224 lines
34 KiB
XML
224 lines
34 KiB
XML
<document id="E723A8D27756053F7DE8B86E6908A774" ID-DOI="10.1111/zoj.12114" ID-ISSN="0024-4082" ID-Zenodo-Dep="10273948" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630109995688" checkinUser="marcus" docAuthor="Fikáček, Martin, Prokin, Alexander, Yan, Evgeny, Yue, Yanli, Wang, Bo, Ren, Dong & Beattie, Robert" docDate="2014" docId="03C287ACFF85D671FC06BD96446D9C00" docLanguage="en" docName="zoj12114.pdf" docOrigin="Zoological Journal of the Linnean Society 170 (4)" docSource="http://doi.wiley.com/10.1111/zoj.12114" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Alegorius Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie 2014, GEN. NOV." docType="treatment" docVersion="3" lastPageNumber="716" masterDocId="FFFBFFD4FF80D676FFD0B961454A9D19" masterDocTitle="Modern hydrophilid clades present and widespread in the Late Jurassic and Early Cretaceous (Coleoptera: Hydrophiloidea: Hydrophilidae)" masterLastPageNumber="734" masterPageNumber="710" pageNumber="715" updateTime="1701826641646" updateUser="ExternalLinkService">
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<mods:title id="F8434EB0C72F1D9FC40B158BF6764150">Modern hydrophilid clades present and widespread in the Late Jurassic and Early Cretaceous (Coleoptera: Hydrophiloidea: Hydrophilidae)</mods:title>
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<mods:namePart id="BA70FD5B216969E8F8DB17A09C8831E1">Fikáček, Martin</mods:namePart>
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<mods:namePart id="3CB7C6DE2F88F3EAA94FFDC513AB0D48">Prokin, Alexander</mods:namePart>
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<mods:namePart id="D088D1E4F5A2F6E245E18052A55315CC">Yan, Evgeny</mods:namePart>
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<mods:roleTerm id="54676107533E04385F51047768EF4761">Author</mods:roleTerm>
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<mods:namePart id="A639A7C144F945629E4142D140595830">Yue, Yanli</mods:namePart>
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<mods:roleTerm id="B476F9E28D33EF19530BF453650CFECA">Author</mods:roleTerm>
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<mods:namePart id="9682CCCC764C4A3B80F757F246739141">Wang, Bo</mods:namePart>
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<mods:namePart id="D5058DBAB52AC66AD4024A89FAA1A05A">Ren, Dong</mods:namePart>
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<mods:namePart id="0BCD81F95C46CBFDA6812B8810A506F3">Beattie, Robert</mods:namePart>
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<mods:typeOfResource id="30F75292B10EF3A2806444758DB04B8D">text</mods:typeOfResource>
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<mods:title id="5FD04306E2C57603BB4C24F5DD6CF526">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="9420E81B9E7BDA95A3FC8648A7AA5868">
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<mods:date id="A2ACDD87973B27CF5092C41048857659">2014</mods:date>
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<mods:number id="DE70D8CCDA94DD6F0C9E11E9A88A6340">2014-04-30</mods:number>
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<mods:number id="02EB7EAE3F6026FD8470076F0ABC599F">170</mods:number>
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<mods:number id="D4005AFA7FB5CB08C8B29B879EA4B844">4</mods:number>
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<mods:start id="2E1C9A6904931F7205382E3B41735EF0">710</mods:start>
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<mods:url id="2FEA51FC6EE0EF1812A353A78531184B">http://doi.wiley.com/10.1111/zoj.12114</mods:url>
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<mods:classification id="39DD8188DEE5D3B5EC871EEBBAB5A717">journal article</mods:classification>
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<mods:identifier id="956B79E6809B27E0C60328FA2AA057E9" type="DOI">10.1111/zoj.12114</mods:identifier>
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<mods:identifier id="F5F70D446763217078E580624D789EF2" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="50026F3D8328CC5805CB7D4865EB9A56" type="Zenodo-Dep">10273948</mods:identifier>
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<subSubSection id="C3716531FF85D673FC06BD9641979814" box="[982,1245,1271,1293]" pageId="5" pageNumber="715" type="nomenclature">
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<paragraph id="8BD436BAFF85D673FC06BD9641979814" blockId="5.[982,1245,1271,1293]" box="[982,1245,1271,1293]" pageId="5" pageNumber="715">
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<heading id="D09C81D6FF85D673FC06BD9641979814" box="[982,1245,1271,1293]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="715" reason="4">
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<taxonomicName id="4C6B4D39FF85D673FC06BD9641149815" ID-CoL="8XJFF" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[982,1118,1271,1292]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="715" phylum="Arthropoda" rank="genus" status="gen. nov.">ALEGORIUS</taxonomicName>
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<emphasis id="B91FEAA8FF85D673FBB8BD9A41979814" bold="true" box="[1128,1245,1272,1293]" pageId="5" pageNumber="715">
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<taxonomicNameLabel id="A22C57D3FF85D673FBB8BD9A41979814" box="[1128,1245,1272,1293]" pageId="5" pageNumber="715" rank="genus">GEN. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3716531FF85D671FCF6BC7E446D9C00" lastPageId="7" lastPageNumber="717" pageId="5" pageNumber="715" type="materials_examined">
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<paragraph id="8BD436BAFF85D673FCF6BC7E46F8984A" blockId="5.[806,1421,1310,1363]" pageId="5" pageNumber="715">
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<emphasis id="B91FEAA8FF85D673FCF6BC7E41D1982D" box="[806,1179,1310,1332]" italics="true" pageId="5" pageNumber="715">
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<typeStatus id="54D08818FF85D673FCF6BC7E4610982D" box="[806,858,1311,1332]" pageId="5" pageNumber="715">Type</typeStatus>
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species:
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<taxonomicName id="4C6B4D39FF85D673FC15BC7F41D1982D" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[965,1179,1310,1332]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="715" phylum="Arthropoda" rank="species" species="yixianus" status="sp. nov.">Alegorius yixianus</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A22C57D3FF85D673FB77BC7F4048982A" box="[1191,1282,1310,1331]" pageId="5" pageNumber="715" rank="species">sp. nov.</taxonomicNameLabel>
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, by present designation.
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</paragraph>
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<paragraph id="8BD436BAFF85D673FCF6BCE541C29880" blockId="5.[806,1160,1412,1434]" box="[806,1160,1412,1434]" pageId="5" pageNumber="715">
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<emphasis id="B91FEAA8FF85D673FCF6BCE546E59880" box="[806,943,1412,1433]" italics="true" pageId="5" pageNumber="715">Time range:</emphasis>
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Early Cretaceous.
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</paragraph>
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<paragraph id="8BD436BAFF85D673FCF6BCAD40259A76" blockId="5.[806,1421,1483,1903]" pageId="5" pageNumber="715">
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<emphasis id="B91FEAA8FF85D673FCF6BCAD46D798F8" box="[806,925,1484,1505]" italics="true" pageId="5" pageNumber="715">Diagnosis:</emphasis>
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Body medium-sized; labrum partly exposed in dorsal view (
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<figureCitation id="13502A3FFF85D673FBB0BC8B419098E6" box="[1120,1242,1514,1536]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Figs 2A–B</figureCitation>
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,
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<figureCitation id="13502A3FFF85D673FB20BC8B40589B19" box="[1264,1298,1514,1536]" captionStart="Figure 8" captionStartId="16.[164,243,909,928]" captionTargetBox="[172,1436,205,871]" captionTargetId="figure-117@16.[164,1444,197,879]" captionTargetPageId="16" captionText="Figure 8. Reconstructions of the clypeus shape and meso- and metathoracic morphology of the Early Cretaceous hydrophilid genera known in adult stage (three-dimensional structure of the mesoventrite not reconstructed). Alegorius gen. nov. (A, B); Cretoxenus gen. nov. (C, D); Hydroyixia gen. nov. (E, F). Head and thorax of the respective genus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10273971" httpUri="https://zenodo.org/record/10273971/files/figure.png" pageId="5" pageNumber="715">8A</figureCitation>
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); anterior margin of clypeus weakly excised mesally (
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<figureCitation id="13502A3FFF85D673FAFFBF6940C29B04" box="[1327,1416,1544,1566]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Figs 2B</figureCitation>
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,
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<figureCitation id="13502A3FFF85D673FCF6BF4646029B24" box="[806,840,1575,1597]" captionStart="Figure 8" captionStartId="16.[164,243,909,928]" captionTargetBox="[172,1436,205,871]" captionTargetId="figure-117@16.[164,1444,197,879]" captionTargetPageId="16" captionText="Figure 8. Reconstructions of the clypeus shape and meso- and metathoracic morphology of the Early Cretaceous hydrophilid genera known in adult stage (three-dimensional structure of the mesoventrite not reconstructed). Alegorius gen. nov. (A, B); Cretoxenus gen. nov. (C, D); Hydroyixia gen. nov. (E, F). Head and thorax of the respective genus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10273971" httpUri="https://zenodo.org/record/10273971/files/figure.png" pageId="5" pageNumber="715">8A</figureCitation>
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); prosternum moderately long, weakly carinate/ elevated medially (
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<figureCitation id="13502A3FFF85D673FBDFBF24412E9B42" box="[1039,1124,1605,1627]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Fig. 2B</figureCitation>
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); scutellar shield small, triangular; anapleural sutures well developed, strongly curved, nearly parallel-sided and widely separate anteriorly (
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<figureCitation id="13502A3FFF85D673FBC7BFC041DA9BAF" box="[1047,1168,1697,1719]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Figs 2A–B</figureCitation>
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,
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<figureCitation id="13502A3FFF85D673FB4FBFC0418A9BAE" box="[1183,1216,1697,1719]" captionStart="Figure 8" captionStartId="16.[164,243,909,928]" captionTargetBox="[172,1436,205,871]" captionTargetId="figure-117@16.[164,1444,197,879]" captionTargetPageId="16" captionText="Figure 8. Reconstructions of the clypeus shape and meso- and metathoracic morphology of the Early Cretaceous hydrophilid genera known in adult stage (three-dimensional structure of the mesoventrite not reconstructed). Alegorius gen. nov. (A, B); Cretoxenus gen. nov. (C, D); Hydroyixia gen. nov. (E, F). Head and thorax of the respective genus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10273971" httpUri="https://zenodo.org/record/10273971/files/figure.png" pageId="5" pageNumber="715">8B</figureCitation>
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); mesocoxae very narrowly separated; abdomen with five ventrites; apical abdominal ventrite with deep and narrow emargination (
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<figureCitation id="13502A3FFF85D673FC0CBF9C417D9A0B" box="[988,1079,1789,1811]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Figs 2A</figureCitation>
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,
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<figureCitation id="13502A3FFF85D673FB80BF9C41389A0A" box="[1104,1138,1789,1811]" captionStart="Figure 7" captionStartId="15.[144,223,1704,1723]" captionTargetBox="[208,1360,195,1674]" captionTargetId="figure-0@15.[208,1360,195,1674]" captionTargetPageId="15" captionText="Figure 7. Hydrophilid fossils from Early Cretaceous Yixian Formation, China. Alegorius yixianus sp. nov.: holotype CNU 2009079, whole specimen (A), detail of meso- and metaventrite (B), detail of abdominal apex (C); paratype CNU 2009078, whole specimen (D), detail of mesoventrite (E). Hydroyixia elongata sp. nov.: paratype CNU 2009075, detail of head and pronotum (F), whole specimen (G); holotype CNU 2010005: whole beetle, piece and counterpiece (H, I), detail of head (J), detail of meso- and metaventrite (K), detail of abdominal apex (L). Hydroyixia latissima sp. nov.: holotype CNU 2009074, detail of abdominal apex (M), whole specimen in dry condition (N) and under alcohol (O). Abbreviations: apls, anapleural suture; abem, apical emargination of abdominal ventrite 5." figureDoi="http://doi.org/10.5281/zenodo.10273968" httpUri="https://zenodo.org/record/10273968/files/figure.png" pageId="5" pageNumber="715">7C</figureCitation>
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); elytron with deeply impressed sutural stria; elytra with series of punctures (
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<figureCitation id="13502A3FFF85D673FCA8BE5B46849A49" box="[888,974,1850,1872]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Fig. 2A</figureCitation>
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); tarsi of mesothoracic leg with five tarsomeres; mesotarsomere 1 very short (
|
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<figureCitation id="13502A3FFF85D673FADDBE3840289A76" box="[1293,1378,1881,1903]" captionStart="Figure 2" captionStartId="6.[164,243,1050,1069]" captionTargetBox="[323,1283,195,1013]" captionTargetId="figure-319@6.[323,1283,195,1020]" captionTargetPageId="6" captionText="Figure 2. Alegorius yixianus gen. nov., sp. nov. from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax." figureDoi="http://doi.org/10.5281/zenodo.10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="5" pageNumber="715">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF146632FF86D670FF74BD7B411C999D" ID-DOI="http://doi.org/10.5281/zenodo.10273954" ID-Zenodo-Dep="10273954" httpUri="https://zenodo.org/record/10273954/files/figure.png" pageId="6" pageNumber="716" startId="6.[164,243,1050,1069]" targetBox="[323,1283,195,1013]" targetPageId="6">
|
||
<paragraph id="8BD436BAFF86D670FF74BD7B411C999D" blockId="6.[164,1441,1049,1157]" pageId="6" pageNumber="716">
|
||
<emphasis id="B91FEAA8FF86D670FF74BD7B479C9934" bold="true" box="[164,726,1049,1069]" pageId="6" pageNumber="716">
|
||
Figure 2.
|
||
<taxonomicName id="4C6B4D39FF86D670FEC7BD7844BE9934" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[279,500,1049,1069]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="species" species="yixianus" status="gen. nov., sp. nov.">
|
||
<emphasis id="B91FEAA8FF86D670FEC7BD7844BE9934" bold="true" box="[279,500,1049,1069]" italics="true" pageId="6" pageNumber="716">Alegorius yixianus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D3FF86D670FDD0BD7B479C9934" box="[512,726,1050,1069]" pageId="6" pageNumber="716" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the Yixian Formation, China. Holotype, CNU 2009079 (A); paratype, CNU 2009078 (B). Abbreviations: abem, apical emargination of abdominal ventrite 5; aes3, metanepisternum; bst, maxillary basistipes; fmtta, free metatibial anterobasal angle; lb, labrum; men, mentum; mstr1, mesotarsomere 1; msv, mesoventrite; pros, prosternum; sstr, sutural stria; tf, transverse fold of prothorax.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD436BAFF86D670FF74BDD744C998D8" blockId="6.[164,779,1206,1473]" pageId="6" pageNumber="716">
|
||
<emphasis id="B91FEAA8FF86D670FF74BDD744D599D2" box="[164,415,1206,1227]" italics="true" pageId="6" pageNumber="716">
|
||
Etymology:
|
||
<taxonomicName id="4C6B4D39FF86D670FEE4BDD744D599D2" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[308,415,1206,1227]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus">Alegorius</taxonomicName>
|
||
</emphasis>
|
||
is an arbitrary combination of letters inspired and loosely derived from the given and patronymic names Alexander Georgievich, referring to three Russian specialists on fossil and aquatic beetles, Alexander Georgievich Ponomarenko, Alexander Georgievich Kirejtshuk and Alexander Georgievich Shatrovskiy, who supported the authors of this paper in their palaeontological and taxonomic studies. Masculine.
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF86D670FF74BC8847559BED" blockId="6.[164,779,1513,1780]" pageId="6" pageNumber="716">
|
||
<emphasis id="B91FEAA8FF86D670FF74BC8844CE98E6" box="[164,388,1513,1535]" italics="true" pageId="6" pageNumber="716">Family assignment:</emphasis>
|
||
The combination of large clypeus, well-developed frontoclypeal sutures reaching lateral margin of head closely before eyes, head not constricted behind eyes, small triangular scutellar shield and femora with anteromesal corner not facing trochanter (
|
||
<figureCitation id="13502A3FFF86D670FEDBBFE344049B8E" box="[267,334,1666,1688]" captionStart="Figure 9" captionStartId="19.[144,223,1853,1872]" captionTargetBox="[327,1240,906,1823]" captionTargetId="figure-313@19.[327,1240,906,1823]" captionTargetPageId="19" captionText="Figure 9. Known distribution of the Hydrophilidae in the Late Jurassic and Early Cretaceous. 1, Solnhofen, Germany; 2, Talbragar, Australia; 3, Baissa, Russia; 4, Yixian Formation, China; 5, Koonwarra, Australia." figureDoi="http://doi.org/10.5281/zenodo.10273975" httpUri="https://zenodo.org/record/10273975/files/figure.png" pageId="6" pageNumber="716">Fig. 9</figureCitation>
|
||
, fmtta) assign the fossil reliably to the family
|
||
<taxonomicName id="4C6B4D39FF86D670FF25BFC144D39BAF" box="[245,409,1696,1718]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
|
||
<emphasis id="B91FEAA8FF86D670FE72BFC0448F9BAF" box="[418,453,1697,1718]" italics="true" pageId="6" pageNumber="716">s.s.</emphasis>
|
||
Additionally, it corresponds well with the modern
|
||
<taxonomicName id="4C6B4D39FF86D670FE62BFDE471C9BCC" box="[434,598,1727,1749]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
|
||
in the general habitus and ventral morphology.
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF86D670FF74BE7D41B29B8E" blockId="6.[164,779,1819,1903]" lastBlockId="6.[826,1441,1206,1901]" pageId="6" pageNumber="716">
|
||
<emphasis id="B91FEAA8FF86D670FF74BE7D47499A28" box="[164,515,1819,1841]" italics="true" pageId="6" pageNumber="716">Comparison with other genera:</emphasis>
|
||
Based on the presence of the apical emargination on the posterior margin of the abdominal ventrite 5,
|
||
<taxonomicName id="4C6B4D39FF86D670FE25BE38472A9A77" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[501,608,1881,1902]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF86D670FE25BE38472A9A77" box="[501,608,1881,1902]" italics="true" pageId="6" pageNumber="716">Alegorius</emphasis>
|
||
</taxonomicName>
|
||
resembles the extant representatives of the
|
||
<taxonomicName id="4C6B4D39FF86D670FB44BDD7406399D5" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[1172,1321,1206,1228]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
and early branching
|
||
<taxonomicName id="4C6B4D39FF86D670FC67BDB4410699F2" authorityName="Latreille" authorityYear="1802" box="[951,1100,1237,1259]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="tribe" tribe="Hydrophilini">Hydrophilini</taxonomicName>
|
||
, and those of the subfamilies
|
||
<taxonomicName id="4C6B4D39FF86D670FCEABD9246F79810" authorityName="Short & Fikacek" authorityYear="2013" box="[826,957,1267,1289]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="subFamily" subFamily="Enochrinae">Enochrinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D39FF86D670FBDCBD9241E89810" baseAuthorityName="Zaitzev" baseAuthorityYear="1908" box="[1036,1186,1267,1289]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
|
||
. In contrast to the
|
||
<taxonomicName id="4C6B4D39FF86D670FCEABC7346859831" baseAuthorityName="Mulsant" baseAuthorityYear="1844" box="[826,975,1298,1320]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="tribe" tribe="Hydrobiusini">Hydrobiusini</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D39FF86D670FBC4BC7341E39831" authorityName="Latreille" authorityYear="1802" box="[1044,1193,1298,1320]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="tribe" tribe="Hydrophilini">Hydrophilini</taxonomicName>
|
||
, the mesoventrite of
|
||
<taxonomicName id="4C6B4D39FF86D670FCEABC5046EF985F" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[826,933,1329,1350]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF86D670FCEABC5046EF985F" box="[826,933,1329,1350]" italics="true" pageId="6" pageNumber="716">Alegorius</emphasis>
|
||
</taxonomicName>
|
||
is wide anteriorly and does not bear a common ventral keel; in contrast to the
|
||
<taxonomicName id="4C6B4D39FF86D670FAC9BC2E40EB987D" authorityName="Bedel" authorityYear="1881" box="[1305,1441,1359,1380]" class="Insecta" family="Hydrophilidae" genus="Cymbiodyta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF86D670FAC9BC2E40EB987D" box="[1305,1441,1359,1380]" italics="true" pageId="6" pageNumber="716">Cymbiodyta</emphasis>
|
||
</taxonomicName>
|
||
group of the
|
||
<taxonomicName id="4C6B4D39FF86D670FC04BC0F4113989D" authorityName="Short & Fikacek" authorityYear="2013" box="[980,1113,1390,1412]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="subFamily" subFamily="Enochrinae">Enochrinae</taxonomicName>
|
||
, the mesothoracic leg bears five tarsomeres. Hence, it seems that
|
||
<taxonomicName id="4C6B4D39FF86D670FB2BBCED402C98B8" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1275,1382,1420,1441]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF86D670FB2BBCED402C98B8" box="[1275,1382,1420,1441]" italics="true" pageId="6" pageNumber="716">Alegorius</emphasis>
|
||
</taxonomicName>
|
||
may be assigned either to the
|
||
<taxonomicName id="4C6B4D39FF86D670FB5EBCCA41B098D9" box="[1166,1274,1451,1472]" class="Insecta" family="Hydrophilidae" genus="Enochrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA8FF86D670FB5EBCCA41B098D9" box="[1166,1274,1451,1472]" italics="true" pageId="6" pageNumber="716">Enochrus</emphasis>
|
||
</taxonomicName>
|
||
-group of the
|
||
<taxonomicName id="4C6B4D39FF86D670FCEABCA846F798C6" authorityName="Short & Fikacek" authorityYear="2013" box="[826,957,1481,1503]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="subFamily" subFamily="Enochrinae">Enochrinae</taxonomicName>
|
||
or to the subfamily
|
||
<taxonomicName id="4C6B4D39FF86D670FB65BCA8400198C6" baseAuthorityName="Zaitzev" baseAuthorityYear="1908" box="[1205,1355,1481,1503]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="subFamily" subFamily="Acidocerinae">Acidocerinae</taxonomicName>
|
||
. It corresponds with extant species of both clades in general body shape, presence of the shallow anterior excision of the clypeus, and the presence of sharply impressed sutural stria. A more detailed comparison with extant genera is not possible because relevant characters are not preserved in the examined fossils.
|
||
</paragraph>
|
||
<paragraph id="8BD436BAFF86D671FC83BFC1446D9C00" blockId="6.[826,1441,1206,1901]" lastBlockId="7.[144,759,198,281]" lastPageId="7" lastPageNumber="717" pageId="6" pageNumber="716">
|
||
<taxonomicName id="4C6B4D39FF86D670FC83BFC146F49BAC" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[851,958,1696,1717]" class="Insecta" family="Hydrophilidae" genus="Alegorius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA8FF86D670FC83BFC146F49BAC" box="[851,958,1696,1717]" italics="true" pageId="6" pageNumber="716">Alegorius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D3FF86D670FC1BBFC1417F9BAC" box="[971,1077,1696,1717]" pageId="6" pageNumber="716" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
differs from other Cretaceous fossil genera of the
|
||
<taxonomicName id="4C6B4D39FF86D670FBFBBFDF41859BCD" box="[1067,1231,1726,1748]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
|
||
especially by the strongly curved anapleural sutures that are nearly parallel and widely separated anteriorly (in contrast to straight anapleural sutures of
|
||
<taxonomicName id="4C6B4D39FF86D670FB34BE7B402B9A36" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[1252,1377,1818,1839]" class="Insecta" family="Hydrophilidae" genus="Cretoxenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA8FF86D670FB34BE7B402B9A36" box="[1252,1377,1818,1839]" italics="true" pageId="6" pageNumber="716">Cretoxenus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D3FF86D670FAA2BE7A46229A57" pageId="6" pageNumber="716" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
and anteriorly joining anapleural sutures of
|
||
<taxonomicName id="4C6B4D39FF86D670FC8FBE3646979A75" authority="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie, 2014" authorityName="Fikáček & Prokin & Yan & Yue & Wang & Ren & Beattie" authorityYear="2014" box="[863,989,1879,1900]" class="Insecta" family="Hydrophilidae" genus="Hydroyixia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="716" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA8FF86D670FC8FBE3646979A75" box="[863,989,1879,1900]" italics="true" pageId="6" pageNumber="716">Hydroyixia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D3FF86D670FC3EBE3941179A74" box="[1006,1117,1880,1901]" pageId="6" pageNumber="716" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, compare
|
||
<figureCitation id="13502A3FFF86D670FB32BE3640DE9A74" box="[1250,1428,1879,1901]" captionStart="Figure 8" captionStartId="16.[164,243,909,928]" captionTargetBox="[172,1436,205,871]" captionTargetId="figure-117@16.[164,1444,197,879]" captionTargetPageId="16" captionText="Figure 8. Reconstructions of the clypeus shape and meso- and metathoracic morphology of the Early Cretaceous hydrophilid genera known in adult stage (three-dimensional structure of the mesoventrite not reconstructed). Alegorius gen. nov. (A, B); Cretoxenus gen. nov. (C, D); Hydroyixia gen. nov. (E, F). Head and thorax of the respective genus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10273971" httpUri="https://zenodo.org/record/10273971/files/figure.png" pageId="6" pageNumber="716">Figs. 8B, D, F</figureCitation>
|
||
). Detailed comparison with Jurassic fossils described above is not possible due to their incomplete preservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |