234448FC5CD87CA9EC" bold="true" box="[597,796,942,969]" italics="true" pageId="39" pageNumber="40">Centris escomeli</emphasis>
<bibRefCitation id="EFFA4B33873CAA23453EFC5CD8BFA9ED" author="Cockerell, T. D. A." box="[803,991,942,968]" journalOrPublisher="Annals and Magazine of Natural History" pageId="39" pageNumber="40" pagination="214 - 226" part="17" refId="ref20022" refString="Cockerell, T. D. A. 1926 a. XXVIII. - Descriptions and records of bees. - CVIII. Annals and Magazine of Natural History 17 (98): 214 - 226. https: // doi. org / 10.1080 / 00222932608633395" title="XXVIII. - Descriptions and records of bees. - CVIII" type="journal article" year="1926">Cockerell, 1926</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3716549873CAA2344EDFC26D823A9CA" box="[752,835,980,1007]" pageId="39" pageNumber="40" type="description">
<paragraph id="8BD436C2873CAA2344EDFC26D823A9CA" blockId="39.[597,991,942,1007]" box="[752,835,980,1007]" pageId="39" pageNumber="40">
<figureCitation id="13502A47873CAA2344EDFC26D823A9CA" box="[752,835,980,1007]" captionStart="Fig" captionStartId="39.[189,232,1965,1991]" captionTargetBox="[189,1398,1470,1925]" captionTargetId="figure@39.[189,1398,1470,1925]" captionTargetPageId="39" captionText="Fig. 29. Centris escomeli Cockerell, 1926, holotype, ♂ (NMNH; USNM ENT 00534193). A. Frontal view. B. Habitus, lateral view. Scale bars: A = 1 mm; B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3723006" httpUri="https://zenodo.org/record/3723006/files/figure.png" pageId="39" pageNumber="40">Fig. 29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3716549873CAA2346A0FBE8D9D3AE16" box="[189,691,1049,1076]" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="8BD436C2873CAA2346A0FBE8D9D3AE16" blockId="39.[189,691,1049,1144]" box="[189,691,1049,1076]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="AB7B11EC873CAA2346A0FBE8D9D3AE16" box="[189,691,1049,1076]" pageId="39" pageNumber="40">
<treatmentCitation id="0ACA10D3873CAA2346A0FBE8D9D0AE16" author="Cockerell T. D. A." box="[189,688,1049,1076]" page="29" pageId="39" pageNumber="40" year="1926">
<taxonomicName id="4C6B4D41873CAA2346A0FBE8D9D0AE16" ID-CoL="S97W" authority="Cockerell, 1926 b: 28 - 29" authorityName="Cockerell" authorityPageNumber="28 - 29" authorityYear="1926" box="[189,688,1049,1076]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="escomeli">
<emphasis id="B91FEAD0873CAA2346A0FBE8DAE2AE16" box="[189,386,1049,1076]" italics="true" pageId="39" pageNumber="40">Centris escomeli</emphasis>
<bibRefCitation id="EFFA4B33873CAA234794FBEBD9D0AE16" author="Cockerell" box="[393,688,1049,1075]" firstAuthor="Cockerell" pageId="39" pageNumber="40" pagination="28 - 29" refId="ref20070" refString="Cockerell T. D. A. 1926 b. The black bees of Peru. The Entomologist 59: 28 - 29." type="journal article" year="1926" yearSuffix="b">Cockerell, 1926b: 2829</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716549873CAA2C46A0FBACDA16ABE8" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BD436C2873CAA2346A0FBACDA59AE5D" blockId="39.[189,691,1049,1144]" box="[189,313,1118,1144]" pageId="39" pageNumber="40">
<emphasis id="B91FEAD0873CAA2346A0FBACDA59AE5D" bold="true" box="[189,313,1118,1144]" pageId="39" pageNumber="40">
<typeStatus id="54D08860873CAA2346A0FBACDB9AAE5D" box="[189,250,1118,1144]" pageId="39" pageNumber="40">Type</typeStatus>
data
</emphasis>
</paragraph>
<paragraph id="8BD436C2873CAA2C46A0FB62DE16AB75" blockId="39.[189,1399,1168,1425]" lastBlockId="40.[189,1399,267,336]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
This species was described based on a couple of female and male specimens collected in
<collectingRegion id="49AFF820873CAA23431FFB62DE12AE8F" box="[1282,1394,1168,1194]" country="Peru" name="Arequipa" pageId="39" pageNumber="40">Arequipa</collectingRegion>
, southern
<collectingCountry id="F37C7652873CAA234736FB45DA05AEF4" box="[299,357,1207,1233]" name="Peru" pageId="39" pageNumber="40">Peru</collectingCountry>
. Both specimens were collected by Edmundo Escomel (
<date id="FFD51002873CAA234219FB44DFEEAEF4" box="[1028,1166,1206,1233]" pageId="39" pageNumber="40" value="1880" valueMax="1959">18801959</date>
), a Peruvian doctor who dedicated his free time as a meticulous observer of the flora, fauna and the natural environment (Velarde 2006). The male
<typeStatus id="54D08860873CAA2347E7FAF1D901AF38" box="[506,609,1283,1309]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
(
<figureCitation id="13502A47873CAA23446EFAF1D9ABAF38" box="[627,715,1283,1309]" captionStart="Fig" captionStartId="39.[189,232,1965,1991]" captionTargetBox="[189,1398,1470,1925]" captionTargetId="figure@39.[189,1398,1470,1925]" captionTargetPageId="39" captionText="Fig. 29. Centris escomeli Cockerell, 1926, holotype, ♂ (NMNH; USNM ENT 00534193). A. Frontal view. B. Habitus, lateral view. Scale bars: A = 1 mm; B = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3723006" httpUri="https://zenodo.org/record/3723006/files/figure.png" pageId="39" pageNumber="40">Fig. 29</figureCitation>
) and the female
<typeStatus id="54D08860873CAA234585FAF6D89DAF3B" box="[920,1021,1284,1310]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
are housed at the
<collectionCode id="ED7AAE07873CAA2342C2FAF6DE5DAF3B" box="[1247,1341,1284,1310]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">NMNH</collectionCode>
. The
<typeStatus id="54D08860873CAA2346A0FADBDA44AF66" box="[189,292,1321,1347]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
has the following data label:
<collectingRegion id="49AFF820873CAA23449AFADBD994AF66" box="[647,756,1321,1347]" country="Peru" name="Arequipa" pageId="39" pageNumber="40">Arequipa</collectingRegion>
<collectingCountry id="F37C7652873CAA2344E0FAD8D853AF61" box="[765,819,1322,1348]" name="Peru" pageId="39" pageNumber="40">Peru</collectingCountry>
(Escomel) [handwritten]\
<taxonomicName id="4C6B4D41873CAA234273FAD8DE16AF61" authority="Ckll." authorityName="Cockerell" authorityYear="1926" box="[1134,1398,1321,1348]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="escomeli">
<emphasis id="B91FEAD0873CAA234273FAD8DE54AF66" box="[1134,1332,1321,1348]" italics="true" pageId="39" pageNumber="40">Centris escomeli</emphasis>
Ckll.
</taxonomicName>
Type [handwritten]\ [red label] Type No. [printed] 40493 [handwritten]
<collectingCountry id="F37C7652873CAA23423BFAA2DF38AF4F" box="[1062,1112,1360,1386]" name="United States of America" pageId="39" pageNumber="40">U.S.</collectingCountry>
N.M. [printed]\ [yellow label] USNM ENT 00534193 [barcode] [printed]\\ DO NOT REMOVE SI DB Reference Not a property tag T. Schultz
<collectionCode id="ED7AAE078733AA2C476EFEFEDAADAB03" box="[371,461,268,294]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="40" pageNumber="41" type="Museum">NMNH</collectionCode>
[printed]. The female
<typeStatus id="54D088608733AA2C44FAFEFED82CAB03" box="[743,844,268,294]" pageId="40" pageNumber="41" type="paratype">paratype</typeStatus>
has the following data label:
<collectingRegion id="49AFF8208733AA2C42DDFEF9DE4DAB00" box="[1216,1325,267,293]" country="Peru" name="Arequipa" pageId="40" pageNumber="41">Arequipa</collectingRegion>
<collectingCountry id="F37C76528733AA2C4324FEFEDE13AB03" box="[1337,1395,268,294]" name="Peru" pageId="40" pageNumber="41">Peru</collectingCountry>
. (Escomel) [handwritten]\
<taxonomicName id="4C6B4D418733AA2C47F4FEC4D98BAB75" authority="Ckll." authorityName="Cockerell" authorityYear="1926" box="[489,747,309,336]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="escomeli">
<emphasis id="B91FEAD08733AA2C47F4FEC4D9CCAB6A" box="[489,684,309,336]" italics="true" pageId="40" pageNumber="41">Centris escomeli</emphasis>
Ckll.
</taxonomicName>
cotype
<specimenCount id="9D6DFD4B8733AA2C4558FEC4D83BAB75" box="[837,859,310,336]" pageId="40" pageNumber="41" type="female"></specimenCount>
[handwritten]\ Probably is a (
<emphasis id="B91FEAD08733AA2C42AEFEC7DE05AB6A" box="[1203,1381,309,335]" italics="true" pageId="40" pageNumber="41">Wagenknechtia</emphasis>
).
</paragraph>
<caption id="DF14664A873CAA2346A0F85FD8CFADCE" ID-DOI="http://doi.org/10.5281/zenodo.3723006" ID-Zenodo-Dep="3723006" httpUri="https://zenodo.org/record/3723006/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1965,1991]" targetBox="[189,1398,1470,1925]" targetPageId="39">
<paragraph id="8BD436C2873CAA2346A0F85FD8CFADCE" blockId="39.[189,1399,1965,2027]" pageId="39" pageNumber="40">
<emphasis id="B91FEAD0873CAA2346A0F85FDA7BADE2" bold="true" box="[189,283,1965,1991]" pageId="39" pageNumber="40">Fig. 29.</emphasis>
<taxonomicName id="4C6B4D41873CAA234739F85FD9D3ADE2" authority="Cockerell, 1926" authorityName="Cockerell" authorityYear="1926" box="[292,691,1965,1991]" class="Insecta" family="Apidae" genus="Centris" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="escomeli">
<emphasis id="B91FEAD0873CAA234739F85FDA8BADE2" box="[292,491,1965,1991]" italics="true" pageId="39" pageNumber="40">Centris escomeli</emphasis>
<bibRefCitation id="EFFA4B33873CAA2347E9F85FD9D3ADE2" author="Cockerell, T. D. A." box="[500,691,1965,1991]" journalOrPublisher="Annals and Magazine of Natural History" pageId="39" pageNumber="40" pagination="214 - 226" part="17" refId="ref20022" refString="Cockerell, T. D. A. 1926 a. XXVIII. - Descriptions and records of bees. - CVIII. Annals and Magazine of Natural History 17 (98): 214 - 226. https: // doi. org / 10.1080 / 00222932608633395" title="XXVIII. - Descriptions and records of bees. - CVIII" type="journal article" year="1926">Cockerell, 1926</bibRefCitation>
</taxonomicName>
, holotype, ♂ (
<collectionCode id="ED7AAE07873CAA23457DF85FD8DFADE2" box="[864,959,1965,1991]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">NMNH</collectionCode>
; USNM ENT 00534193).
<emphasis id="B91FEAD0873CAA2342E1F85FDE71ADE2" bold="true" box="[1276,1297,1965,1991]" pageId="39" pageNumber="40">A</emphasis>
. Frontal view.
<emphasis id="B91FEAD0873CAA23471EF823DA77ADCE" bold="true" box="[259,279,2001,2027]" pageId="39" pageNumber="40">B</emphasis>
. Habitus, lateral view. Scale bars: A = 1 mm; B = 2 mm.
</paragraph>
</caption>
<paragraph id="8BD436C28733AA2C46A0FE8CDA3DABBD" blockId="40.[189,374,382,462]" box="[189,349,382,408]" pageId="40" pageNumber="41">
<heading id="D09C81AE8733AA2C46A0FE8CDA3DABBD" bold="true" box="[189,349,382,408]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B91FEAD08733AA2C46A0FE8CDA3DABBD" bold="true" box="[189,349,382,408]" pageId="40" pageNumber="41">
<typeStatus id="54D088608733AA2C46A0FE8CDB9AABBD" box="[189,250,382,408]" pageId="40" pageNumber="41">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD436C28733AA2C46A0FE46DA16ABE8" blockId="40.[189,374,382,462]" box="[189,374,435,462]" pageId="40" pageNumber="41">
<collectingCountry id="F37C76528733AA2C46A0FE46DB97ABEB" box="[189,247,436,462]" name="Peru" pageId="40" pageNumber="41">Peru</collectingCountry>
:
<collectingRegion id="49AFF8208733AA2C471CFE41DA12ABE8" box="[257,370,435,461]" country="Peru" name="Arequipa" pageId="40" pageNumber="41">Arequipa</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37165498733AA2C46A0FE09DE28A86E" box="[189,1352,507,587]" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BD436C28733AA2C46A0FE09DA59A830" blockId="40.[189,1352,507,587]" box="[189,313,507,533]" pageId="40" pageNumber="41">
<heading id="D09C81AE8733AA2C46A0FE09DA59A830" bold="true" box="[189,313,507,533]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="B91FEAD08733AA2C46A0FE09DA59A830" bold="true" box="[189,313,507,533]" pageId="40" pageNumber="41">Comment</emphasis>
</heading>
</paragraph>
<paragraph id="8BD436C28733AA2C46A0FDC2DE28A86E" blockId="40.[189,1352,507,587]" box="[189,1352,560,587]" pageId="40" pageNumber="41">
This species was correctly interpreted as a valid species by
<bibRefCitation id="EFFA4B338733AA2C456BFDC3DF3CA86E" author="Moure" box="[886,1116,560,587]" etAl="et al." firstAuthor="Moure" pageId="40" pageNumber="41" pagination="83 - 142" refId="ref21502" refString="Moure J. S., Melo G. A. R. &amp; Vivallo F. 2007. Centridini Cockerell &amp; Cockerell. In: Moure J. S., Urban D. &amp; Melo G. A. R. (orgs) Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical Region: 83 - 142. Sociedade Brasileira de Entomologia, Curitibrazil." type="book chapter" year="2007">
Moure
<emphasis id="B91FEAD08733AA2C45D1FDC3DF67A86F" box="[972,1031,560,586]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
(2007)
</bibRefCitation>
and
<bibRefCitation id="EFFA4B338733AA2C4289FDC2DE24A86E" author="Vivallo" box="[1172,1348,560,587]" firstAuthor="Vivallo" pageId="40" pageNumber="41" pagination="501 - 537" refId="ref22300" refString="Vivallo F. 2013. Revision of the bee subgenus Centris (Wagenknechtia) Moure, 1950 (Hymenoptera: Apidae: Centridini). Zootaxa 3683 (5): 501 - 537. https: // doi. org / 10.11646 / zootaxa. 3683.5.1" type="journal article" year="2013">Vivallo (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>