619 lines
78 KiB
XML
619 lines
78 KiB
XML
<document id="4B006472631FBB76F9DB9090CB7872CF" ID="10.11646/zootaxa.4169.3.11" ID-DOI="10.11646/zootaxa.4169.3.11" ID-GBIF-Dataset="afa0887e-f70a-4605-83f5-950003e5f668" ID-ISSN="1175-5326" ID-Zenodo-Dep="272404" ID-ZooBank="982AADB9-A2A3-4E61-9B23-9C85BF044200" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474441700122" checkinUser="plazi" docAuthor="Boonsoong, Boonsatien & Sartori, Michel" docDate="2016" docId="03C287AC2B3EFFABFF3653F9FCA519A6" docLanguage="en" docName="zootaxa.4169.3.11.pdf" docOrigin="Zootaxa 4169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sangpradubina Boonsoong & Sartori, 2016, gen. nov." docType="treatment" docVersion="7" lastPageNumber="591" masterDocId="FFFBFFD42B3FFFAFFFA1536EFFBB1821" masterDocTitle="Sangpradubina, an astonishing new mayfly genus from Thailand (Ephemeroptera: Leptophlebiidae: Atalophlebiinae)" masterLastPageNumber="599" masterPageNumber="587" pageNumber="588" updateTime="1698386711426" updateUser="plazi">
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<mods:mods id="9F12B59C1F05CBE49BBEDEB1F4D96C3C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A0EF696F12ED6E5F938E0C5DFAF6482A">
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<mods:title id="779B8977EF7930C4A00B7036DA308CD0">Sangpradubina, an astonishing new mayfly genus from Thailand (Ephemeroptera: Leptophlebiidae: Atalophlebiinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="056C4C01BFD75CC6B2E014A65D8B5F9A" type="personal">
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<mods:roleTerm id="449B5DB7183C0276FC3871B164E2EA81">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="75A03ECDBD9CE98A4C254D8F6556F49E">Boonsoong, Boonsatien</mods:namePart>
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</mods:name>
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<mods:name id="CC8A123079500909CFBB20AC87D8ED99" type="personal">
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<mods:role id="732BEB7CA03F2B670BA726AE79726B36">
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<mods:roleTerm id="65C3EF811696EE012CC978FD4724B4A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0BF654C408A931ABC6691C66E7784B92">Sartori, Michel</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="6669A7A23007A387A54986D6F7FDC6FC">text</mods:typeOfResource>
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<mods:title id="EE41F7204E0788C9C7DF5D8669FEDBFA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EF528B4DA9ADF71A5774183C3B219E78">
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<mods:date id="B9CF74D08E3D6F0CCFA721D4C65919D4">2016</mods:date>
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<mods:detail id="2C138D35E19DB06100D9091E355615BA" type="volume">
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<mods:number id="A9B3CF8050901E1D8DA6FB868A90C7C4">4169</mods:number>
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</mods:detail>
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<mods:detail id="498578A1E8731CA74A8AC432BFDB3BDF" type="issue">
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<mods:number id="F9DBEF2BA2B926B8628C0119FE19C418">3</mods:number>
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<mods:start id="C4850854C5A079B8360F4AD9AEBEC3E1">587</mods:start>
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<mods:classification id="72FC7F6EFEF8A461A0027D0BDBC8D1AE">journal article</mods:classification>
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<mods:identifier id="BC0003B80A61833E5D4120456F08D2EE" type="DOI">10.11646/zootaxa.4169.3.11</mods:identifier>
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<mods:identifier id="2272B0C64F16EEC8B2B37AE354AC2CA7" type="GBIF-Dataset">afa0887e-f70a-4605-83f5-950003e5f668</mods:identifier>
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<mods:identifier id="29EAC5F07FF32D6863A4FB3A90236913" type="Zenodo-Dep">272404</mods:identifier>
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</mods:mods>
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<treatment id="03C287AC2B3EFFABFF3653F9FCA519A6" ID-DOI="http://doi.org/10.5281/zenodo.6075150" ID-GBIF-Taxon="124552539" ID-Zenodo-Dep="6075150" LSID="urn:lsid:plazi:treatment:03C287AC2B3EFFABFF3653F9FCA519A6" httpUri="http://treatment.plazi.org/id/03C287AC2B3EFFABFF3653F9FCA519A6" lastPageId="4" lastPageNumber="591" pageId="1" pageNumber="588">
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<subSubSection id="C37165312B3EFFAEFF3653F9FE7C1893" box="[151,455,151,178]" pageId="1" pageNumber="588" type="nomenclature">
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<paragraph id="8BD436BA2B3EFFAEFF3653F9FE7C1893" blockId="1.[151,455,151,178]" box="[151,455,151,178]" pageId="1" pageNumber="588">
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<heading id="D09C81D62B3EFFAEFF3653F9FE7C1893" bold="true" box="[151,455,151,178]" fontSize="11" level="1" pageId="1" pageNumber="588" reason="1">
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<emphasis id="B91FEAA82B3EFFAEFF3653F9FE7C1893" bold="true" box="[151,455,151,178]" pageId="1" pageNumber="588">
|
||
<taxonomicName id="4C6B4D392B3EFFAEFF3653F9FEE81890" box="[151,339,151,177]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3EFFAEFF3653F9FEE81890" bold="true" box="[151,339,151,177]" italics="true" pageId="1" pageNumber="588">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3EFFAEFEF853F6FE7C1893" box="[345,455,152,178]" pageId="1" pageNumber="588" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37165312B3EFFAEFF3653B1FDE41A5F" pageId="1" pageNumber="588" type="materials_examined">
|
||
<paragraph id="8BD436BA2B3EFFAEFF3653B1FCC618D5" blockId="1.[151,893,222,244]" box="[151,893,222,244]" pageId="1" pageNumber="588">
|
||
Type species:
|
||
<taxonomicName id="4C6B4D392B3EFFAEFE8753B0FD8018D5" box="[294,571,222,244]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="thailandica" status="sp. nov.">
|
||
<emphasis id="B91FEAA82B3EFFAEFE8753B0FD8018D5" box="[294,571,222,244]" italics="true" pageId="1" pageNumber="588">Sangpradubina thailandica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3EFFAEFDE353B1FD2918D5" box="[578,658,223,244]" pageId="1" pageNumber="588" rank="species">
|
||
<emphasis id="B91FEAA82B3EFFAEFDE353B1FD3718D5" bold="true" box="[578,652,223,244]" pageId="1" pageNumber="588">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by present designation
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3EFFAEFF36524FFDE41A5F" blockId="1.[151,1437,289,857]" pageId="1" pageNumber="588">
|
||
<emphasis id="B91FEAA82B3EFFAEFF36524FFEF7191B" bold="true" box="[151,332,289,314]" pageId="1" pageNumber="588">Mature nymph</emphasis>
|
||
.
|
||
<emphasis id="B91FEAA82B3EFFAEFEFC524FFE19191B" box="[349,418,289,314]" italics="true" pageId="1" pageNumber="588">Head:</emphasis>
|
||
Head prognathous. Labrum with 5 flat denticles in anterior emargination. Mandible with tuft of long setae in middle of lateral margin. Maxillae with one comb-shaped dentiseta, and with well-developed anterolateral projection, as long as dentiseta; palp 3-segmented. Hypopharynx with superlingua well-developed laterally; lingua with pair of lateral processes near base and pointed distally. Labium with glossae dorsal to paraglossae; labial palp 3-segmented.
|
||
<emphasis id="B91FEAA82B3EFFAEFDEF52DFFD1319EB" box="[590,680,433,458]" italics="true" pageId="1" pageNumber="588">Thorax:</emphasis>
|
||
Fore- and middle femora each with dense rows of long setae on outer margin. All legs without tibio-patellar suture, but with stout setae on outer margin of femur and stout bristles scattered on dorsal surface. Claws stout, hooked and narrowed apically, 4 denticles at base and 9 at apex.
|
||
<emphasis id="B91FEAA82B3EFFAEFA885297FA271A33" box="[1321,1436,505,530]" italics="true" pageId="1" pageNumber="588">Abdomen:</emphasis>
|
||
Posterolateral spines present on segments 6–9. Abdominal gill 1 single, consisting of slender lamella; gills 2–7 alike, dorsal and ventral lamellae plate-like, apical half of each lamella with fringed margins. Caudal filaments with whorls of thin setae on each articulation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37165312B3EFFADFF6651E7FE641E60" lastPageId="2" lastPageNumber="589" pageId="1" pageNumber="588" type="description">
|
||
<paragraph id="8BD436BA2B3EFFAEFF6651E7FA261B79" blockId="1.[151,1437,289,857]" pageId="1" pageNumber="588">
|
||
<emphasis id="B91FEAA82B3EFFAEFF6651E7FEA81A83" bold="true" box="[199,275,649,674]" pageId="1" pageNumber="588">Imago</emphasis>
|
||
.
|
||
<emphasis id="B91FEAA82B3EFFAEFE8451E7FED01A83" box="[293,363,649,674]" italics="true" pageId="1" pageNumber="588">Head:</emphasis>
|
||
Eyes of male meet on meson.
|
||
<emphasis id="B91FEAA82B3EFFAEFD4351E7FC871A83" box="[738,828,649,674]" italics="true" pageId="1" pageNumber="588">Thorax:</emphasis>
|
||
Tibio-patellar suture absent on forelegs, present on middle and hind legs. Tarsal claws of all legs similar, one apically hooked, other obtuse, pad-like. Forewing with vein MA symmetrically forked, vein MP not forked; MP2 independent of vein MP1;
|
||
<specimenCount id="9D6DFD332B3EFFAEFBF851BFFB421ACB" box="[1113,1273,721,746]" pageId="1" pageNumber="588" type="intercalary">5 intercalaries</specimenCount>
|
||
between CuA and CuP. Hind wings with rounded apex, costal margin convex, with weakly developed projection.
|
||
<emphasis id="B91FEAA82B3EFFAEFB485196FAE71B30" box="[1257,1372,760,785]" italics="true" pageId="1" pageNumber="588">Abdomen:</emphasis>
|
||
Male genitalia with forceps 3-segmented, styliger strongly convex; penes lobes stocky, with widest part at base; gonopore ventral, in subapical position. Female subanal plate distally narrow and rounded, with small cleft at apex.
|
||
</paragraph>
|
||
<caption id="DF1466322B3EFFAEFF3654AFFF751FD7" httpUri="https://zenodo.org/record/272405/files/figure.png" pageId="1" pageNumber="588" targetBox="[340,1246,909,1963]" targetPageId="1">
|
||
<paragraph id="8BD436BA2B3EFFAEFF3654AFFF751FD7" blockId="1.[151,1436,1985,2038]" pageId="1" pageNumber="588">
|
||
<emphasis id="B91FEAA82B3EFFAEFF3654AFFE8E1FF7" bold="true" box="[151,309,1985,2008]" pageId="1" pageNumber="588">FIGURES 1–2</emphasis>
|
||
.
|
||
<taxonomicName id="4C6B4D392B3EFFAEFEE554AFFDE61FF6" box="[324,605,1985,2007]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="thailandica" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3EFFAEFEE554AFFDE61FF6" box="[324,605,1985,2007]" italics="true" pageId="1" pageNumber="588">Sangpradubina thailandica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91FEAA82B3EFFAEFDC554ACFD7E1FF6" bold="true" box="[612,709,1986,2007]" pageId="1" pageNumber="588">
|
||
<taxonomicNameLabel id="A22C57D32B3EFFAEFDC554ACFD7E1FF6" box="[612,709,1986,2007]" pageId="1" pageNumber="588" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<emphasis id="B91FEAA82B3EFFAEFD6F54ACFD5B1FF6" box="[718,736,1986,2007]" italics="true" pageId="1" pageNumber="588">et</emphasis>
|
||
<emphasis id="B91FEAA82B3EFFAEFD4954ACFC801FF6" bold="true" box="[744,827,1986,2007]" pageId="1" pageNumber="588">sp. nov.</emphasis>
|
||
(1) female mature nymph; (2) male mature nymph, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1466322B3DFFADFF3656EAFEE61D9B" httpUri="https://zenodo.org/record/272406/files/figure.png" pageId="2" pageNumber="589" targetBox="[340,1246,183,1391]" targetPageId="2">
|
||
<paragraph id="8BD436BA2B3DFFADFF3656EAFEE61D9B" blockId="2.[151,1436,1412,1466]" pageId="2" pageNumber="589">
|
||
<emphasis id="B91FEAA82B3DFFADFF3656EAFE8D1DBB" bold="true" box="[151,310,1412,1435]" pageId="2" pageNumber="589">FIGURES 3–4</emphasis>
|
||
.
|
||
<taxonomicName id="4C6B4D392B3DFFADFEE756EBFDE41DBA" box="[326,607,1413,1435]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="thailandica" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3DFFADFEE756EBFDE41DBA" box="[326,607,1413,1435]" italics="true" pageId="2" pageNumber="589">Sangpradubina thailandica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91FEAA82B3DFFADFDC956E8FD721DBA" bold="true" box="[616,713,1414,1435]" pageId="2" pageNumber="589">
|
||
<taxonomicNameLabel id="A22C57D32B3DFFADFDC956E8FD721DBA" box="[616,713,1414,1435]" pageId="2" pageNumber="589" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<emphasis id="B91FEAA82B3DFFADFD7256E8FD5E1DBA" box="[723,741,1414,1435]" italics="true" pageId="2" pageNumber="589">et</emphasis>
|
||
<emphasis id="B91FEAA82B3DFFADFD4E56E8FCF91DBA" bold="true" box="[751,834,1414,1435]" pageId="2" pageNumber="589">sp. nov.</emphasis>
|
||
(3-4) habitat of the nymphs, the Pachi River, Ratchaburi province, Thailand.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD436BA2B3DFFADFF66568FFE641E60" blockId="2.[151,1437,1505,2034]" pageId="2" pageNumber="589">
|
||
<emphasis id="B91FEAA82B3DFFADFF66568FFF4D1DDB" bold="true" box="[199,246,1505,1530]" pageId="2" pageNumber="589">Egg</emphasis>
|
||
. General shape elongated, chorionic surface with uniform longitudinal ridges, each ridge with distinct groove between ridges. Eggs with large KCT’s only on one pole, but absent from elsewhere (
|
||
<figureCitation id="13502A3F2B3DFFADFAA0556BFA2B1E3F" box="[1281,1424,1541,1566]" captionStart="FIGURES 36 – 41" captionStartId="10.[151,264,1935,1958]" captionTargetBox="[227,1360,625,1903]" captionTargetId="figure@10.[227,1361,625,1904]" captionTargetPageId="10" captionText="FIGURES 36 – 41. SEM pictures of eggs extracted from a female imago of Sangpradubina thailandica gen. nov. et sp. nov., (36) general outline; (37 – 40) detail of pole; abbreviation: KCT = Knob-coiled Thread; (41) detail of the chorionic structure." httpUri="https://zenodo.org/record/272416/files/figure.png" pageId="2" pageNumber="589">Figs. 36–41</figureCitation>
|
||
). Micropyle close to polar cap.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37165312B3DFFADFF665523FED41E8F" pageId="2" pageNumber="589" type="etymology">
|
||
<paragraph id="8BD436BA2B3DFFADFF665523FED41E8F" blockId="2.[151,1437,1505,2034]" pageId="2" pageNumber="589">
|
||
<emphasis id="B91FEAA82B3DFFADFF665523FEF51E47" bold="true" box="[199,334,1613,1638]" pageId="2" pageNumber="589">Etymology.</emphasis>
|
||
The genus name is an arbitrary combination of letter to honour Prof. Narumon Sangpradub (
|
||
<collectingRegion id="49AFF8582B3DFFADFAFD5523FF691EA8" country="Thailand" name="Khon Kaen" pageId="2" pageNumber="589">Khon Kaen</collectingRegion>
|
||
University) for her outstanding contributions to the ecology and taxonomy of aquatic insects in
|
||
<collectingCountry id="F37C762A2B3DFFADFAA1551FFADC1EAB" box="[1280,1383,1649,1674]" name="Thailand" pageId="2" pageNumber="589">Thailand</collectingCountry>
|
||
. The gender is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37165312B3DFFADFF6655D7FC6A1EF3" box="[199,977,1721,1746]" pageId="2" pageNumber="589" type="distribution">
|
||
<paragraph id="8BD436BA2B3DFFADFF6655D7FC6A1EF3" blockId="2.[151,1437,1505,2034]" box="[199,977,1721,1746]" pageId="2" pageNumber="589">
|
||
<emphasis id="B91FEAA82B3DFFADFF6655D7FEDB1EF3" bold="true" box="[199,352,1721,1746]" pageId="2" pageNumber="589">Distribution.</emphasis>
|
||
<collectingCountry id="F37C762A2B3DFFADFEC655D7FE701EF3" box="[359,459,1721,1746]" name="Thailand" pageId="2" pageNumber="589">Thailand</collectingCountry>
|
||
(
|
||
<collectingRegion id="49AFF8582B3DFFADFE7855D7FD791EF3" box="[473,706,1721,1746]" country="Thailand" name="Ratchaburi" pageId="2" pageNumber="589">Ratchaburi province</collectingRegion>
|
||
,
|
||
<collectingRegion id="49AFF8582B3DFFADFD6C55D7FC7E1EF3" box="[717,965,1721,1746]" country="Thailand" name="Chanthaburi" pageId="2" pageNumber="589">Chanthaburi province</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37165312B3DFFABFF6655B3FCA519A6" lastPageId="4" lastPageNumber="591" pageId="2" pageNumber="589" type="discussion">
|
||
<paragraph id="8BD436BA2B3DFFADFF6655B3FCC61F40" blockId="2.[151,1437,1505,2034]" pageId="2" pageNumber="589">
|
||
<emphasis id="B91FEAA82B3DFFADFF6655B3FDE01ED7" bold="true" box="[199,603,1757,1782]" pageId="2" pageNumber="589">Discussion. Nymphal affinities.</emphasis>
|
||
According to
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFC9155B3FC611ED7" author="Kluge" box="[816,986,1757,1782]" pageId="2" pageNumber="589" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge (2012)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C6B4D392B3DFFADFC5655B3FB1F1ED7" box="[1015,1188,1757,1782]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3DFFADFC5655B3FB1F1ED7" box="[1015,1188,1757,1782]" italics="true" pageId="2" pageNumber="589">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
clearly belong to Alatophlebomaxillata, Atalophlebolinguata and
|
||
<taxonomicName id="4C6B4D392B3DFFADFD15546FFCFA1F3B" box="[692,833,1793,1818]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg1 by the following characters: 1) maxillae with one proximal comb-shaped dentiseta (distal dentiseta lost); 2) lingua with a pair of lateral processes arising near base and pointed distally; 3) shape of gills 1 different from gills II–VII.
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3DFFACFF665403FE3D193C" blockId="2.[151,1437,1505,2034]" lastBlockId="3.[151,1437,151,2018]" lastPageId="3" lastPageNumber="590" pageId="2" pageNumber="589">
|
||
Based on gill morphology,
|
||
<taxonomicName id="4C6B4D392B3DFFADFE575403FD181FA7" box="[502,675,1901,1926]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3DFFADFE575403FD181FA7" box="[502,675,1901,1926]" italics="true" pageId="2" pageNumber="589">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
is more closely related to the
|
||
<taxonomicName id="4C6B4D392B3DFFADFC535403FBEC1FA7" box="[1010,1111,1901,1926]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3DFFADFC535403FBEC1FA7" box="[1010,1111,1901,1926]" italics="true" pageId="2" pageNumber="589">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
group (sensu
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFB555400FF6B1F8B" author="Grant" pageId="2" pageNumber="589" refString="Grant, P. M. & Peters, W. L. (1993) Description of four new genera of Thraulus group mayflies from the Eastern Hemisphere and redescription of Simothraulus and Chiusanophlebia (Ephemeroptera: Leptophlebiidae: Atalophlebiidae). Transactions of the American Entomological Society, 119, 131 - 168." type="journal article" year="1993">Grant & Peters 1993</bibRefCitation>
|
||
) than to other members of
|
||
<taxonomicName id="4C6B4D392B3DFFADFDE854FFFD6D1F8B" box="[585,726,1937,1962]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg1 (
|
||
<taxonomicName id="4C6B4D392B3DFFADFCBA54FFFBF31F8B" authority="Eaton 1881" authorityName="Eaton" authorityYear="1881" box="[795,1096,1937,1962]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3DFFADFCBA54FFFC1D1F8B" box="[795,934,1937,1962]" italics="true" pageId="2" pageNumber="589">Choroterpes</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFC1854FCFBF31F8B" author="Eaton" box="[953,1096,1938,1962]" pageId="2" pageNumber="589" refString="Eaton, A. E. (1881) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 17, 191 - 197." type="journal article" year="1881">Eaton 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B4D392B3DFFADFBC154FFFA2F1F8B" authority="Barnard 1932" authorityName="Barnard" authorityYear="1932" box="[1120,1428,1937,1962]" class="Insecta" family="Leptophlebiidae" genus="Euthraulus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3DFFADFBC154FFFB671F8B" box="[1120,1244,1937,1962]" italics="true" pageId="2" pageNumber="589">Euthraulus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFB4E54FFFA2F1F8B" author="Barnard" box="[1263,1428,1937,1962]" pageId="2" pageNumber="589" refString="Barnard, K. H. (1932) South African mayflies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. http: // dx. doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B91FEAA82B3DFFADFF3654DBFE8C1FEF" box="[151,311,1973,1998]" italics="true" pageId="2" pageNumber="589">Dilatognathus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFE9F54DBFE791FEF" author="Kluge" box="[318,450,1973,1998]" pageId="2" pageNumber="589" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge 2012</bibRefCitation>
|
||
,
|
||
<emphasis id="B91FEAA82B3DFFADFE7154DBFD2D1FEF" box="[464,662,1973,1998]" italics="true" pageId="2" pageNumber="589">Monochoroterpes</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFD3C54DBFC181FEF" author="Kluge" box="[669,931,1973,1998]" pageId="2" pageNumber="589" refString="Kluge, N. J. & Jacobus, L. M. (2015) Monochoroterpes, a replacement name for Monophyllus Kluge, 2012 (Insecta: Ephemeroptera), nec Monophyllus Leach, 1821 (Mammalia: Chiroptera). Zootaxa, 3946 (1), 150 - 150. http: // dx. doi. org / 10.11646 / zootaxa. 3946.1.11" type="journal article" year="2015">Kluge & Jacobus 2015</bibRefCitation>
|
||
). The
|
||
<taxonomicName id="4C6B4D392B3DFFADFC4954DBFBF61FEF" box="[1000,1101,1973,1998]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3DFFADFC4954DBFBF61FEF" box="[1000,1101,1973,1998]" italics="true" pageId="2" pageNumber="589">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
group currently encompasses seven genera, of which only three are known from the nymphal stage:
|
||
<taxonomicName id="4C6B4D392B3DFFADFC1C54B7FB411FD3" authority="Grant & Peters 1993" authorityName="Grant & Peters" authorityYear="1993" box="[957,1274,2009,2034]" class="Insecta" family="Leptophlebiidae" genus="Barba" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3DFFADFC1C54B7FBB81FD3" box="[957,1027,2009,2034]" italics="true" pageId="2" pageNumber="589">Barba</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3DFFADFBAD54B4FB411FD3" author="Grant" box="[1036,1274,2010,2034]" pageId="2" pageNumber="589" refString="Grant, P. M. & Peters, W. L. (1993) Description of four new genera of Thraulus group mayflies from the Eastern Hemisphere and redescription of Simothraulus and Chiusanophlebia (Ephemeroptera: Leptophlebiidae: Atalophlebiidae). Transactions of the American Entomological Society, 119, 131 - 168." type="journal article" year="1993">Grant & Peters 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B4D392B3DFFACFAA754B7FE391891" authority="Grant & Peters 1993" authorityName="Grant & Peters" authorityYear="1993" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" lastPageId="3" lastPageNumber="590" order="Ephemeroptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3DFFADFAA754B7FA201FD3" box="[1286,1435,2009,2034]" italics="true" pageId="2" pageNumber="589">Nonnullidens</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFF3653F6FE391891" author="Grant" box="[151,386,152,176]" pageId="3" pageNumber="590" refString="Grant, P. M. & Peters, W. L. (1993) Description of four new genera of Thraulus group mayflies from the Eastern Hemisphere and redescription of Simothraulus and Chiusanophlebia (Ephemeroptera: Leptophlebiidae: Atalophlebiidae). Transactions of the American Entomological Society, 119, 131 - 168." type="journal article" year="1993">Grant & Peters 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D392B3CFFACFE1B53F9FD171891" authority="Eaton 1881" authorityName="Eaton" authorityYear="1881" box="[442,684,151,176]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFE1B53F9FDA41891" box="[442,543,151,176]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFD8953F6FD171891" author="Eaton" box="[552,684,152,176]" pageId="3" pageNumber="590" refString="Eaton, A. E. (1881) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 17, 191 - 197." type="journal article" year="1881">Eaton 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
. The
|
||
<collectingCountry id="F37C762A2B3CFFACFD4D53F6FC381891" box="[748,899,151,176]" name="New Zealand" pageId="3" pageNumber="590">New Zealand</collectingCountry>
|
||
genus
|
||
<taxonomicName id="4C6B4D392B3CFFACFC7453F9FAE91891" authority="Towns & Peters 1979" authorityName="Towns & Peters" authorityYear="1979" box="[981,1362,151,176]" class="Insecta" family="Leptophlebiidae" genus="Isothraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFC7453F9FBEF1891" box="[981,1108,151,176]" italics="true" pageId="3" pageNumber="590">Isothraulus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFBFC53F6FAE91891" author="Towns" box="[1117,1362,152,176]" pageId="3" pageNumber="590" refString="Towns, D. R. & Peters, W. L. (1979) New genera and species of Leptophlebiidae (Ephemeroptera) from New Zealand. New Zealand Journal of Zoology, 6, 439 - 452. http: // dx. doi. org / 10.1080 / 03014223.1979.10428383" type="journal article" year="1979">Towns & Peters 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
might also belong to the
|
||
<taxonomicName id="4C6B4D392B3CFFACFEC253D2FE7318F4" box="[355,456,188,213]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFEC253D2FE7318F4" box="[355,456,188,213]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
lineage, but its phylogenetic position is not completely clear (
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFBDD53D2FAC918F4" author="Towns" box="[1148,1394,188,213]" pageId="3" pageNumber="590" refString="Towns, D. R. & Peters, W. L. (1996) Leptophlebiidae (Insecta: Ephemeroptera). Fauna of New Zealand, 36, 1 - 141." type="journal article" year="1996">Towns & Peters 1996</bibRefCitation>
|
||
). In peculiar
|
||
<taxonomicName id="4C6B4D392B3CFFACFF5853B1FEC218D9" box="[249,377,223,248]" class="Insecta" family="Leptophlebiidae" genus="Isothraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFF5853B1FEC218D9" box="[249,377,223,248]" italics="true" pageId="3" pageNumber="590">Isothraulus</emphasis>
|
||
</taxonomicName>
|
||
imaginal tarsi bear two hooked claws, while all
|
||
<taxonomicName id="4C6B4D392B3CFFACFC3C53B1FBB918D9" box="[925,1026,223,248]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFC3C53B1FBB918D9" box="[925,1026,223,248]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
-related genera bear one hooked and one paddle-like claw.
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3CFFACFF665249FD7B19D4" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="590">
|
||
Within the
|
||
<taxonomicName id="4C6B4D392B3CFFACFEE55249FE121961" box="[324,425,295,320]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFEE55249FE121961" box="[324,425,295,320]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
group, the genus
|
||
<taxonomicName id="4C6B4D392B3CFFACFDD25249FD631961" box="[627,728,295,320]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFDD25249FD631961" box="[627,728,295,320]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
is the most rich in species, with a total of 15 species described from the Palaearctic (3), Afrotropical (3) and Oriental (9) regions. The genera
|
||
<taxonomicName id="4C6B4D392B3CFFACFB8C5222FBC81944" box="[1069,1139,332,357]" class="Insecta" family="Leptophlebiidae" genus="Barba" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB8C5222FBC81944" box="[1069,1139,332,357]" italics="true" pageId="3" pageNumber="590">Barba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B91FEAA82B3CFFACFB105222FAF61944" box="[1201,1357,332,357]" italics="true" pageId="3" pageNumber="590">
|
||
<taxonomicName id="4C6B4D392B3CFFACFB105222FAF21944" box="[1201,1353,332,357]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Nonnullidens</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which encompass 7 species together, have been reported from
|
||
<collectingCountry id="F37C762A2B3CFFACFCA7521EFC5A19A9" box="[774,993,367,392]" name="Papua New Guinea" pageId="3" pageNumber="590">Papua New Guinea</collectingCountry>
|
||
, and recently
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFBDD5201FAA919A9" author="Kluge" box="[1148,1298,367,392]" pageId="3" pageNumber="590" refString="Kluge, N. J. (2013) Mayflies of the genus Nonnullidens Grant & Peters 1993 (Ephemeroptera: Leptophlebiidae) from New Guinea. Zootaxa, 3722 (1), 33 - 49. http: // dx. doi. org / 10.11646 / zootaxa. 3722.1.3" type="journal article" year="2013">Kluge (2013)</bibRefCitation>
|
||
proposed to consider
|
||
<taxonomicName id="4C6B4D392B3CFFACFEA552FAFEF1198C" box="[260,330,404,429]" class="Insecta" family="Leptophlebiidae" genus="Barba" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFEA552FAFEF1198C" box="[260,330,404,429]" italics="true" pageId="3" pageNumber="590">Barba</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName id="4C6B4D392B3CFFACFD2952FAFCA6198C" box="[648,797,404,429]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFD2952FAFCA6198C" box="[648,797,404,429]" italics="true" pageId="3" pageNumber="590">Nonnullidens</emphasis>
|
||
</taxonomicName>
|
||
. It is our opinion that this synonymy needs further investigation, because study of our extensive material from
|
||
<collectingCountry id="F37C762A2B3CFFACFCCB52D6FBF419F1" box="[874,1103,439,464]" name="Papua New Guinea" pageId="3" pageNumber="590">Papua New Guinea</collectingCountry>
|
||
reveals that some nymphal characters could allow separating the two genera.
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3CFFACFF66516EFDAF1C14" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="590">
|
||
<materialsCitation id="3B033CE72B3CFFACFF66516EFB521A41" ID-GBIF-Occurrence="1313880553" country="Thailand" county="Timor" location="Borneo" municipality="Sumatra" pageId="3" pageNumber="590" specimenCount="1" typeStatus="holotype">
|
||
We compared the nymph of
|
||
<taxonomicName id="4C6B4D392B3CFFACFDAD516EFD121A39" box="[524,681,511,536]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="thailandica">
|
||
<emphasis id="B91FEAA82B3CFFACFDAD516EFD121A39" box="[524,681,511,536]" italics="true" pageId="3" pageNumber="590">S. thailandica</emphasis>
|
||
</taxonomicName>
|
||
to those of several species of
|
||
<taxonomicName id="4C6B4D392B3CFFACFBA55291FBD21A39" box="[1028,1129,511,536]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFBA55291FBD21A39" box="[1028,1129,511,536]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="4C6B4D392B3CFFACFB4A5291FE991A1D" authority="Eaton, 1881" authorityName="Eaton" authorityYear="1881" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="bellus">
|
||
<emphasis id="B91FEAA82B3CFFACFB4A5291FA271A39" box="[1259,1436,511,536]" italics="true" pageId="3" pageNumber="590">Thraulus bellus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFF36514BFE991A1D" author="Eaton" box="[151,290,548,572]" pageId="3" pageNumber="590" refString="Eaton, A. E. (1881) An announcement of new genera of the Ephemeridae. Entomologist's Monthly Magazine, 17, 191 - 197." type="journal article" year="1881">Eaton, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54D088182B3CFFACFE95514AFEDD1A1D" box="[308,358,548,572]" pageId="3" pageNumber="590">type</typeStatus>
|
||
species of the genus), several unnamed species from
|
||
<collectingCountry id="F37C762A2B3CFFACFC7C514AFBF81A1C" box="[989,1091,548,573]" name="Thailand" pageId="3" pageNumber="590">Thailand</collectingCountry>
|
||
,
|
||
<location id="8EB460612B3CFFACFBF3514BFB1C1A1D" LSID="urn:lsid:plazi:treatment:03C287AC2B3EFFABFF3653F9FCA519A6:8EB460612B3CFFACFBF3514BFB1C1A1D" box="[1106,1191,549,572]" country="Thailand" county="Timor" municipality="Sumatra" name="Borneo" pageId="3" pageNumber="590">Borneo</location>
|
||
,
|
||
<collectingMunicipality id="6BB0ACC02B3CFFACFB17514AFAAD1A1D" box="[1206,1302,548,572]" pageId="3" pageNumber="590">Sumatra</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62B54E362B3CFFACFA85514AFAD21A1C" box="[1316,1385,548,573]" pageId="3" pageNumber="590">Timor</collectingCounty>
|
||
and
|
||
<collectingCountry id="F37C762A2B3CFFACFF365126FECA1A41" box="[151,369,583,608]" name="Papua New Guinea" pageId="3" pageNumber="590">Papua New Guinea</collectingCountry>
|
||
, as well as several species of
|
||
<taxonomicName id="4C6B4D392B3CFFACFD1B5129FCBB1A41" box="[698,768,583,608]" class="Insecta" family="Leptophlebiidae" genus="Barba" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFD1B5129FCBB1A41" box="[698,768,583,608]" italics="true" pageId="3" pageNumber="590">Barba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D392B3CFFACFC945129FC711A41" box="[821,970,583,608]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFC945129FC711A41" box="[821,970,583,608]" italics="true" pageId="3" pageNumber="590">Nonnullidens</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F37C762A2B3CFFACFBAF5126FB521A41" box="[1038,1257,583,608]" name="Papua New Guinea" pageId="3" pageNumber="590">Papua New Guinea</collectingCountry>
|
||
</materialsCitation>
|
||
. Our new genus present unique characters among the
|
||
<taxonomicName id="4C6B4D392B3CFFACFD995102FD261AA4" box="[568,669,620,645]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFD995102FD261AA4" box="[568,669,620,645]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
lineage, such as: 1) maxillae with few pectinate setae (ca. 11–13) in the subapical row and with well-developed anterolateral projection; 2) tarsal claws with 4 denticles at the base and 9 near the apex; 3) gill 1 comprised only of a single and slender lamella; 4) cerci and terminal filament bearing additional long, thin setae. The mandibles with only a tuft of setae in the middle of the lateral margin separate our new genus also from
|
||
<taxonomicName id="4C6B4D392B3CFFACFE305192FE6C1B34" box="[401,471,764,789]" class="Insecta" family="Leptophlebiidae" genus="Barba" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFE305192FE6C1B34" box="[401,471,764,789]" italics="true" pageId="3" pageNumber="590">Barba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D392B3CFFACFDB55192FD121B34" box="[532,681,764,789]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFDB55192FD121B34" box="[532,681,764,789]" italics="true" pageId="3" pageNumber="590">Nonnullidens</emphasis>
|
||
</taxonomicName>
|
||
. The nymph of
|
||
<taxonomicName id="4C6B4D392B3CFFACFCC95192FBAE1B34" box="[872,1045,764,789]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFCC95192FBAE1B34" box="[872,1045,764,789]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
appears to be more closely allied with
|
||
<taxonomicName id="4C6B4D392B3CFFACFF705071FEAC1B19" box="[209,279,799,824]" class="Insecta" family="Leptophlebiidae" genus="Barba" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFF705071FEAC1B19" box="[209,279,799,824]" italics="true" pageId="3" pageNumber="590">Barba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6B4D392B3CFFACFEEE5071FE5F1B19" box="[335,484,799,824]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFEEE5071FE5F1B19" box="[335,484,799,824]" italics="true" pageId="3" pageNumber="590">Nonnullidens</emphasis>
|
||
</taxonomicName>
|
||
than
|
||
<emphasis id="B91FEAA82B3CFFACFD855071FD2B1B19" box="[548,656,799,824]" italics="true" pageId="3" pageNumber="590">
|
||
<taxonomicName id="4C6B4D392B3CFFACFD855071FD371B19" box="[548,652,799,824]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Thraulus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
based on gill II–VII morphology, where the fimbriate part is located in the distal half (present on the whole margin in
|
||
<taxonomicName id="4C6B4D392B3CFFACFD62502AFC931B7C" box="[707,808,836,861]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFD62502AFC931B7C" box="[707,808,836,861]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C6B4D392B3CFFACFCE1502AFC561B7C" box="[832,1005,836,861]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFCE1502AFC561B7C" box="[832,1005,836,861]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
gill 1 clearly differs from those of the other three genera, with the notable exception of
|
||
<taxonomicName id="4C6B4D392B3CFFACFD1A5009FB301BA1" authority="Li, Liu & Zhou 2006" authorityName="Li, Liu & Zhou" authorityYear="2006" box="[699,1163,871,896]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="femoratus">
|
||
<emphasis id="B91FEAA82B3CFFACFD1A5009FC2D1BA1" box="[699,918,871,896]" italics="true" pageId="3" pageNumber="590">Thraulus femoratus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFC3C5009FB301BA1" author="Li" box="[925,1163,871,896]" pageId="3" pageNumber="590" refString="Li, P., Liu, D. & Zhou, C. - F. (2006) A new species of Thraulus from Nanjing (Eastern China) with single first gill (Insecta: Ephemeroptera: Leptophlebiidae). Zoological Science, 23, 641 - 645. http: // dx. doi. org / 10.2108 / zsj. 23.641" type="journal article" year="2006">Li, Liu & Zhou 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, from China, which also possesses a single and slender lamella, but in which all other characters clearly fit the genus
|
||
<taxonomicName id="4C6B4D392B3CFFACFB6650E2FF671BE9" authority="Li et al. 2006" authorityName="Li et al." authorityYear="2006" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB6650E2FA971B84" box="[1223,1324,908,933]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFA9F50E2FF6F1BE9" author="Li" pageId="3" pageNumber="590" refString="Li, P., Liu, D. & Zhou, C. - F. (2006) A new species of Thraulus from Nanjing (Eastern China) with single first gill (Insecta: Ephemeroptera: Leptophlebiidae). Zoological Science, 23, 641 - 645. http: // dx. doi. org / 10.2108 / zsj. 23.641" type="journal article" year="2006">
|
||
Li
|
||
<emphasis id="B91FEAA82B3CFFACFAC350E3FA271B84" box="[1378,1436,908,933]" italics="true" pageId="3" pageNumber="590">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The first gill of most
|
||
<taxonomicName id="4C6B4D392B3CFFACFE7950C1FD861BE9" box="[472,573,943,968]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFE7950C1FD861BE9" box="[472,573,943,968]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
species is comprised of long and slender dorsal and ventral lamellae, except for
|
||
<taxonomicName id="4C6B4D392B3CFFACFF1E50BAFCB41BCC" authority="Grant & Sivamarakrishnan, 1985" authorityName="Grant & Sivamarakrishnan" authorityYear="1985" box="[191,783,980,1005]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="gopalani">
|
||
<emphasis id="B91FEAA82B3CFFACFF1E50BAFE341BCC" box="[191,399,980,1005]" italics="true" pageId="3" pageNumber="590">Thraulus gopalani</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFE3750BAFCB41BCC" author="Grant" box="[406,783,980,1005]" pageId="3" pageNumber="590" refString="Grant, P. M. & Sivaramakrishnan, K. G. (1985) A new species of Thraulus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Southern India. The Florida Entomologist, 68 (3), 424 - 432. http: // dx. doi. org / 10.2307 / 3495129" type="journal article" year="1985">Grant & Sivamarakrishnan, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, from India, and
|
||
<taxonomicName id="4C6B4D392B3CFFACFC7150BBFB5C1BCC" authority="Gillies, 1964" authorityName="Gillies" authorityYear="1964" box="[976,1255,980,1005]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="torrentis">
|
||
<emphasis id="B91FEAA82B3CFFACFC7150BBFBF01BCD" box="[976,1099,981,1004]" italics="true" pageId="3" pageNumber="590">T. torrentis</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFBF250BAFB5C1BCC" author="Gillies" box="[1107,1255,980,1005]" pageId="3" pageNumber="590" refString="Peters, W. L., Gillies, M. T. & Edmunds, G. F. Jr. (1964) Two new genera of mayflies from the Ethiopian and Oriental Regions (Ephemeroptera: Leptophlebiidae). Proceedings of the Royal Entomological Society of London (B), 33 (7 - 8), 117 - 124. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1964. tb 01623. x" type="journal article" year="1964">Gillies, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Tanzania, (
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFF3F5096FED71C31" author="Peters" box="[158,364,1015,1040]" pageId="3" pageNumber="590" refString="Peters, W. L., Gillies, M. T. & Edmunds, G. F. Jr. (1964) Two new genera of mayflies from the Ethiopian and Oriental Regions (Ephemeroptera: Leptophlebiidae). Proceedings of the Royal Entomological Society of London (B), 33 (7 - 8), 117 - 124. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1964. tb 01623. x" type="journal article" year="1964">
|
||
Peters
|
||
<emphasis id="B91FEAA82B3CFFACFF4F5097FE921C31" box="[238,297,1015,1040]" italics="true" pageId="3" pageNumber="590">et al.</emphasis>
|
||
1964
|
||
</bibRefCitation>
|
||
) which have a dorsal lanceolate portion and a ventral fimbriate portion (
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFB155096FA2C1C31" author="Peters" box="[1204,1431,1015,1040]" pageId="3" pageNumber="590" refString="Peters, W. L. & Tsui, P. T. P. (1972) New species of Thraulus from Asia (Leptophlebiidae: Ephemeroptera). Oriental Insects, 6, 1 - 17. http: // dx. doi. org / 10.1080 / 00305316.1972.10434048" type="journal article" year="1972">Peters & Tsui 1972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFF365772FDB31C14" author="Grant" box="[151,520,1052,1077]" pageId="3" pageNumber="590" refString="Grant, P. M. & Sivaramakrishnan, K. G. (1985) A new species of Thraulus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Southern India. The Florida Entomologist, 68 (3), 424 - 432. http: // dx. doi. org / 10.2307 / 3495129" type="journal article" year="1985">Grant & Sivaramakrishnan 1985</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3CFFACFF665751FBE51D74" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="590">
|
||
<materialsCitation id="3B033CE72B3CFFACFF665751FBE51D74" ID-GBIF-Occurrence="1313880556" country="Papua New Guinea" county="In" location="The" municipality="western Pacific Ocean" pageId="3" pageNumber="590" specimenCount="7" specimenCount-intercalary="7">
|
||
<emphasis id="B91FEAA82B3CFFACFF665751FE1D1C79" bold="true" box="[199,422,1087,1112]" pageId="3" pageNumber="590">Imaginal affinities.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C6B4D392B3CFFACFD865751FD691C79" box="[551,722,1087,1112]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFD865751FD691C79" box="[551,722,1087,1112]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
differs from other genera of the
|
||
<taxonomicName id="4C6B4D392B3CFFACFB9D5751FB1B1C79" box="[1084,1184,1087,1112]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB9D5751FB1B1C79" box="[1084,1184,1087,1112]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
group by the presence of
|
||
<specimenCount id="9D6DFD332B3CFFACFF15570AFE851C5C" box="[180,318,1124,1149]" pageId="3" pageNumber="590" type="intercalary">5 intercalary</specimenCount>
|
||
veins in the cubital field, whereas other genera possess only
|
||
<specimenCount id="9D6DFD332B3CFFACFC7B570AFBC21C5C" box="[986,1145,1124,1149]" pageId="3" pageNumber="590" type="intercalary">2 intercalaries</specimenCount>
|
||
, including those for which nymphs remain unknown, such as
|
||
<taxonomicName id="4C6B4D392B3CFFACFDB657E9FC871C81" authority="Ulmer, 1939" authorityName="Ulmer" authorityYear="1939" box="[535,828,1159,1184]" class="Insecta" family="Leptophlebiidae" genus="Simothraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFDB657E9FD101C81" box="[535,683,1159,1184]" italics="true" pageId="3" pageNumber="590">Simothraulus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFD1357E9FC871C81" author="Ulmer" box="[690,828,1159,1184]" pageId="3" pageNumber="590" refString="Ulmer, G. (1939) Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln. Archiv fur Hydrobiologie, Supplement 16, 443 - 692." type="book chapter" year="1939">Ulmer, 1939</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6B4D392B3CFFACFCEA57E9FBD41C81" authority="Grant & Peters, 1993" authorityName="Grant & Peters" authorityYear="1993" box="[843,1135,1159,1184]" class="Insecta" family="Leptophlebiidae" genus="Sulu" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFCEA57E9FCC71C81" box="[843,892,1159,1184]" italics="true" pageId="3" pageNumber="590">Sulu</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFC2257E6FBD41C81" author="Grant" box="[899,1135,1160,1184]" pageId="3" pageNumber="590" refString="Grant, P. M. & Peters, W. L. (1993) Description of four new genera of Thraulus group mayflies from the Eastern Hemisphere and redescription of Simothraulus and Chiusanophlebia (Ephemeroptera: Leptophlebiidae: Atalophlebiidae). Transactions of the American Entomological Society, 119, 131 - 168." type="journal article" year="1993">Grant & Peters, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C6B4D392B3CFFACFB3557E9FF691CE4" authority="Ueno, 1969" authorityName="Ueno" authorityYear="1969" class="Insecta" family="Leptophlebiidae" genus="Chiusanophlebia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB3557E9FAEA1C81" box="[1172,1361,1159,1184]" italics="true" pageId="3" pageNumber="590">Chiusanophlebia</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFAF857E6FF691CE4" author="Ueno" pageId="3" pageNumber="590" refString="Ueno, M. (1969) Mayflies (Ephemeroptera) from various regions of Southeast Asia. Oriental Insects, 3, 221 - 238. http: // dx. doi. org / 10.1080 / 00305316.1969.10433911" type="journal article" year="1969">Uéno, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<location id="8EB460612B3CFFACFF7D57C2FEBC1CE4" LSID="urn:lsid:plazi:treatment:03C287AC2B3EFFABFF3653F9FCA519A6:8EB460612B3CFFACFF7D57C2FEBC1CE4" box="[220,263,1196,1221]" country="Papua New Guinea" county="In" municipality="western Pacific Ocean" name="The" pageId="3" pageNumber="590">The</location>
|
||
shape of the hind wing also separates
|
||
<taxonomicName id="4C6B4D392B3CFFACFD1557C2FCE41CE4" box="[692,863,1196,1221]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFD1557C2FCE41CE4" box="[692,863,1196,1221]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
easily from the enigmatic genus
|
||
<taxonomicName id="4C6B4D392B3CFFACFB7057C2FE861CC9" authority="Grant & Peters, 1993" authorityName="Grant & Peters" authorityYear="1993" class="Insecta" family="Leptophlebiidae" genus="Magnilobus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB7057C2FAEE1CE4" box="[1233,1365,1196,1221]" italics="true" pageId="3" pageNumber="590">Magnilobus</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFAFC57C2FE861CC9" author="Grant" pageId="3" pageNumber="590" refString="Grant, P. M. & Peters, W. L. (1993) Description of four new genera of Thraulus group mayflies from the Eastern Hemisphere and redescription of Simothraulus and Chiusanophlebia (Ephemeroptera: Leptophlebiidae: Atalophlebiidae). Transactions of the American Entomological Society, 119, 131 - 168." type="journal article" year="1993">Grant & Peters, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is known only from a female subimago from an island in the
|
||
<collectingCountry id="F37C762A2B3CFFACFB8F57A1FA931CC9" box="[1070,1320,1231,1256]" name="Papua New Guinea" pageId="3" pageNumber="590">Bismarck Archipelago</collectingCountry>
|
||
(northeast of
|
||
<location id="8EB460612B3CFFACFF17579BFEF81D2C" LSID="urn:lsid:plazi:treatment:03C287AC2B3EFFABFF3653F9FCA519A6:8EB460612B3CFFACFF17579BFEF81D2C" box="[182,323,1268,1293]" country="Papua New Guinea" county="In" municipality="western Pacific Ocean" name="New Guinea" pageId="3" pageNumber="590">New Guinea</location>
|
||
in the
|
||
<collectingMunicipality id="6BB0ACC02B3CFFACFE32579BFD281D2D" box="[403,659,1268,1293]" pageId="3" pageNumber="590">western Pacific Ocean</collectingMunicipality>
|
||
).
|
||
<collectingCounty id="62B54E362B3CFFACFD07579BFD061D2D" box="[678,701,1269,1292]" pageId="3" pageNumber="590">In</collectingCounty>
|
||
the male imaginal stage, all other genera of the
|
||
<taxonomicName id="4C6B4D392B3CFFACFB41579AFAFF1D2C" box="[1248,1348,1268,1293]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB41579AFAFF1D2C" box="[1248,1348,1268,1293]" italics="true" pageId="3" pageNumber="590">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
lineage have penes lobes elongate, with the widest part generally at the apex (more rarely in the middle), whereas in
|
||
<taxonomicName id="4C6B4D392B3CFFACFF365652FEF91D74" box="[151,322,1340,1365]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFF365652FEF91D74" box="[151,322,1340,1365]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
, the penes lobes are more stocky, with the widest part being at the base.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3CFFACFF665631FB681FC3" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="590">
|
||
The puzzling thing is that the
|
||
<taxonomicName id="4C6B4D392B3CFFACFD9E5631FD571D59" box="[575,748,1375,1400]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFD9E5631FD571D59" box="[575,748,1375,1400]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
male imago closely resembles those of
|
||
<taxonomicName id="4C6B4D392B3CFFACFB7A5631FADD1D59" box="[1243,1382,1375,1400]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B91FEAA82B3CFFACFB7A5631FADD1D59" box="[1243,1382,1375,1400]" italics="true" pageId="3" pageNumber="590">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3CFFACFAD55631FA2D1D59" box="[1396,1430,1375,1400]" pageId="3" pageNumber="590" sensu="lato">s.l.</taxonomicNameLabel>
|
||
, especially with regards to venation of the cubital field of the wing, and general shape of male genitalia. When using the key provided in
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFEDF56C6FD121DE1" author="Peters" box="[382,681,1447,1472]" pageId="3" pageNumber="590" refString="Peters, W. L. & Edmunds, G. F. Jr. (1970) Revision of the generic classification of the Eastern Hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970">Peters & Edmunds (1970)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C6B4D392B3CFFACFD1656C9FCDF1DE1" box="[695,868,1447,1472]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFD1656C9FCDF1DE1" box="[695,868,1447,1472]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
will key to
|
||
<taxonomicName id="4C6B4D392B3CFFACFC5056C9FA951DE1" authority="Gillies, 1951" authorityName="Gillies" authorityYear="1951" box="[1009,1326,1447,1472]" class="Insecta" family="Leptophlebiidae" genus="Cryptopenella" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFC5056C9FB2A1DE1" box="[1009,1169,1447,1472]" italics="true" pageId="3" pageNumber="590">Cryptopenella</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFB3856C9FA951DE1" author="Gillies" box="[1177,1326,1447,1472]" pageId="3" pageNumber="590" refString="Gillies, M. T. (1951) Further notes on Ephemeroptera from India and South East Asia. Proceedings of the Royal Entomological Society of London (B), 20, 121 - 130. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.1951. tb 01009. x" type="journal article" year="1951">Gillies, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is now considered as a subgenus of
|
||
<taxonomicName id="4C6B4D392B3CFFACFD8656A2FCEA1DC4" authority="Zhou 2006" authorityName="Zhou" authorityYear="2006" box="[551,849,1484,1509]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFD8656A2FD091DC4" box="[551,690,1484,1509]" italics="true" pageId="3" pageNumber="590">Choroterpes</emphasis>
|
||
(Zhou 2006)
|
||
</taxonomicName>
|
||
, or even a synonym of the
|
||
<taxonomicName id="4C6B4D392B3CFFACFB3C56A2FA931DC4" box="[1181,1320,1484,1509]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFB3C56A2FA931DC4" box="[1181,1320,1484,1509]" italics="true" pageId="3" pageNumber="590">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
subgenus
|
||
<taxonomicName id="4C6B4D392B3CFFACFF365681FDB51E29" authority="Kang & Yang 1994" authorityName="Kang & Yang" authorityYear="1994" box="[151,526,1519,1544]" class="Insecta" family="Leptophlebiidae" genus="Euthraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFF365681FEA81E29" box="[151,275,1519,1544]" italics="true" pageId="3" pageNumber="590">Euthraulus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFE82569EFDBD1E29" author="Kang" box="[291,518,1520,1544]" pageId="3" pageNumber="590" refString="Kang, S. C. & Yang, C. T. (1994) Leptophlebiidae of Taiwan (Ephemeroptera). Journal of Taiwan Museum, 47 (1), 57 - 82." type="journal article" year="1994">Kang & Yang 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Our new genus however greatly differs from
|
||
<taxonomicName id="4C6B4D392B3CFFACFBBC5681FB061E29" box="[1053,1213,1519,1544]" class="Insecta" family="Leptophlebiidae" genus="Cryptopenella" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFBBC5681FB061E29" box="[1053,1213,1519,1544]" italics="true" pageId="3" pageNumber="590">Cryptopenella</emphasis>
|
||
</taxonomicName>
|
||
species because, in the latter, the penes lobes are almost completely hidden under the styliger plate (Zhou 2006,
|
||
<figureCitation id="13502A3F2B3CFFACFAA0557AFA2B1E0D" box="[1281,1424,1556,1581]" captionStart-0="FIGURES 19 – 22" captionStart-1="FIGURES 23 – 25" captionStart-2="FIGURES 26 – 31" captionStartId-0="7.[151,264,1479,1502]" captionStartId-1="8.[151,264,929,952]" captionStartId-2="8.[151,264,1904,1927]" captionTargetBox-0="[151,1427,209,1443]" captionTargetBox-1="[327,1240,200,890]" captionTargetBox-2="[320,1230,1023,1872]" captionTargetId-0="figure@7.[151,1436,195,1458]" captionTargetId-1="figure@8.[306,1278,193,897]" captionTargetId-2="figure@8.[282,1254,1023,1882]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="8" captionText-0="FIGURES 19 – 22. Mature nymph of Sangpradubina thailandica gen. nov. et sp. nov., 19) foreleg; (20) middle leg; (21) hindleg; (22) foreleg claw." captionText-1="FIGURES 23 – 25. Mature nymph of Sangpradubina thailandica gen. nov. et sp. nov., (23) gill I; (24) gill IV; (25) gill VII." captionText-2="FIGURES 26 – 31. Male imago of Sangpradubina thailandica gen. nov. et sp. nov., (26) forewing; (27) hindwing; (28) foreleg; (29) middle leg; (30) hindleg; (31) tarsal claw." httpUri-0="https://zenodo.org/record/272412/files/figure.png" httpUri-1="https://zenodo.org/record/272413/files/figure.png" httpUri-2="https://zenodo.org/record/272414/files/figure.png" pageId="3" pageNumber="590">Figs. 21–28</figureCitation>
|
||
).
|
||
<taxonomicName id="4C6B4D392B3CFFACFF365559FEFF1E71" box="[151,324,1591,1616]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFF365559FEFF1E71" box="[151,324,1591,1616]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
male adults also differ from those of
|
||
<taxonomicName id="4C6B4D392B3CFFACFCAF5559FC221E71" box="[782,921,1591,1616]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B91FEAA82B3CFFACFCAF5559FC221E71" box="[782,921,1591,1616]" italics="true" pageId="3" pageNumber="590">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3CFFACFC055556FC731E6E" box="[932,968,1592,1615]" pageId="3" pageNumber="590" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C6B4D392B3CFFACFBA95559FB3F1E71" box="[1032,1156,1591,1616]" class="Insecta" family="Leptophlebiidae" genus="Euthraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFBA95559FB3F1E71" box="[1032,1156,1591,1616]" italics="true" pageId="3" pageNumber="590">Euthraulus</emphasis>
|
||
</taxonomicName>
|
||
by the vein MP2 being independent of vein MP1, and the styliger plate margin being strongly convex.
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFBE35530FB661E56" author="Kluge" box="[1090,1245,1630,1655]" pageId="3" pageNumber="590" refString="Kluge, N. J. (2016) Phylogeny of Ephemeroptera. Available from: http: // www. insecta. bio. spbu. ru (accessed 1 June 2016)" type="journal article" year="2016">Kluge (2016)</bibRefCitation>
|
||
considers as an autapomorphy of
|
||
<taxonomicName id="4C6B4D392B3CFFACFEC055EBFE511EBF" box="[353,490,1669,1694]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg2 (
|
||
<taxonomicName id="4C6B4D392B3CFFACFD8955EBFD081EBF" box="[552,691,1669,1694]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B91FEAA82B3CFFACFD8955EBFD081EBF" box="[552,691,1669,1694]" italics="true" pageId="3" pageNumber="590">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3CFFACFD1D55EBFD601EBF" box="[700,731,1669,1694]" pageId="3" pageNumber="590" sensu="lato">s.l.</taxonomicNameLabel>
|
||
) the presence of what he calls a projection distal to the penes gonopore, and
|
||
<taxonomicName id="4C6B4D392B3CFFACFEFB55C7FDBC1EE3" box="[346,519,1705,1730]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFEFB55C7FDBC1EE3" box="[346,519,1705,1730]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
also possesses this character (
|
||
<figureCitation id="13502A3F2B3CFFACFC3E55C7FBBA1EE3" box="[927,1025,1705,1730]" captionStart="FIGURES 32 – 35" captionStartId="9.[151,264,1514,1537]" captionTargetBox="[185,1386,193,1471]" captionTargetId="figure@9.[185,1401,193,1493]" captionTargetPageId="9" captionText="FIGURES 32 – 35. Imago of Sangpradubina thailandica gen. nov. et sp. nov., (32) photograph of male genitalia, ventral view, abbreviation: gp = gonopore; (33) male genitalia, ventral view; (34) male genitalia, lateral view; (35) female last sternites, ventral view." httpUri="https://zenodo.org/record/272415/files/figure.png" pageId="3" pageNumber="590">Fig. 32</figureCitation>
|
||
). Therefore, one could consider
|
||
<taxonomicName id="4C6B4D392B3CFFACFF3655A3FEFF1EC7" box="[151,324,1741,1766]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFF3655A3FEFF1EC7" box="[151,324,1741,1766]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
as belonging to
|
||
<taxonomicName id="4C6B4D392B3CFFACFDB455A3FD1B1EC7" box="[533,672,1741,1766]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B91FEAA82B3CFFACFDB455A3FD1B1EC7" box="[533,672,1741,1766]" italics="true" pageId="3" pageNumber="590">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3CFFACFD0C55A3FD751EC7" box="[685,718,1741,1766]" pageId="3" pageNumber="590" sensu="lato">s.l.</taxonomicNameLabel>
|
||
However, inclusion of this genus in
|
||
<taxonomicName id="4C6B4D392B3CFFACFB3255A3FAA71EC7" box="[1171,1308,1741,1766]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg2 would destabilize the present concept of this monophyletic group by invalidating two of the three proposed apomorphies for the group, namely gill shape and egg papillae. Species placed in
|
||
<taxonomicName id="4C6B4D392B3CFFACFC3E547BFB931F0F" box="[927,1064,1813,1838]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg2 never possess dense rows of setae on the outer margins of fore- and middle legs, which indicate that
|
||
<taxonomicName id="4C6B4D392B3CFFACFC425457FB2B1F73" box="[995,1168,1849,1874]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFC425457FB2B1F73" box="[995,1168,1849,1874]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
may also be related to
|
||
<taxonomicName id="4C6B4D392B3CFFACFF365433FDB61F57" authority="Peters & Edmunds, 1970" authorityName="Peters & Edmunds" authorityYear="1970" box="[151,525,1885,1910]" class="Insecta" family="Leptophlebiidae" genus="Indialis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFF365433FF561F57" box="[151,237,1885,1910]" italics="true" pageId="3" pageNumber="590">Indialis</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFF555430FDB61F57" author="Peters" box="[244,525,1885,1910]" pageId="3" pageNumber="590" refString="Peters, W. L. & Edmunds, G. F. Jr. (1970) Revision of the generic classification of the Eastern Hemisphere Leptophlebiidae (Ephemeroptera). Pacific Insects, 12, 157 - 240." type="journal article" year="1970">Peters & Edmunds, 1970</bibRefCitation>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C6B4D392B3CFFACFD955433FC601F57" authority="Sivaramakrishnan, 1985" authorityName="Sivaramakrishnan" authorityYear="1985" box="[564,987,1885,1910]" class="Insecta" family="Leptophlebiidae" genus="Edmundsula" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3CFFACFD955433FD7B1F57" box="[564,704,1885,1910]" italics="true" pageId="3" pageNumber="590">Edmundsula</emphasis>
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFD695433FC601F57" author="Sivaramakrishnan" box="[712,987,1885,1910]" pageId="3" pageNumber="590" refString="Grant, P. M. & Sivaramakrishnan, K. G. (1985) A new species of Thraulus (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Southern India. The Florida Entomologist, 68 (3), 424 - 432. http: // dx. doi. org / 10.2307 / 3495129" type="journal article" year="1985">Sivaramakrishnan, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, two genera endemic to Southern
|
||
<collectingCountry id="F37C762A2B3CFFACFAFA5433FA231F57" box="[1371,1432,1885,1910]" name="India" pageId="3" pageNumber="590">India</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C6B4D392B3CFFACFF3654EFFEFF1FBB" box="[151,324,1921,1946]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFF3654EFFEFF1FBB" box="[151,324,1921,1946]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
differs from these by the shape of the gills (each lamella lanceolate and not fringed as in the latter’s). In our opinion,
|
||
<taxonomicName id="4C6B4D392B3CFFACFE0C54CBFDE11F9F" box="[429,602,1957,1982]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3CFFACFE0C54CBFDE11F9F" box="[429,602,1957,1982]" italics="true" pageId="3" pageNumber="590">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
probably belongs to
|
||
<taxonomicName id="4C6B4D392B3CFFACFCEA54CBFC631F9F" box="[843,984,1957,1982]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg
|
||
<quantity id="4C939B5F2B3CFFACFC5954CBFBA41F9F" box="[1016,1055,1957,1982]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="590" unit="in" value="1.0">1 in</quantity>
|
||
Kluge’s (2016) scheme, and may have rank equal to
|
||
<taxonomicName id="4C6B4D392B3CFFACFECD54A7FE4C1FC3" box="[364,503,1993,2018]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Choroterpes</taxonomicName>
|
||
/fg2,
|
||
<taxonomicName id="4C6B4D392B3CFFACFD9154A7FD311FC3" box="[560,650,1993,2018]" class="Insecta" family="Leptophlebiidae" genus="Indialis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Indialis</taxonomicName>
|
||
/g1,
|
||
<taxonomicName id="4C6B4D392B3CFFACFD1754A7FCA51FC3" box="[694,798,1993,2018]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">Thraulus</taxonomicName>
|
||
/g1 and
|
||
<taxonomicName id="4C6B4D392B3CFFACFCD254A7FBB41FC3" box="[883,1039,1993,2018]" class="Insecta" family="Leptophlebiidae" genus="Nonnullidens" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="599" phylum="Arthropoda" rank="genus">Nonnullidens</taxonomicName>
|
||
/g1 (
|
||
<bibRefCitation id="EFFA4B4B2B3CFFACFB9C54A7FB7C1FC3" author="Kluge" box="[1085,1223,1993,2018]" pageId="3" pageNumber="590" refString="Kluge, N. J. (2016) Phylogeny of Ephemeroptera. Available from: http: // www. insecta. bio. spbu. ru (accessed 1 June 2016)" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD436BA2B3BFFABFF6653F9FCA519A6" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="591">
|
||
The egg chorionic structure of
|
||
<taxonomicName id="4C6B4D392B3BFFABFD8953F9FD6E1891" box="[552,725,151,176]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="588" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B91FEAA82B3BFFABFD8953F9FD6E1891" box="[552,725,151,176]" italics="true" pageId="4" pageNumber="591">Sangpradubina</emphasis>
|
||
</taxonomicName>
|
||
also appears to be very peculiar. The general shape resembles that of many
|
||
<taxonomicName id="4C6B4D392B3BFFABFE8C53D2FE2918F4" box="[301,402,188,213]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91FEAA82B3BFFABFE8C53D2FE2918F4" box="[301,402,188,213]" italics="true" pageId="4" pageNumber="591">Thraulus</emphasis>
|
||
</taxonomicName>
|
||
species, being elongate with longitudinal ridges. They differ from those of
|
||
<taxonomicName id="4C6B4D392B3BFFABFB4853D2FACF18F4" box="[1257,1396,188,213]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="genus" sensu="lato">
|
||
<emphasis id="B91FEAA82B3BFFABFB4853D2FACF18F4" box="[1257,1396,188,213]" italics="true" pageId="4" pageNumber="591">Choroterpes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22C57D32B3BFFABFADA53D2FA2718F4" box="[1403,1436,188,213]" pageId="4" pageNumber="591" sensu="lato">s.l.</taxonomicNameLabel>
|
||
by the absence of papillae connected by ridges, in the form of a flower-like protuberance (
|
||
<bibRefCitation id="EFFA4B4B2B3BFFABFB3C53B1FF0C193D" author="Kluge" pageId="4" pageNumber="591" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">
|
||
Kluge 2012,
|
||
<figureCitation id="13502A3F2B3BFFABFA9353B1FF0C193D" captionStart-0="FIGURES 19 – 22" captionStart-1="FIGURES 23 – 25" captionStartId-0="7.[151,264,1479,1502]" captionStartId-1="8.[151,264,929,952]" captionTargetBox-0="[151,1427,209,1443]" captionTargetBox-1="[327,1240,200,890]" captionTargetId-0="figure@7.[151,1436,195,1458]" captionTargetId-1="figure@8.[306,1278,193,897]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 19 – 22. Mature nymph of Sangpradubina thailandica gen. nov. et sp. nov., 19) foreleg; (20) middle leg; (21) hindleg; (22) foreleg claw." captionText-1="FIGURES 23 – 25. Mature nymph of Sangpradubina thailandica gen. nov. et sp. nov., (23) gill I; (24) gill IV; (25) gill VII." httpUri-0="https://zenodo.org/record/272412/files/figure.png" httpUri-1="https://zenodo.org/record/272413/files/figure.png" pageId="4" pageNumber="591">Figs. 20– 24</figureCitation>
|
||
</bibRefCitation>
|
||
). The presence of large KCT’s on one pole is almost unheard of in the family
|
||
<taxonomicName id="4C6B4D392B3BFFABFBD7526AFA97193C" box="[1142,1324,260,285]" class="Insecta" family="Leptophlebiidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="family">Leptophlebiidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFA4B4B2B3BFFABFAE1526AFEF81961" author="Koss" pageId="4" pageNumber="591" refString="Koss, R. W. & Edmunds, G. F. Jr. (1974) Ephemeroptera eggs and their contribution to phylogenetic studies of the order. Zoological Journal of the Linnean Society, 55 (4), 267 - 349. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1974. tb 01648. x" type="journal article" year="1974">Koss & Edmunds 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFA4B4B2B3BFFABFEEE5249FD361961" author="Dominguez" box="[335,653,295,320]" pageId="4" pageNumber="591" refString="Dominguez, E. & Cuezzo, M. G. (2002) Ephemeroptera egg chorion characters: a test of their importance in assessing phylogenetic relationships. Journal of Morphology, 253, 148 - 165. http: // dx. doi. org / 10.1002 / jmor. 1117" type="journal article" year="2002">Dominguez & Cuezzo 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFA4B4B2B3BFFABFD385249FC8E1961" author="Li" box="[665,821,295,320]" pageId="4" pageNumber="591" refString="Li, P., Liu, D. & Zhou, C. - F. (2006) A new species of Thraulus from Nanjing (Eastern China) with single first gill (Insecta: Ephemeroptera: Leptophlebiidae). Zoological Science, 23, 641 - 645. http: // dx. doi. org / 10.2108 / zsj. 23.641" type="journal article" year="2006">
|
||
Li
|
||
<emphasis id="B91FEAA82B3BFFABFD185247FD491961" box="[697,754,295,320]" italics="true" pageId="4" pageNumber="591">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
), but this condition has been reported by
|
||
<bibRefCitation id="EFFA4B4B2B3BFFABFAA75249FA271961" author="Kluge" box="[1286,1436,295,320]" pageId="4" pageNumber="591" refString="Kluge, N. J. (2012) Contribution to the knowledge of Choroterpes (Ephemeroptera, Leptophlebiidae). Russian Entomological Journal, 21 (3), 273 - 306." type="journal article" year="2012">Kluge (2012)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="4C6B4D392B3BFFABFF1F5222FDC01945" box="[190,635,332,357]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="mercatorius" subGenus="Choroterpes">
|
||
<emphasis id="B91FEAA82B3BFFABFF1F5222FDC01945" box="[190,635,332,357]" italics="true" pageId="4" pageNumber="591">Choroterpes (Choroterpes) mercatorius</emphasis>
|
||
</taxonomicName>
|
||
from Sulawesi. We note that in
|
||
<taxonomicName id="4C6B4D392B3BFFABFC455223FB311945" box="[996,1162,333,356]" class="Insecta" family="Leptophlebiidae" genus="Choroterpes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="mercatorius">
|
||
<emphasis id="B91FEAA82B3BFFABFC455223FB311945" box="[996,1162,333,356]" italics="true" pageId="4" pageNumber="591">C. mercatorius</emphasis>
|
||
</taxonomicName>
|
||
the papillae are present, contrary to the absence of such papillae on
|
||
<taxonomicName id="4C6B4D392B3BFFABFDDD521EFCAC19A9" box="[636,791,367,392]" class="Insecta" family="Leptophlebiidae" genus="Sangpradubina" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="thailandica">
|
||
<emphasis id="B91FEAA82B3BFFABFDDD521EFCAC19A9" box="[636,791,367,392]" italics="true" pageId="4" pageNumber="591">S. thailandica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |