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<document id="B9B788D33C9FB70199A0711478C1EE33" ID-DOI="10.5281/zenodo.5348151" ID-ISSN="2345-7600" ID-Zenodo-Dep="5348151" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630203888444" checkinUser="tatiana" docAuthor="Polhemus, Dan A. &amp; Polhemus, John T." docDate="2012" docId="03C287CFFFB2C815FF31FC5FFB0EFB75" docLanguage="en" docName="60rbz343-359.pdf" docOrigin="Raffles Bulletin of Zoology 60 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Ochteridae Kirkaldy 1906" docType="treatment" docVersion="4" lastPageNumber="344" masterDocId="FFFBFFB7FFB3C814FFC0FF82FFDDFFE0" masterDocTitle="Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Ix. Infraorder Nepomorpha, Families Ochteridae And Gelastocoridae" masterLastPageNumber="359" masterPageNumber="343" pageNumber="344" updateTime="1699183961091" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8BA4AB9DE98E9BFAF39501087FD98200">Guide To The Aquatic Heteroptera Of Singapore And Peninsular Malaysia. Ix. Infraorder Nepomorpha, Families Ochteridae And Gelastocoridae</mods:title>
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<mods:namePart id="5C2EF033FF47E898969014A0C8031AE6">Polhemus, Dan A.</mods:namePart>
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<subSubSection id="C3716552FFB2C815FF31FC5FFDADFC15" box="[241,624,989,1013]" pageId="1" pageNumber="344" type="nomenclature">
<paragraph id="8BD436D9FFB2C815FF31FC5FFDADFC15" blockId="1.[241,624,989,1013]" box="[241,624,989,1013]" pageId="1" pageNumber="344">
<heading id="D09C81B5FFB2C815FF31FC5FFDADFC15" bold="true" box="[241,624,989,1013]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="344" reason="2">
<emphasis id="B91FEACBFFB2C815FF31FC5FFDADFC15" bold="true" box="[241,624,989,1013]" pageId="1" pageNumber="344">
Family
<taxonomicName id="4C6B4D5AFFB2C815FE89FC5FFDADFC15" ID-CoL="DGZ" authority="Kirkaldy" authorityName="Kirkaldy" authorityYear="1906" box="[329,624,989,1013]" class="Insecta" family="Ochteridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="family">OCHTERIDAE Kirkaldy</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3716552FFB2C815FFB1FB9FFB0EFB75" pageId="1" pageNumber="344" type="discussion">
<paragraph id="8BD436D9FFB2C815FFB1FB9FFE7DFA15" blockId="1.[113,752,1053,1525]" pageId="1" pageNumber="344">
<emphasis id="B91FEACBFFB2C815FFB1FB9FFF27FBD5" bold="true" box="[113,250,1053,1077]" pageId="1" pageNumber="344">
<emphasis id="B91FEACBFFB2C815FFB1FB9FFF2FFBD5" bold="true" box="[113,242,1053,1077]" italics="true" pageId="1" pageNumber="344">Discussion</emphasis>
.
</emphasis>
— The
<taxonomicName id="4C6B4D5AFFB2C815FEA3FB9FFE39FBD5" authorityName="Kirkaldy" authorityYear="1906" box="[355,484,1053,1077]" class="Insecta" family="Ochteridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="family">Ochteridae</taxonomicName>
, or velvet shore bugs, comprise a small family containing three extant genera: the widespread
<taxonomicName id="4C6B4D5AFFB2C815FEDAFBDCFEA1FB95" authorityName="Latreille" authorityYear="1807" box="[282,380,1118,1141]" class="Insecta" family="Ochteridae" genus="Ochterus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEACBFFB2C815FEDAFBDCFEA1FB95" box="[282,380,1118,1141]" italics="true" pageId="1" pageNumber="344">Ochterus</emphasis>
</taxonomicName>
, which is pantropical and the only genus occurring in
<collectingCountry id="F37C7649FFB2C815FE93FBFFFE19FB75" box="[339,452,1149,1173]" name="Singapore" pageId="1" pageNumber="344">Singapore</collectingCountry>
and Peninsular
<collectingCountry id="F37C7649FFB2C815FD42FBFFFD36FB75" box="[642,747,1149,1173]" name="Malaysia" pageId="1" pageNumber="344">Malaysia</collectingCountry>
;
<emphasis id="B91FEACBFFB2C815FFB1FB1CFECCFB55" box="[113,273,1182,1205]" italics="true" pageId="1" pageNumber="344">Megaochterus</emphasis>
, endemic to
<collectingCountry id="F37C7649FFB2C815FE6DFB1FFDC5FB55" box="[429,536,1181,1205]" name="Australia" pageId="1" pageNumber="344">Australia</collectingCountry>
; and
<taxonomicName id="4C6B4D5AFFB2C815FD9BFB1CFD34FB55" authorityName="Drake &amp; Gomez-Menor" authorityYear="1954" box="[603,745,1182,1205]" class="Insecta" family="Ochteridae" genus="Ocyochterus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEACBFFB2C815FD9BFB1CFD34FB55" box="[603,745,1182,1205]" italics="true" pageId="1" pageNumber="344">Ocyochterus</emphasis>
</taxonomicName>
, confined to the Neotropical region.As a whole, the
<taxonomicName id="4C6B4D5AFFB2C815FDBDFB3FFD2DFB35" authorityName="Kirkaldy" authorityYear="1906" box="[637,752,1213,1237]" class="Insecta" family="Ochteridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="family">Ochteridae</taxonomicName>
exhibit two major geographic foci of species radiation, in South America (
<bibRefCitation id="EFFA4B28FFB2C815FEDEFB7FFE78FAF5" author="Schell, D" box="[286,421,1277,1301]" pageId="1" pageNumber="344" pagination="29 - 47" refId="ref13944" refString="Schell, D. V., 1943. The Ochteridae of the Western Hemisphere. Journal of the Kansas Entomological Society, 16: 29 - 47." type="journal article" year="1943">Schell, 1943</bibRefCitation>
) and Melanesia (
<bibRefCitation id="EFFA4B28FFB2C815FD9BFB7FFD31FAF5" author="Baehr, M" box="[603,748,1277,1301]" pageId="1" pageNumber="344" pagination="91 - 93" refId="ref12800" refString="Baehr, M., 1990 a. Ochterus noualhieri sp. n., a new ochterid from Java [Heteroptera, Ochteridae]. Revue Francaise d'Entomologie (N. S.), 12: 91 - 93." type="journal article" year="1990">Baehr, 1990a</bibRefCitation>
,
<bibRefCitation id="EFFA4B28FFB2C815FFB1FA9FFF6AFAD5" author="Baehr, M" box="[113,183,1309,1333]" pageId="1" pageNumber="344" pagination="449 - 477" refId="ref12842" refString="Baehr, M., 1990 b. Revision of the genus Ochterus Latreille in the Australian Region (Heteroptera: Ochteridae). Entomologica Scandinavica, 20: 449 - 477." type="journal article" year="1990">1990b</bibRefCitation>
;
<bibRefCitation id="EFFA4B28FFB2C815FF05FA9FFEACFAD5" author="Kormilev, N" box="[197,369,1309,1333]" pageId="1" pageNumber="344" pagination="429 - 444" refId="ref13296" refString="Kormilev, N. A., 1971. Ochteridae from the Oriental and Australian Regions. Pacific Insects, 13: 429 - 444." type="journal article" year="1971">Kormilev, 1971</bibRefCitation>
,
<bibRefCitation id="EFFA4B28FFB2C815FEBFFA9FFE6BFAD5" author="Kormilev, N" box="[383,438,1309,1333]" pageId="1" pageNumber="344" pagination="1 - 9" refId="ref13322" refString="Kormilev, N. A., 1973. Ochteridae from Western and Southern Africa (Hemiptera: Heteroptera). Occasional Papers of the California Academy of Sciences, 106: 1 - 9." type="journal article" year="1973">1973</bibRefCitation>
) respectively. In the Eastern Hemisphere, species richness diminishes markedly west of Huxleys Line (
<bibRefCitation id="EFFA4B28FFB2C815FE88FADFFE00FA95" author="Schell, D" box="[328,477,1373,1397]" pageId="1" pageNumber="344" pagination="29 - 47" refId="ref13944" refString="Schell, D. V., 1943. The Ochteridae of the Western Hemisphere. Journal of the Kansas Entomological Society, 16: 29 - 47." type="journal article" year="1943">Schell, 1943</bibRefCitation>
;
<bibRefCitation id="EFFA4B28FFB2C815FE2CFADFFD7CFA95" author="Kormilev, N" box="[492,673,1373,1397]" pageId="1" pageNumber="344" pagination="429 - 444" refId="ref13296" refString="Kormilev, N. A., 1971. Ochteridae from the Oriental and Australian Regions. Pacific Insects, 13: 429 - 444." type="journal article" year="1971">Kormilev, 1971</bibRefCitation>
,
<bibRefCitation id="EFFA4B28FFB2C815FD71FADFFD36FA95" author="Kormilev, N" box="[689,747,1373,1397]" pageId="1" pageNumber="344" pagination="1 - 9" refId="ref13322" refString="Kormilev, N. A., 1973. Ochteridae from Western and Southern Africa (Hemiptera: Heteroptera). Occasional Papers of the California Academy of Sciences, 106: 1 - 9." type="journal article" year="1973">1973</bibRefCitation>
; Gapud &amp; San Valentin, 1977;
<bibRefCitation id="EFFA4B28FFB2C815FE04FAFFFD88FA75" author="Gapud, V. P" box="[452,597,1405,1429]" pageId="1" pageNumber="344" pagination="300 - 309" refId="ref12969" refString="Gapud, V. P., 1981. Contribution to the taxonomy of the genus Ochterus Latreille (Hemiptera: Ochteridae). Kalikasan, Philippine Journal of Biology, 10: 300 - 309." type="journal article" year="1981">Gapud, 1981</bibRefCitation>
,
<bibRefCitation id="EFFA4B28FFB2C815FDA3FAFFFD41FA75" author="Gapud, V. P" box="[611,668,1405,1429]" pageId="1" pageNumber="344" pagination="41 - 44" refId="ref13006" refString="Gapud, V. P., 1995. A new species of Ochterus Latreille (Hemiptera: Ochteridae) from the Philippines. Asia Life Sciences, 4: 41 - 44." type="journal article" year="1995">1995</bibRefCitation>
;
<bibRefCitation id="EFFA4B28FFB2C815FD6AFAFFFF68FA55" author="Baehr, M" pageId="1" pageNumber="344" pagination="91 - 93" refId="ref12800" refString="Baehr, M., 1990 a. Ochterus noualhieri sp. n., a new ochterid from Java [Heteroptera, Ochteridae]. Revue Francaise d'Entomologie (N. S.), 12: 91 - 93." type="journal article" year="1990">Baehr, 1990a</bibRefCitation>
,
<bibRefCitation id="EFFA4B28FFB2C815FF00FA1FFEDBFA55" author="Baehr, M" box="[192,262,1437,1461]" pageId="1" pageNumber="344" pagination="449 - 477" refId="ref12842" refString="Baehr, M., 1990 b. Revision of the genus Ochterus Latreille in the Australian Region (Heteroptera: Ochteridae). Entomologica Scandinavica, 20: 449 - 477." type="journal article" year="1990">1990b</bibRefCitation>
; Nieser &amp; Chen, 1992), although the fauna of
<collectingCountry id="F37C7649FFB2C815FFB1FA3FFF00FA35" box="[113,221,1469,1493]" name="Singapore" pageId="1" pageNumber="344">Singapore</collectingCountry>
and
<collectingCountry id="F37C7649FFB2C815FED3FA3FFEA8FA35" box="[275,373,1469,1493]" name="Malaysia" pageId="1" pageNumber="344">Malaysia</collectingCountry>
is undersampled and contains new species, as described below.
</paragraph>
<paragraph id="8BD436D9FFB2C815FFB1F99FFB93FED5" blockId="1.[113,752,1565,2101]" lastBlockId="1.[812,1451,189,309]" pageId="1" pageNumber="344">
<taxonomicName id="4C6B4D5AFFB2C815FFB1F99FFF3BF9D5" authorityName="Kirkaldy" authorityYear="1906" box="[113,230,1565,1589]" class="Insecta" family="Ochteridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="family">Ochteridae</taxonomicName>
are recognised among the nepomorphan families by their large eyes, presence of ocelli, slender four-segmented antennae, transversely rugose frons and clypeus, long labium with segment III much longer than the others, slender legs with the fore femora enlarged, a tarsal formula of 2-2-3, the forewing membrane with several closed cells but no anastamosing veins, and unique genitalia with the right paramere greatly enlarged in relation to the left and bearing distal appendices. Despite being superficially similar in external appearance to members of the Leptopodomorpha, and inhabiting similar littoral and hygropetric habitats, the
<taxonomicName id="4C6B4D5AFFB2C815FFB1F8FFFF3AF875" authorityName="Kirkaldy" authorityYear="1906" box="[113,231,1917,1941]" class="Insecta" family="Ochteridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="family">Ochteridae</taxonomicName>
possesses a suite of character states typical of
<taxonomicName id="4C6B4D5AFFB2C815FFB1F81FFF23F855" box="[113,254,1949,1973]" class="Insecta" infraOrder="Nepomorpha" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="343" phylum="Arthropoda" rank="infraOrder">Nepomorpha</taxonomicName>
, including the short, ventrally inserted antennae and asymmetrical male genitalia, as well as a tubular spermatheca (Schuh &amp; Slater, 1995). The primary characters currently used for species separation are the shape of the male right paramere (
<figureCitation id="13502A5CFFB2C815FE9DF79FFE1BF7D5" box="[349,454,2077,2101]" captionStart="Figs" captionStartId="4.[136,181,1919,1941]" captionTargetBox="[138,774,649,1878]" captionTargetId="figure-8@4.[107,796,629,1898]" captionTargetPageId="4" captionText="Figs. 59. O. marginatus (Latreille), male, structural details. 5, Male right paramere, specimen from Vietnam, CL 4368.6, Superior appendage of right paramere, specimen from Vietnam, CL 4368. 7, Superior appendage of right paramere, specimen from Indonesia, Bali, CL 2170. 8, Medial process of pygophore, specimen from Vietnam, CL 4368. 9, Medial process of pygophore, specimen from Indonesia, Bali, CL 2170." figureDoi="http://doi.org/10.5281/zenodo.5348157" httpUri="https://zenodo.org/record/5348157/files/figure.png" pageId="1" pageNumber="344">Figs. 57</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FE13F79FFDC5F7D5" box="[467,536,2077,2101]" captionStart="Figs" captionStartId="5.[114,159,2006,2028]" captionTargetBox="[117,763,820,1979]" captionTargetId="figure-15@5.[95,766,802,1989]" captionTargetPageId="5" captionText="Figs. 1012. O. signatus, new species, male, structural details, specimen from Malaysia, Perak, CL 2081. 10, Male right paramere. 11, Superior appendage of right paramere. 12, Medial process of pygophore." figureDoi="http://doi.org/10.5281/zenodo.5348159" httpUri="https://zenodo.org/record/5348159/files/figure.png" pageId="1" pageNumber="344">10, 11</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FDE6F79FFDB0F7D5" box="[550,621,2077,2101]" captionStart="Figs" captionStartId="6.[833,877,2004,2026]" captionTargetBox="[830,1474,645,1971]" captionTargetId="figure-686@6.[826,1506,625,1988]" captionTargetPageId="6" captionText="Figs. 1316. O. pseudomarginatus, new species, male, structural details, specimen from Malaysia, Perak, CL 2078. 13, Male right paramere. 14, Superior appendage of right paramere. 15, Medial process of pygophore. 16, Endosoma." figureDoi="http://doi.org/10.5281/zenodo.5348161" httpUri="https://zenodo.org/record/5348161/files/figure.png" pageId="1" pageNumber="344">13, 14</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FDBAF79FFD1CF7D5" box="[634,705,2077,2101]" captionStart="Figs" captionStartId="8.[835,879,2006,2028]" captionTargetBox="[838,1470,681,1952]" captionTargetId="figure-8@8.[820,1497,669,1967]" captionTargetPageId="8" captionText="Figs. 1720. O. singaporensis, new species, male, structural details, specimen from Singapore, Bukit Timah. 17, Male right paramere. 18, Superior appendage of right paramere. 19, Medial process of pygophore. 20, Endosoma." figureDoi="http://doi.org/10.5281/zenodo.5348165" httpUri="https://zenodo.org/record/5348165/files/figure.png" pageId="1" pageNumber="344">17, 18</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FD0EF79FFC94FF35" box="[718,841,189,2101]" captionStart="Figs" captionStartId="10.[835,880,2003,2025]" captionTargetBox="[834,1473,791,1966]" captionTargetId="figure-8@10.[815,1514,778,1975]" captionTargetPageId="10" captionText="Figs. 2325. O. thienemanni Jaczewski, male, structural details, specimen from Indonesia, Bali, CL 2170. 23, Male right paramere. 24, Superior appendage of right paramere. 25, Medial process of pygophore." figureDoi="http://doi.org/10.5281/zenodo.5348169" httpUri="https://zenodo.org/record/5348169/files/figure.png" pageId="1" pageNumber="344">23, 24</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FC98FF3FFC7FFF35" box="[856,930,189,213]" captionStart="Figs" captionStartId="11.[811,855,2005,2027]" captionTargetBox="[811,1454,541,1948]" captionTargetId="figure-15@11.[789,1477,521,1954]" captionTargetPageId="11" captionText="Figs.2628. O. noualhieri Baehr, male, structural details, specimen from Indonesia, Celebes, CL 2127. 26, Male right paramere. 27, Superior appendage of right paramere. 28, Medial process of pygophore." figureDoi="http://doi.org/10.5281/zenodo.5348173" httpUri="https://zenodo.org/record/5348173/files/figure.png" pageId="1" pageNumber="344">26, 27</figureCitation>
), the shape of the medial process on the male pygophore (
<figureCitation id="13502A5CFFB2C815FC6EFF5FFBC5FF15" box="[942,1048,221,245]" captionStart="Figs" captionStartId="4.[136,181,1919,1941]" captionTargetBox="[138,774,649,1878]" captionTargetId="figure-8@4.[107,796,629,1898]" captionTargetPageId="4" captionText="Figs. 59. O. marginatus (Latreille), male, structural details. 5, Male right paramere, specimen from Vietnam, CL 4368.6, Superior appendage of right paramere, specimen from Vietnam, CL 4368. 7, Superior appendage of right paramere, specimen from Indonesia, Bali, CL 2170. 8, Medial process of pygophore, specimen from Vietnam, CL 4368. 9, Medial process of pygophore, specimen from Indonesia, Bali, CL 2170." figureDoi="http://doi.org/10.5281/zenodo.5348157" httpUri="https://zenodo.org/record/5348157/files/figure.png" pageId="1" pageNumber="344">Figs. 8, 9</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FBE5FF5FFB9FFF15" box="[1061,1090,221,245]" captionStart="Figs" captionStartId="5.[114,159,2006,2028]" captionTargetBox="[117,763,820,1979]" captionTargetId="figure-15@5.[95,766,802,1989]" captionTargetPageId="5" captionText="Figs. 1012. O. signatus, new species, male, structural details, specimen from Malaysia, Perak, CL 2081. 10, Male right paramere. 11, Superior appendage of right paramere. 12, Medial process of pygophore." figureDoi="http://doi.org/10.5281/zenodo.5348159" httpUri="https://zenodo.org/record/5348159/files/figure.png" pageId="1" pageNumber="344">12</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FB8EFF5FFBB6FF15" box="[1102,1131,221,245]" captionStart="Figs" captionStartId="6.[833,877,2004,2026]" captionTargetBox="[830,1474,645,1971]" captionTargetId="figure-686@6.[826,1506,625,1988]" captionTargetPageId="6" captionText="Figs. 1316. O. pseudomarginatus, new species, male, structural details, specimen from Malaysia, Perak, CL 2078. 13, Male right paramere. 14, Superior appendage of right paramere. 15, Medial process of pygophore. 16, Endosoma." figureDoi="http://doi.org/10.5281/zenodo.5348161" httpUri="https://zenodo.org/record/5348161/files/figure.png" pageId="1" pageNumber="344">15</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FBB8FF5FFB48FF15" box="[1144,1173,221,245]" captionStart="Figs" captionStartId="8.[835,879,2006,2028]" captionTargetBox="[838,1470,681,1952]" captionTargetId="figure-8@8.[820,1497,669,1967]" captionTargetPageId="8" captionText="Figs. 1720. O. singaporensis, new species, male, structural details, specimen from Singapore, Bukit Timah. 17, Male right paramere. 18, Superior appendage of right paramere. 19, Medial process of pygophore. 20, Endosoma." figureDoi="http://doi.org/10.5281/zenodo.5348165" httpUri="https://zenodo.org/record/5348165/files/figure.png" pageId="1" pageNumber="344">19</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FB61FF5FFB63FF15" box="[1185,1214,221,245]" captionStart="Figs" captionStartId="10.[835,880,2003,2025]" captionTargetBox="[834,1473,791,1966]" captionTargetId="figure-8@10.[815,1514,778,1975]" captionTargetPageId="10" captionText="Figs. 2325. O. thienemanni Jaczewski, male, structural details, specimen from Indonesia, Bali, CL 2170. 23, Male right paramere. 24, Superior appendage of right paramere. 25, Medial process of pygophore." figureDoi="http://doi.org/10.5281/zenodo.5348169" httpUri="https://zenodo.org/record/5348169/files/figure.png" pageId="1" pageNumber="344">25</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FB0BFF5FFB35FF15" box="[1227,1256,221,245]" captionStart="Figs" captionStartId="11.[811,855,2005,2027]" captionTargetBox="[811,1454,541,1948]" captionTargetId="figure-15@11.[789,1477,521,1954]" captionTargetPageId="11" captionText="Figs.2628. O. noualhieri Baehr, male, structural details, specimen from Indonesia, Celebes, CL 2127. 26, Male right paramere. 27, Superior appendage of right paramere. 28, Medial process of pygophore." figureDoi="http://doi.org/10.5281/zenodo.5348173" httpUri="https://zenodo.org/record/5348173/files/figure.png" pageId="1" pageNumber="344">28</figureCitation>
), the striations on the frons and clypeus (
<figureCitation id="13502A5CFFB2C815FBE9FF7FFB73FEF5" box="[1065,1198,253,277]" captionStart="Figs" captionStartId="12.[136,180,1976,1998]" captionTargetBox="[136,1473,222,1952]" captionTargetId="figure-107@12.[136,1474,222,1953]" captionTargetPageId="12" captionText="Figs. 2934. Ochterus species, colour photographs of head, anterior view, showing pattern of striations on frons.29, O. marginatus (Latreille), female, specimen from Vietnam, Cao Bang Prov., CL 4368. 30, O. thienemanni Jaczewski, female, specimen from Indonesia, Bali Prov., CL 2170. 31, O. signatus, new species, female, specimen from Vietnam, Lam Dong Prov., CL3085. 32, O. pseudomarginatus, new species, female, specimen from Malaysia, Perak, CL 2078. 33, O. noualhieri Baehr, female, specimen from Indonesia, Sulawesi Utara Prov., CL 2114. 34, O. singaporensis, new species, male, specimen from Singapore, Bukit Timah." figureDoi="http://doi.org/10.5281/zenodo.5348175" httpUri="https://zenodo.org/record/5348175/files/figure.png" pageId="1" pageNumber="344">Figs. 2934</figureCitation>
), and the dorsal colour pattern (
<figureCitation id="13502A5CFFB2C815FC46FE9FFC33FED5" box="[902,1006,285,309]" captionStart="Figs" captionStartId="3.[114,158,2033,2055]" captionTargetBox="[113,1453,257,1991]" captionTargetId="figure-15@3.[113,1454,257,1991]" captionTargetPageId="3" captionText="Figs. 14. Ochterus species, colour photographs of dorsum. 1, O. marginatus (Latreille), specimen from Thailand, Chonburi Prov. 2, O. signatus, new species, specimen from Malaysia, Perak, CL 2081. 3, O. pseudomarginatus, new species, specimen from Malaysia, Pahang, CL 2072. 4, O. singaporensis, new species, specimen from Singapore, Bukit Timah." figureDoi="http://doi.org/10.5281/zenodo.5348155" httpUri="https://zenodo.org/record/5348155/files/figure.png" pageId="1" pageNumber="344">Figs. 14</figureCitation>
,
<figureCitation id="13502A5CFFB2C815FC3AFE9FFB9FFED5" box="[1018,1090,285,309]" captionStart="Figs" captionStartId="9.[813,857,2004,2026]" captionTargetBox="[813,1451,337,1981]" captionTargetId="figure-15@9.[813,1452,337,1982]" captionTargetPageId="9" captionText="Figs. 2122. Ochterus species, colour photographs of dorsum. 21, O. thienemanni Jaczewski, specimen from Indonesia, Bali, CL 2170. 22, O. noualhieri Baehr, specimen from Indonesia, Celebes, CL 2127." figureDoi="http://doi.org/10.5281/zenodo.5348167" httpUri="https://zenodo.org/record/5348167/files/figure.png" pageId="1" pageNumber="344">21, 22</figureCitation>
).
</paragraph>
<paragraph id="8BD436D9FFB2C815FCECFEDFFC53FC35" blockId="1.[812,1452,349,981]" pageId="1" pageNumber="344">
Ochterids are usually beautifully marked with spots and flecks of blue, lavendar, green, orange, or yellow on a darker background, and such markings are often distinctive among species and useful in taxon separation. The markings consist of two distinct
<typeStatus id="54D0887BFFB2C815FC1BFE5CFBC5FE16" box="[987,1048,478,502]" pageId="1" pageNumber="344">types</typeStatus>
: depigmented integument (as in the case of yellowish or whitish spots), and pruinose patches of bluish, purplish, or greenish structural colouration occurring on the surface of the integument. The former can become less conspicuous over time in specimens stored for many decades in alcohol, which fades dark colors, while the latter can be degraded or lost in specimens killed by use of ethyl acetate, or pinned through the body so that internal greases and oils spread across the body surface, thereby compromising the structural colors. The best preservation of colour patterns is therefore obtained by collecting specimens directly into 80% ethanol, and then drying them under a lamp in the lab and point-mounting them for long-term vouchering as dry pinned specimens. The photographs of dorsal colour patterns in this work are taken from specimens prepared using such methods.
</paragraph>
<paragraph id="8BD436D9FFB2C815FCECFC7FFB0EFB75" blockId="1.[812,1452,1021,1173]" pageId="1" pageNumber="344">
The family name
<taxonomicName id="4C6B4D5AFFB2C815FC28FC7FFB80FBF5" authorityName="Kirkaldy" authorityYear="1906" box="[1000,1117,1021,1045]" class="Insecta" family="Ochteridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="family">Ochteridae</taxonomicName>
was first proposed by
<bibRefCitation id="EFFA4B28FFB2C815FA8CFC7FFCAAFBD5" author="Kirkaldy, G" pageId="1" pageNumber="344" pagination="117 - 156" refId="ref13191" refString="Kirkaldy, G. W., 1906. List of the pagiopodus Hemiptera- Heteroptera, with their type species, from 1758 to 1901 (and also of the aquatic and semi-aquatic Trochalopoda). Transactions of the American Entomological Society, 32: 117 - 156." type="journal article" year="1906">Kirkaldy (1906)</bibRefCitation>
. Because
<bibRefCitation id="EFFA4B28FFB2C815FC28FB9FFB48FBD5" author="Latreille, P" box="[1000,1173,1053,1077]" pageId="1" pageNumber="344" refId="ref13405" refString="Latreille, P. A., 1809. Genera crustaceorum et insectorum secundum ordinem naturalem in familias disposit, iconibus exemplisque plurimis explicata, Volume 4. A. Konig, Parisiis et Argentorati. 399 pp." type="book" year="1809">Latreille (1809)</bibRefCitation>
had earlier proposed the unnecessary replacement name
<taxonomicName id="4C6B4D5AFFB2C815FBBCFBBCFB30FBB5" authorityName="Latreille" authorityYear="1809" box="[1148,1261,1086,1109]" class="Insecta" family="Ochteridae" genus="Pelogonus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEACBFFB2C815FBBCFBBCFB30FBB5" box="[1148,1261,1086,1109]" italics="true" pageId="1" pageNumber="344">Pelogonus</emphasis>
</taxonomicName>
for
<taxonomicName id="4C6B4D5AFFB2C815FAD6FBBCFAA5FBB5" authorityName="Latreille" authorityYear="1807" box="[1302,1400,1086,1109]" class="Insecta" family="Ochteridae" genus="Ochterus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEACBFFB2C815FAD6FBBCFAA5FBB5" box="[1302,1400,1086,1109]" italics="true" pageId="1" pageNumber="344">Ochterus</emphasis>
</taxonomicName>
, this family is treated under the name Pelogonidae in many works published prior to Kirkaldys checklist.
</paragraph>
</subSubSection>
</treatment>
</document>