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<document id="2A6388378F4796ADBD556422B6E96678" ID-DOI="10.11646/zootaxa.3847.1.4" ID-GBIF-Dataset="c2754097-3bc5-445d-8b98-56ce18cd608e" ID-ISSN="1175-5326" ID-Zenodo-Dep="231549" ID-ZooBank="4C151005-01FC-4503-961A-0A8CBB2F263E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460469941492" checkinUser="plazi" docAuthor="Gnezdilov, Vladimir M., Thierry, Bourgoin &amp; Soulier-Perkins, Adeline" docDate="2014" docId="03C28797B15CFFCBFF50F89EFA8EFD2C" docLanguage="en" docName="zt03847p096.pdf" docOrigin="Zootaxa 3847 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thabena litaoensis Yang 1994" docType="treatment" docVersion="9" masterDocId="FFFBFFEFB154FFC6FFC7FFEBFFC7FFDC" masterDocTitle="Vietnamese Issidae (Hemiptera, Fulgoroidea): new taxa, new records and new distribution data" updateTime="1698305520964" updateUser="plazi">
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<mods:title id="0EB644E05393596C8AF5B2BE56E4B28A">Vietnamese Issidae (Hemiptera, Fulgoroidea): new taxa, new records and new distribution data</mods:title>
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<mods:namePart id="FE50C133D2D597B0E50E86DC0C007F81">Gnezdilov, Vladimir M.</mods:namePart>
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<mods:namePart id="9D1F4446BFB9CF6D08BD842AC298208C">Thierry, Bourgoin</mods:namePart>
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<mods:namePart id="2E3794120EBCD7D1410828F3A019DDA7">Soulier-Perkins, Adeline</mods:namePart>
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<mods:title id="F073365D59D4AAE2CF441588648FC6DC">Zootaxa</mods:title>
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<mods:date id="5565C412195B842102201BE95248AC3F">2014</mods:date>
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<mods:number id="C6D4FB097154551D7F0DA9B6E02ABC00">3847</mods:number>
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<treatment id="03C28797B15CFFCBFF50F89EFA8EFD2C" ID-DOI="http://doi.org/10.5281/zenodo.5697968" ID-GBIF-Taxon="119514667" ID-Zenodo-Dep="5697968" LSID="urn:lsid:plazi:treatment:03C28797B15CFFCBFF50F89EFA8EFD2C" httpUri="http://treatment.plazi.org/id/03C28797B15CFFCBFF50F89EFA8EFD2C" lastPageId="13" pageId="8">
<subSubSection id="C371650AB15CFFCEFF50F89EFDCCF853" box="[151,523,1909,1935]" pageId="8" type="nomenclature">
<paragraph id="8BD43681B15CFFCEFF50F89EFDCCF853" blockId="8.[151,523,1909,1935]" box="[151,523,1909,1935]" pageId="8">
<heading id="D09C81EDB15CFFCEFF50F89EFDCCF853" bold="true" box="[151,523,1909,1935]" fontSize="11" level="1" pageId="8" reason="1">
<taxonomicName id="4C6B4D02B15CFFCEFF50F89EFDCCF853" ID-CoL="55YJ6" authority="Yang, 1994" authorityName="Yang" authorityYear="1994" box="[151,523,1909,1935]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="8" phylum="Arthropoda" rank="species" species="litaoensis">
<emphasis id="B91FEA93B15CFFCEFF50F89EFDCCF853" bold="true" box="[151,523,1909,1935]" pageId="8">
<emphasis id="B91FEA93B15CFFCEFF50F89EFEBBF853" bold="true" box="[151,380,1909,1935]" italics="true" pageId="8">Thabena litaoensis</emphasis>
<bibRefCitation id="EFFA4B70B15CFFCEFE45F89EFDCCF853" author="Yang" box="[386,523,1909,1935]" pageId="8" refString="Chan, M. - L. &amp; Yang, C. - T. (1994) Issidae of Taiwan (Homoptera: Fulgoroidea). Chen Chung Book, Taichung, Taiwan, 188 pp." type="book" year="1994">Yang, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C371650AB15CFFCEFF50F846FBF2F83E" pageId="8" type="reference_group">
<paragraph id="8BD43681B15CFFCEFF50F846FBF2F83E" blockId="8.[151,1436,1965,2018]" pageId="8">
<treatmentCitationGroup id="AB7B11AFB15CFFCEFF50F846FBF2F83E" pageId="8">
<taxonomicName id="4C6B4D02B15CFFCEFF50F846FDDEF81F" ID-CoL="3FHLN" authority="Yang, 1994: 90" authorityName="Yang" authorityPageNumber="90" authorityYear="1994" box="[151,537,1965,1987]" class="Insecta" family="Issidae" genus="Gelastyrella" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" phylum="Arthropoda" rank="species" species="litaoensis">
<emphasis id="B91FEA93B15CFFCEFF50F846FEBDF81F" box="[151,378,1965,1987]" italics="true" pageId="8">Gelastyrella litaoensis</emphasis>
<treatmentCitation id="0ACA1090B15CFFCEFE47F845FDDEF81F" author="Yang" box="[384,537,1965,1987]" page="90" pageId="8" year="1994">
<bibRefCitation id="EFFA4B70B15CFFCEFE47F845FE30F81F" author="Yang" box="[384,503,1965,1987]" pageId="8" refString="Chan, M. - L. &amp; Yang, C. - T. (1994) Issidae of Taiwan (Homoptera: Fulgoroidea). Chen Chung Book, Taichung, Taiwan, 188 pp." type="book" year="1994">Yang, 1994</bibRefCitation>
: 90
</treatmentCitation>
</taxonomicName>
(in
<bibRefCitation id="EFFA4B70B15CFFCEFD86F846FCC9F81F" author="Chan" box="[577,782,1965,1987]" pageId="8" refString="Chan, M. - L. &amp; Yang, C. - T. (1994) Issidae of Taiwan (Homoptera: Fulgoroidea). Chen Chung Book, Taichung, Taiwan, 188 pp." type="book" year="1994">Chan &amp; Yang, 1994</bibRefCitation>
).
<taxonomicName id="4C6B4D02B15CFFCEFCD9F846FACFF81F" authority="Ran et Liang, 2006: 65" authorityName="Ran et Liang" authorityPageNumber="65" authorityYear="2006" box="[798,1288,1965,1987]" class="Insecta" family="Issidae" genus="Gelastyrella" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B91FEA93B15CFFCEFCD9F846FBDEF81F" box="[798,1049,1965,1987]" italics="true" pageId="8">Gelastyrella hainanensis</emphasis>
Ran et Liang, 2006: 65
</taxonomicName>
, figs 18,
<emphasis id="B91FEA93B15CFFCEFAB6F846FF1CF83E" bold="true" pageId="8">syn. n.</emphasis>
<taxonomicName id="4C6B4D02B15CFFCEFF25F827FDBFF83E" ID-CoL="55YJ6" authority="Gnezdilov, 2009: 80" authorityName="Gnezdilov" authorityPageNumber="80" authorityYear="2009" box="[226,632,1996,2018]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="8" phylum="Arthropoda" rank="species" species="litaoensis">
<emphasis id="B91FEA93B15CFFCEFF25F827FE66F83E" box="[226,417,1996,2018]" italics="true" pageId="8">Thabena litaoensis</emphasis>
<treatmentCitation id="0ACA1090B15CFFCEFE60F827FDBFF83E" author="Gnezdilov" box="[423,632,1996,2018]" page="80" pageId="8" year="2009">
<bibRefCitation id="EFFA4B70B15CFFCEFE60F827FD95F83E" author="Gnezdilov" box="[423,594,1996,2018]" pageId="8" refString="Gnezdilov, V. M. (2009) Revisionary notes on some tropical Issidae and Nogodinidae (Hemiptera: Fulgoroidea). Acta Entomologica Musei Nationalis Pragae, 49 (1), 75 - 92." type="journal article" year="2009">Gnezdilov, 2009</bibRefCitation>
: 80
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="4C6B4D02B15CFFCEFD45F827FBF6F83E" ID-CoL="55YJ3" authority="Gnezdilov, 2009: 80" authorityName="Gnezdilov" authorityPageNumber="80" authorityYear="2009" box="[642,1073,1996,2018]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="8" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B91FEA93B15CFFCEFD45F827FC9DF83E" box="[642,858,1996,2018]" italics="true" pageId="8">Thabena hainanensis</emphasis>
<treatmentCitation id="0ACA1090B15CFFCEFCA7F827FBF6F83E" author="Gnezdilov" box="[864,1073,1996,2018]" page="80" pageId="8" year="2009">
<bibRefCitation id="EFFA4B70B15CFFCEFCA7F827FBCCF83E" author="Gnezdilov" box="[864,1035,1996,2018]" pageId="8" refString="Gnezdilov, V. M. (2009) Revisionary notes on some tropical Issidae and Nogodinidae (Hemiptera: Fulgoroidea). Acta Entomologica Musei Nationalis Pragae, 49 (1), 75 - 92." type="journal article" year="2009">Gnezdilov, 2009</bibRefCitation>
: 80
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF146609B15DFFCFFF50F8D1FD09F851" httpUri="https://zenodo.org/record/231555/files/figure.png" pageId="9" targetBox="[151,1436,193,1828]" targetPageId="9">
<paragraph id="8BD43681B15DFFCFFF50F8D1FD09F851" blockId="9.[151,1436,1850,1933]" pageId="9">
<emphasis id="B91FEA93B15DFFCFFF50F8D1FE92F88C" bold="true" box="[151,341,1850,1872]" pageId="9">FIGURES 1521.</emphasis>
<taxonomicName id="4C6B4D02B15DFFCFFE9AF8D1FD8CF88C" box="[349,587,1850,1872]" class="Insecta" family="Issidae" genus="Dactylissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="9" phylum="Arthropoda" rank="species" species="armillarius">
<emphasis id="B91FEA93B15DFFCFFE9AF8D1FD8CF88C" box="[349,587,1850,1872]" italics="true" pageId="9">Dactylissus armillarius</emphasis>
</taxonomicName>
<emphasis id="B91FEA93B15DFFCFFD93F8D1FD19F88C" bold="true" box="[596,734,1850,1872]" pageId="9">gen. et sp. n.</emphasis>
, paratype, male genitalia. 15—penis, in ventral view; 16—penis, in lateral view; 17—penis, in dorsal view; 18—style, in lateral view; 19—capitulum of style, in dorsal view; 20—pygofer and anal tube, in lateral view; 21—anal tube, in dorsal view.
</paragraph>
</caption>
<caption id="DF146609B15EFFCCFF50F84FFCB8F805" httpUri="https://zenodo.org/record/231556/files/figure.png" pageId="10" targetBox="[226,1373,208,1913]" targetPageId="10">
<paragraph id="8BD43681B15EFFCCFF50F84FFCB8F805" blockId="10.[151,1436,1956,2009]" pageId="10">
<emphasis id="B91FEA93B15EFFCCFF50F84FFE9CF866" bold="true" box="[151,347,1956,1978]" pageId="10">FIGURES 2225.</emphasis>
<taxonomicName id="4C6B4D02B15EFFCCFEA1F84FFE09F866" box="[358,462,1956,1978]" class="Insecta" family="Issidae" genus="Darwallia" kingdom="Animalia" order="Hemiptera" pageId="10" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEA93B15EFFCCFEA1F84FFE09F866" box="[358,462,1956,1978]" italics="true" pageId="10">Darwallia</emphasis>
</taxonomicName>
spp. 22—
<taxonomicName id="4C6B4D02B15EFFCCFD86F84EFD72F866" box="[577,693,1956,1978]" class="Insecta" family="Issidae" genus="Darwallia" kingdom="Animalia" order="Hemiptera" pageId="10" phylum="Arthropoda" rank="species" species="barbata" status="sp. nov.">
<emphasis id="B91FEA93B15EFFCCFD86F84EFD72F866" box="[577,693,1956,1978]" italics="true" pageId="10">D. barbata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57E8B15EFFCCFD07F84FFD39F866" box="[704,766,1956,1978]" pageId="10" rank="species">
<emphasis id="B91FEA93B15EFFCCFD07F84FFD39F866" bold="true" box="[704,766,1956,1978]" pageId="10">sp. n.</emphasis>
</taxonomicNameLabel>
, holotype, female, in dorsal view; 23—same, in frontal view; 24—same, in lateral view; 25—
<taxonomicName id="4C6B4D02B15EFFCCFE1CF82FFD5BF805" authority="Walker" authorityName="Walker" box="[475,668,1987,2009]" class="Insecta" family="Issidae" genus="Darwallia" kingdom="Animalia" order="Hemiptera" pageId="10" phylum="Arthropoda" rank="species" species="patula">
<emphasis id="B91FEA93B15EFFCCFE1CF82FFDFCF805" box="[475,571,1987,2009]" italics="true" pageId="10">D. patula</emphasis>
(Walker)
</taxonomicName>
, male, in frontal view.
</paragraph>
</caption>
<caption id="DF146609B15FFFCDFF50F84EFF09F806" httpUri="https://zenodo.org/record/231557/files/figure.png" pageId="11" targetBox="[219,1384,213,1932]" targetPageId="11">
<paragraph id="8BD43681B15FFFCDFF50F84EFF09F806" blockId="11.[151,1436,1957,2010]" pageId="11">
<emphasis id="B91FEA93B15FFFCDFF50F84EFE92F867" bold="true" box="[151,341,1957,1979]" pageId="11">FIGURES 2629.</emphasis>
<taxonomicName id="4C6B4D02B15FFFCDFE9CF84EFD80F867" box="[347,583,1957,1979]" class="Insecta" family="Issidae" genus="Dactylissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="11" phylum="Arthropoda" rank="species" species="armillarius">
<emphasis id="B91FEA93B15FFFCDFE9CF84EFD80F867" box="[347,583,1957,1979]" italics="true" pageId="11">Dactylissus armillarius</emphasis>
</taxonomicName>
<emphasis id="B91FEA93B15FFFCDFD8AF84EFD14F867" bold="true" box="[589,723,1957,1979]" pageId="11">gen. et sp. n.</emphasis>
, paratypes, males. 26, 27—dorsal view; 28—fronatl view; 29—lateral view.
</paragraph>
</caption>
<caption id="DF146609B158FFCAFF50F84AFEB2F80A" httpUri="https://zenodo.org/record/231558/files/figure.png" pageId="12" targetBox="[258,1322,193,1918]" targetPageId="12">
<paragraph id="8BD43681B158FFCAFF50F84AFEB2F80A" blockId="12.[151,1436,1953,2006]" pageId="12">
<emphasis id="B91FEA93B158FFCAFF50F84AFE92F86B" bold="true" box="[151,341,1953,1975]" pageId="12">FIGURES 3033.</emphasis>
<taxonomicName id="4C6B4D02B158FFCAFE9BF84AFD8DF86B" box="[348,586,1953,1975]" class="Insecta" family="Issidae" genus="Dactylissus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="12" phylum="Arthropoda" rank="species" species="armillarius">
<emphasis id="B91FEA93B158FFCAFE9BF84AFD8DF86B" box="[348,586,1953,1975]" italics="true" pageId="12">Dactylissus armillarius</emphasis>
</taxonomicName>
<emphasis id="B91FEA93B158FFCAFD95F84AFD25F86B" bold="true" box="[594,738,1953,1975]" pageId="12">gen. et sp. n.,</emphasis>
paratypes, females. 30—dorsal view; 31—fronatl view; 32—lateral view; 33—ovipositor.
</paragraph>
</caption>
<subSubSection id="C371650AB159FFCBFF50FF7CFAFCFF24" pageId="13" type="materials_examined">
<paragraph id="8BD43681B159FFCBFF50FF7CFAFCFF24" blockId="13.[151,1437,151,752]" pageId="13">
<emphasis id="B91FEA93B159FFCBFF50FF7CFEBCFF6C" bold="true" box="[151,379,151,176]" pageId="13">Material examined</emphasis>
(
<collectionCode id="ED7AAE44B159FFCBFE4BFF73FE2FFF6C" LSID="urn:lsid:biocol.org:col:34988" box="[396,488,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13">MNHN</collectionCode>
).
<collectingCountry id="F37C7611B159FFCBFE3BFF7CFD98FF6C" box="[508,607,151,176]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
:
<specimenCount id="9D6DFD08B159FFCBFDABFF73FD49FF6C" box="[620,654,152,176]" pageId="13" type="generic">1♂</specimenCount>
1♀, “Museum Paris,
<collectingCountry id="F37C7611B159FFCBFC4DFF7CFC1BFF6C" box="[906,988,151,176]" name="Vietnam" pageId="13">Tonkin</collectingCountry>
, Hoa Binh, J. de Cooman, R. Oberthur 1919”;
<specimenCount id="9D6DFD08B159FFCBFF2DFF57FEC9FF08" box="[234,270,188,212]" pageId="13" type="generic">2♂</specimenCount>
2♀, “Museum Paris,
<collectingCountry id="F37C7611B159FFCBFDC6FF57FD94FF09" box="[513,595,188,213]" name="Vietnam" pageId="13">Tonkin</collectingCountry>
, rég. de Hoa Binh, A. de Cooman 1926”; 2♀, “Museum Paris,
<collectingCountry id="F37C7611B159FFCBFAD2FF57FAA0FF09" box="[1301,1383,188,213]" name="Vietnam" pageId="13">Tonkin</collectingCountry>
, rég. de Hoa Binh, A. de Cooman 1928”; 1♀, “Museum Paris,
<collectingCountry id="F37C7611B159FFCBFCDBFF34FCA9FF24" box="[796,878,223,248]" name="Vietnam" pageId="13">Tonkin</collectingCountry>
, rég. de Hoa Binh, A. de Cooman 1929”.
</paragraph>
</subSubSection>
<subSubSection id="C371650AB159FFCBFF00FEEFFA8EFD2C" pageId="13" type="distribution">
<paragraph id="8BD43681B159FFCBFF00FEEFFE53FE9C" blockId="13.[151,1437,151,752]" pageId="13">
<emphasis id="B91FEA93B159FFCBFF00FEEFFEA7FEC1" bold="true" box="[199,352,260,285]" pageId="13">Distribution.</emphasis>
<collectingCountry id="F37C7611B159FFCBFEACFEEFFE69FEC1" box="[363,430,260,285]" name="China" pageId="13">China</collectingCountry>
(Hainan) (
<bibRefCitation id="EFFA4B70B159FFCBFDE9FEEFFCE5FEC1" author="Ran" box="[558,802,260,285]" pageId="13" refString="Ran, H. - F. &amp; Liang, A. - P. (2006 a) The issid genus Gelastyrella Yang (Hemiptera: Fulgoroidea: Issidae) from China. Zootaxa, 1238, 63 - 68." type="journal article" year="2006" yearSuffix="a">Ran &amp; Liang, 2006a</bibRefCitation>
;
<bibRefCitation id="EFFA4B70B159FFCBFCF6FEEFFBE2FEC0" author="Zhang" box="[817,1061,260,285]" pageId="13" refString="Zhang, Z. - G. &amp; Chen, X. - S. (2012) A review of the genus Thabena Stal (Hemiptera: Fulgoromorpha: Issidae) from China with description of one new species. Entomotaxonomia, 34 (2), 227 - 232." type="journal article" year="2012">Zhang &amp; Chen 2012</bibRefCitation>
),
<collectingCountry id="F37C7611B159FFCBFBFBFEEFFB49FEC1" box="[1084,1166,260,285]" name="Taiwan" pageId="13">Taiwan</collectingCountry>
(
<bibRefCitation id="EFFA4B70B159FFCBFB67FEEFFA57FEC0" author="Chan" box="[1184,1424,260,285]" pageId="13" refString="Chan, M. - L. &amp; Yang, C. - T. (1994) Issidae of Taiwan (Homoptera: Fulgoroidea). Chen Chung Book, Taichung, Taiwan, 188 pp." type="book" year="1994">Chan &amp; Yang, 1994</bibRefCitation>
),
<collectingCountry id="F37C7611B159FFCBFF50FECCFF31FE9C" box="[151,246,295,320]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
(new record).
</paragraph>
<paragraph id="8BD43681B159FFCBFF00FEA7FE0EFDE1" blockId="13.[151,1437,151,752]" pageId="13">
<emphasis id="B91FEA93B159FFCBFF00FEA7FEC1FEB9" bold="true" box="[199,262,332,357]" pageId="13">Note.</emphasis>
The drawings of male genitalia of
<taxonomicName id="4C6B4D02B159FFCBFD57FEA6FCD0FEB9" box="[656,791,332,357]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="13" phylum="Arthropoda" rank="species" species="litaoensis">
<emphasis id="B91FEA93B159FFCBFD57FEA6FCD0FEB9" box="[656,791,332,357]" italics="true" pageId="13">T. litaoensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6B4D02B159FFCBFC89FEA6FC28FEB9" box="[846,1007,332,357]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="13" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B91FEA93B159FFCBFC89FEA6FC28FEB9" box="[846,1007,332,357]" italics="true" pageId="13">T. hainanensis</emphasis>
</taxonomicName>
were compared and the species found conspecific as the structure of penis is very peculiar according to the cap-shaped apical lobe and very different from other species of the genus. Diagnostic characters mentioned for
<taxonomicName id="4C6B4D02B159FFCBFC05FE7EFBA0FE71" box="[962,1127,404,429]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="13" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B91FEA93B159FFCBFC05FE7EFBA0FE71" box="[962,1127,404,429]" italics="true" pageId="13">T. hainanensis</emphasis>
</taxonomicName>
in the original description (Ran &amp; Liang, 2006) are only the coloration of coryphe (vertex) and the number of apical spines on hind tibia. These charactres are more variable in comparison with male genitalia and may be treated as interspecific variation between the island populations. Finally both names were mentioned from Hainan (
<bibRefCitation id="EFFA4B70B159FFCBFBE8FE14FAD1FDC4" author="Zhang" box="[1071,1302,511,536]" pageId="13" refString="Zhang, Z. - G. &amp; Chen, X. - S. (2012) A review of the genus Thabena Stal (Hemiptera: Fulgoromorpha: Issidae) from China with description of one new species. Entomotaxonomia, 34 (2), 227 - 232." type="journal article" year="2012">Zhang &amp; Chen 2012</bibRefCitation>
) which also confirms our point of view.
</paragraph>
<paragraph id="8BD43681B159FFCBFF00FDACFA8EFD2C" blockId="13.[151,1437,151,752]" pageId="13">
The genus
<taxonomicName id="4C6B4D02B159FFCBFE95FDACFE72FDBC" box="[338,437,583,608]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="13" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEA93B159FFCBFE95FDACFE72FDBC" box="[338,437,583,608]" italics="true" pageId="13">Thabena</emphasis>
</taxonomicName>
is Indo-Malayan and currently groups 14 species distributed in
<collectingCountry id="F37C7611B159FFCBFB08FDACFAD2FDBC" box="[1231,1301,583,608]" name="China" pageId="13">China</collectingCountry>
,
<collectingCountry id="F37C7611B159FFCBFAEFFDACFA5FFDBC" box="[1320,1432,583,608]" name="Indonesia" pageId="13">Indonesia</collectingCountry>
,
<collectingCountry id="F37C7611B159FFCBFF50FD87FEC6FD59" box="[151,257,620,645]" name="Malaysia" pageId="13">Malaysia</collectingCountry>
,
<collectingCountry id="F37C7611B159FFCBFECBFD87FE4AFD59" box="[268,397,620,645]" name="Philippines" pageId="13">Philippines</collectingCountry>
,
<collectingCountry id="F37C7611B159FFCBFE5EFD87FDCAFD59" box="[409,525,620,645]" name="Singapore" pageId="13">Singapore</collectingCountry>
,
<collectingCountry id="F37C7611B159FFCBFDDEFD87FDA9FD59" box="[537,622,620,645]" name="Taiwan" pageId="13">Taiwan</collectingCountry>
, and
<collectingCountry id="F37C7611B159FFCBFD6DFD87FCC9FD59" box="[682,782,620,645]" name="Thailand" pageId="13">Thailand</collectingCountry>
(
<bibRefCitation id="EFFA4B70B159FFCBFCDAFD87FC1CFD58" author="Gnezdilov" box="[797,987,620,645]" pageId="13" refString="Gnezdilov, V. M. (2009) Revisionary notes on some tropical Issidae and Nogodinidae (Hemiptera: Fulgoroidea). Acta Entomologica Musei Nationalis Pragae, 49 (1), 75 - 92." type="journal article" year="2009">Gnezdilov, 2009</bibRefCitation>
,
<bibRefCitation id="EFFA4B70B159FFCBFC20FD87FBF5FD59" author="Gnezdilov" box="[999,1074,620,645]" pageId="13" refString="Gnezdilov, V. M. (2013 b) Modern classification and the distribution of the family Issidae Spinola (Homoptera, Auchenorrhyncha: Fulgoroidea). Entomologicheskoe Obozrenie, 92 (4), 724 - 738. [in Russian with English summary]" type="journal article" year="2013" yearSuffix="b">2013b</bibRefCitation>
).
<taxonomicName id="4C6B4D02B159FFCBFB83FD87FE64FD74" authority="Bonfils, Attie et Reynaud, 2001" authorityName="Bonfils, Attie et Reynaud" authorityYear="2001" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="13" phylum="Arthropoda" rank="species" species="brunnifrons">
<emphasis id="B91FEA93B159FFCBFB83FD87FAF6FD59" box="[1092,1329,620,645]" italics="true" pageId="13">Thabena brunnifrons</emphasis>
(Bonfils, Attié et Reynaud, 2001)
</taxonomicName>
described from
<collectingCountry id="F37C7611B159FFCBFD9FFD64FD70FD74" box="[600,695,655,680]" name="Réunion" pageId="13">Réunion</collectingCountry>
was probably imported from Asia to this island (
<bibRefCitation id="EFFA4B70B159FFCBFB1EFD64FA52FD74" author="Gnezdilov" box="[1241,1429,655,680]" pageId="13" refString="Gnezdilov, V. M. (2009) Revisionary notes on some tropical Issidae and Nogodinidae (Hemiptera: Fulgoroidea). Acta Entomologica Musei Nationalis Pragae, 49 (1), 75 - 92." type="journal article" year="2009">Gnezdilov, 2009</bibRefCitation>
) and
<taxonomicName id="4C6B4D02B159FFCBFF0BFD5FFDF0FD11" authority="Fieber, 1876" authorityName="Fieber" authorityYear="1876" box="[204,567,692,717]" class="Insecta" family="Issidae" genus="Thabena" kingdom="Animalia" order="Hemiptera" pageId="13" phylum="Arthropoda" rank="species" species="fissala">
<emphasis id="B91FEA93B159FFCBFF0BFD5FFE43FD11" box="[204,388,692,717]" italics="true" pageId="13">Thabena fissala</emphasis>
(Fieber, 1876)
</taxonomicName>
described from
<collectingCountry id="F37C7611B159FFCBFCC7FD5FFC98FD11" box="[768,863,692,717]" name="Portugal" pageId="13">Portugal</collectingCountry>
by Fieber might be due to an error of locality (
<bibRefCitation id="EFFA4B70B159FFCBFF59FD3CFE5AFD2C" author="Gnezdilov" box="[158,413,727,752]" pageId="13" refString="Gnezdilov, V. M., Soulier-Perkins, A. &amp; Bourgoin, T. (2011) Fieber's original drawings and their corresponding types for the family Issidae (Hemiptera, Fulgoromorpha) in the Museum national d'Histoire naturelle of Paris, France. Zootaxa, 2806, 24 - 34." type="journal article" year="2011">
Gnezdilov
<emphasis id="B91FEA93B159FFCBFEDBFD32FE93FD2C" box="[284,340,727,752]" italics="true" pageId="13">et al.</emphasis>
, 2011
</bibRefCitation>
). As it could have been expected, this genus is now found also in northern
<collectingCountry id="F37C7611B159FFCBFB24FD3CFA81FD2C" box="[1251,1350,727,752]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>