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<document id="822F4E65D4C117D02595C2F6DDF6645A" ID-DOI="10.11646/zootaxa.3986.2.4" ID-GBIF-Dataset="99ab0619-d0df-4186-a2be-5198484ca988" ID-ISSN="1175-5326" ID-Zenodo-Dep="245930" ID-ZooBank="02CCFD53-C564-4DEF-86B5-21B0951768EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461262132840" checkinUser="plazi" docAuthor="Hoese, Douglass F. &amp; Allen, Gerald R." docDate="2015" docId="03C28793FFCAFFBEFF4CFF4145B6FA4C" docLanguage="en" docName="zt03986p216.pdf" docOrigin="Zootaxa 3986 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glossogobius multipapillus Hoese &amp; Allen, 2015, n. sp." docType="treatment" docUuid="E39E4210-7D95-4A3B-9C4A-513BBF030943" docUuidSource="ZooBank" docVersion="9" lastPageNumber="210" masterDocId="FFFBFFEBFFCDFFB7FFDBFFD64417FF8C" masterDocTitle="Descriptions of three new species of Glossogobius (Teleostei: Gobiidae) from New Guinea" masterLastPageNumber="216" masterPageNumber="201" pageNumber="208" updateTime="1698609731161" updateUser="plazi">
<mods:mods id="DA61DF7E45455FF0722222C45DA047D6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7AC1BDB1F4948C0B5C3C1B8DFA98511F">Descriptions of three new species of Glossogobius (Teleostei: Gobiidae) from New Guinea</mods:title>
</mods:titleInfo>
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<mods:role id="64D1585825CBDEBF1EFC4A0537B341EA">
<mods:roleTerm id="1319CD1A05280ECC660014A672DE931A">Author</mods:roleTerm>
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<mods:namePart id="BF3006E2320878B256A463C65916F716">Hoese, Douglass F.</mods:namePart>
</mods:name>
<mods:name id="7C0A5D2D10D18AB4DC0EE47C7988522B" type="personal">
<mods:role id="983ED45463E2D005618157DB6B6F96AC">
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<mods:namePart id="9893D440A9D64513C446203ADF463F85">Allen, Gerald R.</mods:namePart>
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<mods:typeOfResource id="322C1F5ABA45C16C39EF56B389295647">text</mods:typeOfResource>
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<mods:title id="006ED62E3D0C69C30240289758C6D89D">Zootaxa</mods:title>
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<mods:date id="909FD256B2A3AC8949C4DB55848B1FAA">2015</mods:date>
<mods:detail id="D47F17F2268CD40F3087D19BD21B7977" type="volume">
<mods:number id="8384DAF648E534F9B665A650D0853AD6">3986</mods:number>
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<mods:classification id="752BA484FE1D3F3E1ED075D69FFE2A07">journal article</mods:classification>
<mods:identifier id="A69978CD3EA6234868EB956A71761D91" type="DOI">10.11646/zootaxa.3986.2.4</mods:identifier>
<mods:identifier id="72BF5618A6C16515CBA4B392E730D7D9" type="GBIF-Dataset">99ab0619-d0df-4186-a2be-5198484ca988</mods:identifier>
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</mods:mods>
<treatment id="03C28793FFCAFFBEFF4CFF4145B6FA4C" ID-DOI="http://doi.org/10.5281/zenodo.6093995" ID-GBIF-Taxon="119637136" ID-Zenodo-Dep="6093995" LSID="urn:lsid:zoobank.org:act:E39E4210-7D95-4A3B-9C4A-513BBF030943" httpUri="http://treatment.plazi.org/id/03C28793FFCAFFBEFF4CFF4145B6FA4C" lastPageId="9" lastPageNumber="210" pageId="7" pageNumber="208">
<subSubSection id="C371650EFFCAFFB0FF4CFF4147B3FEB0" pageId="7" pageNumber="208" type="nomenclature">
<paragraph id="8BD43685FFCAFFB0FF4CFF41463EFF3E" blockId="7.[151,553,151,247]" box="[151,553,151,178]" pageId="7" pageNumber="208">
<heading id="D09C81E9FFCAFFB0FF4CFF41463EFF3E" bold="true" box="[151,553,151,178]" fontSize="11" level="1" pageId="7" pageNumber="208" reason="1">
<emphasis id="B91FEA97FFCAFFB0FF4CFF41463EFF3E" bold="true" box="[151,553,151,178]" pageId="7" pageNumber="208">
<taxonomicName id="4C6B4D06FFCAFFB0FF4CFF4145C9FF3D" box="[151,478,151,177]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="multipapillus" status="sp. nov.">
<emphasis id="B91FEA97FFCAFFB0FF4CFF4145C9FF3D" bold="true" box="[151,478,151,177]" italics="true" pageId="7" pageNumber="208">Glossogobius multipapillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22C57ECFFCAFFB0FE3DFF4E463EFF3E" box="[486,553,152,178]" pageId="7" pageNumber="208" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD43685FFCAFFB0FF4CFF6C4529FF5F" blockId="7.[151,553,151,247]" box="[151,318,186,211]" pageId="7" pageNumber="208">Papillate Goby</paragraph>
<paragraph id="8BD43685FFCAFFB0FF4CFF0845A6FF7B" blockId="7.[151,553,151,247]" box="[151,433,222,247]" pageId="7" pageNumber="208">
<figureCitation id="13502A00FFCAFFB0FF4CFF0844F3FF7B" box="[151,228,222,247]" captionStart="FIGURE 2" captionStartId="4.[151,250,1954,1976]" captionTargetBox="[309,1256,221,1906]" captionTargetId="figure@4.[294,1292,194,1933]" captionTargetPageId="4" captionText="FIGURE 2. Sensory papillae of three species of Glossogobius: a. Glossogobius macrocephalus, based on holotype WAMP. 26972 – 003 and paratype WAMP. 26972 – 003; b. Glossogobius multipapillus, composite based on holotype and several Paratypes; c. Glossogobius sentaniensis, based largely on holotype, WAMP. 27852 – 004." httpUri="https://zenodo.org/record/245932/files/figure.png" pageId="7" pageNumber="208">Figs. 2</figureCitation>
b, 5, 6,
<tableCitation id="C6E9033EFFCAFFB0FEECFF0845A6FF7B" box="[311,433,222,247]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStart-5="TABLE 6" captionStartId-0="2.[151,239,1621,1644]" captionStartId-1="3.[151,239,152,175]" captionStartId-2="3.[151,239,538,561]" captionStartId-3="3.[151,239,925,948]" captionStartId-4="3.[151,239,1312,1335]" captionStartId-5="3.[151,239,1699,1722]" captionText-0="TABLE 1. Second dorsal fin-ray counts of 3 species of Glossogobius." captionText-1="TABLE 2. Anal fin-ray counts of 3 species of Glossogobius." captionText-2="TABLE 3. Pectoral fin-ray counts of 3 species of Glossogobius." captionText-3="TABLE 4. Predorsal scale counts of 3 species of Glossogobius." captionText-4="TABLE 5. Longitudinal scale count for three species of Glossogobius." captionText-5="TABLE 6. Transvers scale count for three species of Glossogobius." pageId="7" pageNumber="208">Tables 1–6</tableCitation>
</paragraph>
<paragraph id="8BD43685FFCAFFB0FF4CFEF047B3FEB0" blockId="7.[151,932,294,316]" box="[151,932,294,316]" pageId="7" pageNumber="208">
<treatmentCitationGroup id="AB7B11ABFFCAFFB0FF4CFEF047B3FEB0" box="[151,932,294,316]" pageId="7" pageNumber="208">
<taxonomicName id="4C6B4D06FFCAFFB0FF4CFEF04536FEB0" ID-CoL="4P99" box="[151,289,294,316]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="208" phylum="Chordata" rank="genus">
<emphasis id="B91FEA97FFCAFFB0FF4CFEF04536FEB0" box="[151,289,294,316]" italics="true" pageId="7" pageNumber="208">Glossogobius</emphasis>
</taxonomicName>
sp. 4. —
<treatmentCitation id="0ACA1094FFCAFFB0FE58FEF04627FEB0" author="Allen" box="[387,560,294,316]" page="184" pageId="7" pageNumber="208" year="1991">
<bibRefCitation id="EFFA4B74FFCAFFB0FE58FEF045E9FEB0" author="Allen" box="[387,510,294,316]" pageId="7" pageNumber="208" refString="Allen, G. R. (1991) Field Guide to the Freshwater Fishes of New Guinea. Publication No. 9. Christensen Research Institute, Madang, 268 pp." type="book" year="1991">Allen, 1991</bibRefCitation>
: 184
</treatmentCitation>
, pl. 15, fig. 11 (
<collectingCountry id="F37C7615FFCAFFB0FD14FEF0478EFEB0" box="[719,921,294,316]" name="Papua New Guinea" pageId="7" pageNumber="208">Papua New Guinea</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C371650EFFCAFFB0FF4CFEBF459BFCDA" pageId="7" pageNumber="208" type="materials_examined">
<paragraph id="8BD43685FFCAFFB0FF4CFEBF44C1FDF2" blockId="7.[151,1437,361,2006]" pageId="7" pageNumber="208">
<emphasis id="B91FEA97FFCAFFB0FF4CFEBF451CFE0D" bold="true" box="[151,267,361,385]" pageId="7" pageNumber="208">
<typeStatus id="54D08827FFCAFFB0FF4CFEBF4511FE0D" box="[151,262,361,385]" pageId="7" pageNumber="208" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED7AAE40FFCAFFB0FEC9FEBF4541FE0D" LSID="urn:lsid:biocol.org:col:34981" box="[274,342,361,385]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27790–009,
<quantity id="4C939B60FFCAFFB0FE27FEBF465CFE0D" box="[508,587,361,386]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="208" unit="mm" value="56.0">56 mm</quantity>
SL male, Kaili Creek,
<quantity id="4C939B60FFCAFFB0FC97FEBF4784FE0E" box="[844,915,361,386]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="208" unit="km" value="12.0">12 km</quantity>
E of Kokoda (
<geoCoordinate id="EE5F5042FFCAFFB0FBE3FEBF4093FE0E" box="[1080,1156,361,386]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.916667">8°55'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FB4BFEBF40EFFE0D" box="[1168,1272,361,385]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="147.78334">147°47'E</geoCoordinate>
), G. Allen,
<date id="FFD51045FFCAFFB0FA5BFEBF44C4FE2A" pageId="7" pageNumber="208" value="2012-09">12 Sept.</date>
, 1982.
<typeStatus id="54D08827FFCAFFB0FEFDFE5B4589FE29" box="[294,414,397,421]" pageId="7" pageNumber="208" type="paratype">
<emphasis id="B91FEA97FFCAFFB0FEFDFE5B4589FE29" bold="true" box="[294,414,397,421]" pageId="7" pageNumber="208">Paratypes</emphasis>
</typeStatus>
AMS I.234328–002, (ex
<collectionCode id="ED7AAE40FFCAFFB0FD18FE58471FFE2A" LSID="urn:lsid:biocol.org:col:34981" box="[707,776,398,422]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27785–003), 16(36–49), Safia Creek,
<quantity id="4C939B60FFCAFFB0FB08FE5B411AFE2A" box="[1235,1293,397,422]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="208" unit="km" value="5.0">5 km</quantity>
NE of Safia station, (
<geoCoordinate id="EE5F5042FFCAFFB0FF22FE674552FE45" box="[249,325,433,457]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-9.566667">9°34'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FE89FE6745ABFE45" box="[338,444,433,457]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.66667">148°40'E</geoCoordinate>
), G. Allen,
<date id="FFD51045FFCAFFB0FDE6FE67468BFE45" box="[573,668,433,457]" pageId="7" pageNumber="208" value="2010-09">10 Sept.</date>
, 1982; AMS I.25872–002, 2 (26–39), Safia, Bulmacau cattle station Creek (
<geoCoordinate id="EE5F5042FFCAFFB0FF31FE00452AFE62" box="[234,317,469,494]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.75">8° 45'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FE93FE0045A4FE62" box="[328,435,470,494]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.33333">148°20'E</geoCoordinate>
), W. Ivantsoff and J. Paska,
<date id="FFD51045FFCAFFB0FD34FE03475AFE62" box="[751,845,469,494]" pageId="7" pageNumber="208" value="1916-09">16 Sept.</date>
, 1985;
<collectionCode id="ED7AAE40FFCAFFB0FC44FE0047E7FE62" LSID="urn:lsid:biocol.org:col:34871" box="[927,1008,470,494]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="208">USNM</collectionCode>
<date id="FFD51045FFCAFFB0FC2CFE03405CFE62" box="[1015,1099,469,494]" pageId="7" pageNumber="208" value="56-05-26">260556</date>
4(31–49), forest stream about
<quantity id="4C939B60FFCAFFB0FF4CFE2F44DEFD9E" box="[151,201,505,530]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="7" pageNumber="208" unit="mi" value="5.0">5 mi</quantity>
W of Popondetta (
<geoCoordinate id="EE5F5042FFCAFFB0FE47FE2F45FFFD9D" box="[412,488,505,529]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.783334">8°47'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FE28FE2F4649FD9D" box="[499,606,505,529]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.16667">148°10'E</geoCoordinate>
), T. Roberts,
<date id="FFD51045FFCAFFB0FD29FE2F4756FD9D" box="[754,833,505,529]" pageId="7" pageNumber="208">3 Aug.</date>
, 1975;
<collectionCode id="ED7AAE40FFCAFFB0FC49FE2F47C0FD9D" LSID="urn:lsid:biocol.org:col:34981" box="[914,983,505,529]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27785–003, 40(19–52), Safia Creek,
<quantity id="4C939B60FFCAFFB0FA55FE2F44ACFDBA" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="208" unit="km" value="5.0">5 km</quantity>
NE of Safia station (
<geoCoordinate id="EE5F5042FFCAFFB0FE63FDC84613FDBA" box="[440,516,542,566]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-9.566667">9°34'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FDC8FDC84669FDBA" box="[531,638,542,566]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.66667">148°40'E</geoCoordinate>
), G. Allen,
<date id="FFD51045FFCAFFB0FCDCFDC8477EFDBA" box="[775,873,542,566]" pageId="7" pageNumber="208" value="2010-09">10 Sept.</date>
, 1982;
<collectionCode id="ED7AAE40FFCAFFB0FC1AFDC84011FDBA" LSID="urn:lsid:biocol.org:col:34981" box="[961,1030,542,566]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27790–002, 7(19–48) taken with
<typeStatus id="54D08827FFCAFFB0FF4CFD9744EAFDD6" box="[151,253,577,602]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED7AAE40FFCAFFB0FEDCFD97455BFDD5" LSID="urn:lsid:biocol.org:col:34981" box="[263,332,577,601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27791–003 27, (21–54), tributary,
<quantity id="4C939B60FFCAFFB0FD3DFD97473BFDD6" box="[742,812,577,602]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="km" value="10.0">10 km</quantity>
S of Popondetta (
<geoCoordinate id="EE5F5042FFCAFFB0FC2FFD974057FDD5" box="[1012,1088,577,601]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.783334">8°47'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FB90FD9740A5FDD5" box="[1099,1202,577,601]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.21666">148°13'E</geoCoordinate>
), G. Allen,
<date id="FFD51045FFCAFFB0FAE1FD97418FFDD5" box="[1338,1432,577,601]" pageId="7" pageNumber="208" value="2012-09">12 Sept.</date>
, 1982.
</paragraph>
<paragraph id="8BD43685FFCAFFB0FF1CFD5F459BFCDA" blockId="7.[151,1437,361,2006]" pageId="7" pageNumber="208">
<emphasis id="B91FEA97FFCAFFB0FF1CFD5F45A3FD2E" bold="true" box="[199,436,649,674]" pageId="7" pageNumber="208">
Non–
<typeStatus id="54D08827FFCAFFB0FEDCFD5F452DFD2D" box="[263,314,649,673]" pageId="7" pageNumber="208">type</typeStatus>
material.
</emphasis>
AMS I.46509-001 (ex
<collectionCode id="ED7AAE40FFCAFFB0FD0EFD5C470EFD2E" LSID="urn:lsid:biocol.org:col:34981" box="[725,793,650,674]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27791–003), 2(41–50),
<quantity id="4C939B60FFCAFFB0FB88FD5C40B6FD2E" box="[1107,1185,649,674]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="208" unit="km" value="10.0">10 km</quantity>
south of Popondetta (
<geoCoordinate id="EE5F5042FFCAFFB0FF7BFD7844FBFD4A" box="[160,236,686,710]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.783334">8°47'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FF23FD7B4574FD4A" box="[248,355,685,710]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.1">148°06'E</geoCoordinate>
), G. Allen and D. Coates,
<date id="FFD51045FFCAFFB0FD50FD78473DFD4A" box="[651,810,686,710]" pageId="7" pageNumber="208" value="1982-09-12">12 Sept. 1982</date>
;
<collectionCode id="ED7AAE40FFCAFFB0FCEDFD784790FD4A" LSID="urn:lsid:biocol.org:col:34871" box="[822,903,686,710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="208">USNM</collectionCode>
<date id="FFD51045FFCAFFB0FC55FD7B47F2FD4A" box="[910,997,685,710]" pageId="7" pageNumber="208" value="28-05-26">260528</date>
, 1(41), Ejava and Ovi Creeks about
<quantity id="4C939B60FFCAFFB0FA5BFD7844ACFD66" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="208" unit="km" value="13.0">13 km</quantity>
ESE of Kokoda, T. Roberts,
<date id="FFD51045FFCAFFB0FE24FD07465AFD66" box="[511,589,721,746]" pageId="7" pageNumber="208">5 Aug.</date>
, 1975;
<collectionCode id="ED7AAE40FFCAFFB0FD45FD0446F4FD66" LSID="urn:lsid:biocol.org:col:34981" box="[670,739,722,746]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.26409–002, 1(30), Auga Creek, about
<quantity id="4C939B60FFCAFFB0FB70FD0740F3FD66" box="[1195,1252,721,746]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="208" unit="km" value="5.0">5 km</quantity>
S of Popondetta (
<geoCoordinate id="EE5F5042FFCAFFB0FF7BFD2044E4FC82" box="[160,243,758,782]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.8">8°48' S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FEDBFD20457DFC82" box="[256,362,758,782]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="148.21666">148°13'E</geoCoordinate>
), G. Allen,
<date id="FFD51045FFCAFFB0FE30FD204624FC82" box="[491,563,758,782]" pageId="7" pageNumber="208">4 Oct.</date>
, 1978;
<collectionCode id="ED7AAE40FFCAFFB0FD5DFD2046DCFC82" LSID="urn:lsid:biocol.org:col:34981" box="[646,715,758,782]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.26417–002, 2(19–22), Hunete Creek,
<quantity id="4C939B60FFCAFFB0FB4AFD2040CEFC82" box="[1169,1241,757,782]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="208" unit="km" value="33.0">33 km</quantity>
from Popondetta (
<geoCoordinate id="EE5F5042FFCAFFB0FF7BFCCF44FCFCBE" box="[160,235,793,818]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.85">8°51'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FF22FCCC4576FCBE" box="[249,353,794,818]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-148.0">148°00'S</geoCoordinate>
), G. Allen
<date id="FFD51045FFCAFFB0FE07FCCF4633FCBE" box="[476,548,793,818]" pageId="7" pageNumber="208">5 Oct.</date>
, 1978;
<collectionCode id="ED7AAE40FFCAFFB0FDACFCCC46ABFCBE" LSID="urn:lsid:biocol.org:col:34981" box="[631,700,794,818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="7" pageNumber="208">WAM</collectionCode>
P.27789–004 1 (55) Iara Creek at Kokoda (
<geoCoordinate id="EE5F5042FFCAFFB0FB77FCCF40EFFCBE" box="[1196,1272,793,818]" direction="south" orientation="latitude" pageId="7" pageNumber="208" precision="925" value="-8.883333">8°53'S</geoCoordinate>
,
<geoCoordinate id="EE5F5042FFCAFFB0FADEFCCC4178FCBE" box="[1285,1391,794,818]" direction="east" orientation="longitude" pageId="7" pageNumber="208" precision="925" value="141.73334">141°44'E</geoCoordinate>
), G. Allen,
<date id="FFD51045FFCAFFB0FF38FCE84555FCDA" box="[227,322,830,854]" pageId="7" pageNumber="208" value="2012-09">12 Sept.</date>
, 1982.
</paragraph>
</subSubSection>
<subSubSection id="C371650EFFCAFFB0FF1CFCB7412DF93A" pageId="7" pageNumber="208" type="diagnosis">
<paragraph id="8BD43685FFCAFFB0FF1CFCB747B9FAA6" blockId="7.[151,1437,361,2006]" pageId="7" pageNumber="208">
<emphasis id="B91FEA97FFCAFFB0FF1CFCB74556FCF6" bold="true" box="[199,321,865,890]" pageId="7" pageNumber="208">Diagnosis.</emphasis>
Preoperculum with a blunt triangular projection at angle of preoperculum, covered by skin. Mental fraenum bilobed, with short lateral lobes. Mouth small; reaching to below anterior quarter of eye, upper jaw 12.0– 15.5% SL (12–14% SL in specimens less than
<quantity id="4C939B60FFCAFFB0FD7DFC7F46E1FC4D" box="[678,758,937,962]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="208" unit="mm" value="50.0">50 mm</quantity>
SL, 13–15% in larger specimens); jaws forming an angle of 45° with body axis; upper margin of upper jaw in line with mideye. Cheeks tapering. Interorbital narrow, less than eye length. Predorsal area scaled forward to just behind eyes. Cheek naked. Operculum naked. Pectoral base covered with small cycloid scales. Prepelvic area with moderate sized cycloid scales in about 6 rows covering all but small anterior triangular patch. Belly fully scaled. First dorsal fin spines 4–5 (usually 4) extending beyond other spines when fin depressed. First dorsal fin with basal half black, sometimes with a light area posteriorly. Pelvic fin, length greater than width; reaching to or slightly beyond anus. Second dorsal rays usually I,8; anal rays usually I,7; pectoral rays usually 17–18; longitudinal scale count 25–28; predorsal scale count 12–16; transverse scale count (TRB) 8.5; gill rakers on outer face of first arch 0–1+1+6–8, raker on upper part of arch rarely present; lower gill rakers on outer face of second arch 6–7; branched caudal rays usually 13–14 (
<quantity id="4C939B60FFCAFFB0FC27FB384022FA8A" box="[1020,1077,1261,1286]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="7" pageNumber="208" unit="in" value="13.0">13 in</quantity>
specimens less than
<quantity id="4C939B60FFCAFFB0FAC4FB384179FA89" box="[1311,1390,1262,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="208" unit="mm" value="20.0">20 mm</quantity>
SL, usually
<quantity id="4C939B60FFCAFFB0FF34FAC4453FFAA6" box="[239,296,1297,1322]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="7" pageNumber="208" unit="in" value="14.0">14 in</quantity>
specimens greater than
<quantity id="4C939B60FFCAFFB0FDEDFAC44692FAA5" box="[566,645,1298,1322]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="208" unit="mm" value="40.0">40 mm</quantity>
SL); vertebrae 11+16=27.
</paragraph>
<paragraph id="8BD43685FFCAFFB0FF1CFAE3412DF93A" blockId="7.[151,1437,361,2006]" pageId="7" pageNumber="208">
<emphasis id="B91FEA97FFCAFFB0FF1CFAE34531FAC1" box="[199,294,1333,1357]" italics="true" pageId="7" pageNumber="208">Papillae</emphasis>
. Line 1 (before nasal pore) composed on 3–4 rows of papillae. Line 2 (between nasal pores) composed of 3 rows of papillae, not connecting to line 2 on opposite side of head. Line 5 (suborbital) composed of 2–3 rows of papillae. Line 6 (suborbital branch) composed of 1–3 pairs of papillae in very short line. Lines 7, 9, 10 (LL cheek rows) composed on 2, 3 and 3 rows of papillae respectively, papillae in 3 lines lining up vertically to produce vertical rows, similar to VT lines (multiple longitudinal pattern of
<bibRefCitation id="EFFA4B74FFCAFFB0FC3EFA104061FA52" author="Hoese" box="[997,1142,1478,1502]" pageId="7" pageNumber="208" refString="Hoese, D. F. (1983) Sensory papilla patterns of the cheek lateralis system in the gobiid fishes Acentrogobius and Glossogobius, and their significance for the classification of gobioid fishes. Records of the Australian Museum, 35, 223 - 229. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.35.1983.319" type="journal article" year="1983">Hoese, 1983</bibRefCitation>
). Lines 8 and 11 (LT row) ending before posterior preopercular margin. Line 12 (Outer POP–mandibular) composed of single row of papillae, without gap behind lips. Line 13 (Inner POP–mandibular) composed of 2–3 rows of papillae. Line 20 (opercular VT) composed of single row of papillae dorsally, becoming double ventrally. Line 21 (Upper opercular OT) composed of single row of papillae, curved, extending along posterior margin of operculum and joining line 22. Line 22 (Lower opercular OT) composed of single row of papillae, without any branches. Several vertical papillae rows on belly. A single curved line anteriorly on most body scales (often obscure dorsally and posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C371650EFFCAFFBEFF1CF917463BFCB4" lastPageId="9" lastPageNumber="210" pageId="7" pageNumber="208" type="description">
<paragraph id="8BD43685FFCAFFB0FF1CF9174500F8CA" blockId="7.[151,1437,361,2006]" pageId="7" pageNumber="208">
<emphasis id="B91FEA97FFCAFFB0FF1CF917454EF956" bold="true" box="[199,345,1729,1754]" pageId="7" pageNumber="208">Description.</emphasis>
Based on
<specimenCount id="9D6DFD0CFFCAFFB0FE08F914466DF956" box="[467,634,1729,1754]" pageId="7" pageNumber="208" type="generic">114 specimens</specimenCount>
<quantity id="4C939B60FFCAFFB0FD58F91746E9F955" box="[643,766,1729,1754]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="5.6" metricValueMin="2.6" pageId="7" pageNumber="208" unit="mm" value="41.0" valueMax="56.0" valueMin="26.0">26–56 mm</quantity>
SL. First dorsal spines 6(30*); gill rakers on outer face of first arch 0+1+6(13), 0+1+7(12), 0+1+8(3*), 1+1+6(1), 1+1+7(1), 1+1+8(1); segmented caudal rays 17(27*); branched caudal rays 7/6(12), 7/7 (16), 8/7(3*); vertebrae 11+16(11*).. Counts of other meristic characters given in
<tableCitation id="C6E9033EFFCAFFB0FF4CF8FB4504F8CA" box="[151,275,1837,1862]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStart-5="TABLE 6" captionStartId-0="2.[151,239,1621,1644]" captionStartId-1="3.[151,239,152,175]" captionStartId-2="3.[151,239,538,561]" captionStartId-3="3.[151,239,925,948]" captionStartId-4="3.[151,239,1312,1335]" captionStartId-5="3.[151,239,1699,1722]" captionText-0="TABLE 1. Second dorsal fin-ray counts of 3 species of Glossogobius." captionText-1="TABLE 2. Anal fin-ray counts of 3 species of Glossogobius." captionText-2="TABLE 3. Pectoral fin-ray counts of 3 species of Glossogobius." captionText-3="TABLE 4. Predorsal scale counts of 3 species of Glossogobius." captionText-4="TABLE 5. Longitudinal scale count for three species of Glossogobius." captionText-5="TABLE 6. Transvers scale count for three species of Glossogobius." pageId="7" pageNumber="208">Tables 1–6</tableCitation>
.
</paragraph>
<paragraph id="8BD43685FFCAFFBFFF1CF887461BFDEC" blockId="7.[151,1437,361,2006]" lastBlockId="8.[151,1437,151,608]" lastPageId="8" lastPageNumber="209" pageId="7" pageNumber="208">
Head depressed, length 31.5–35.2% SL, width 17.5–20.2% SL, depth 16.0–18.4% SL. Snout short, rounded in dorsal view; slightly convex to straight in side view; 8.4–11.3% SL. Eye about 1.0–
<quantity id="4C939B60FFCAFFB0FBE4F8A04069F802" box="[1087,1150,1909,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="208" unit="in" value="1.2">1.2 in</quantity>
snout, 8.8–10.8% SL (9– 11% SL in specimens less than
<quantity id="4C939B60FFCAFFB0FDD7F84F4649F83D" box="[524,606,1945,1970]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="208" unit="mm" value="50.0">50 mm</quantity>
SL, 8–10% SL in larger specimens. Small bump below anterior nostril absent, area covered with papillae. Anterior nostril at end of short tube, just above upper lip. Posterior nostril with raised rim, midway between lips and eye and 2 nostril diameters from anterior nostril. Preoperculum short, distance from end of eye to upper posterior preopercular margin less than eye length. Postorbital short, subequal to distance from tip of snout to mideye. Gill opening reaching to below or just behind posterior preoperculum margin. Teeth in upper jaw: outer row of teeth conical, slightly enlarged and wide-set, 2 inner rows of smaller depressible, inwardly directed teeth, innermost row teeth larger than those in middle row. Teeth in lower jaw: teeth in outer row conical, slightly enlarged and wide-set anteriorly, 2 inner rows of smaller depressible teeth, teeth in innermost row slightly larger than teeth in middle row. Tongue tip bilobed. Gill rakers on outer face of first arch short and broad; less than one–third filament length. Rakers on inner face of first arch and other arches short and denticulate. Body covered mostly with large ctenoid scales, cycloid on predorsal region (some predorsal scales ctenoid in specimens less than
<quantity id="4C939B60FFC5FFBFFF4CFE0A44F2FE78" box="[151,229,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="209" unit="mm" value="30.0">30 mm</quantity>
SL), midbelly, pectoral base and prepelvic area. First dorsal fin low, with rounded margin. Second dorsal fin slightly higher than first dorsal fin. Anal fin height slightly less than second dorsal height. Pectoral fin with rounded margin, reaching to above a point just before anal origin. Pelvic disc, thin, oval, reaching to anus; fifth ray with 11 terminal tips in
<typeStatus id="54D08827FFC5FFBFFE79FD91461FFDEC" box="[418,520,583,608]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF14660DFFC5FFBFFF4CFB8A4057FBFE" box="[151,1088,1116,1138]" httpUri="https://zenodo.org/record/245935/files/figure.png" pageId="8" pageNumber="209" targetBox="[151,1436,661,1094]" targetPageId="8">
<paragraph id="8BD43685FFC5FFBFFF4CFB8A4057FBFE" blockId="8.[151,1088,1116,1138]" box="[151,1088,1116,1138]" pageId="8" pageNumber="209">
<emphasis id="B91FEA97FFC5FFBFFF4CFB8A4505FBFE" bold="true" box="[151,274,1116,1138]" pageId="8" pageNumber="209">FIGURE 5.</emphasis>
Holotype of
<taxonomicName id="4C6B4D06FFC5FFBFFE42FB8A46B8FBFE" box="[409,687,1116,1138]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="multipapillus">
<emphasis id="B91FEA97FFC5FFBFFE42FB8A46B8FBFE" box="[409,687,1116,1138]" italics="true" pageId="8" pageNumber="209">Glossogobius multipapillus</emphasis>
</taxonomicName>
, WAM P.27790–009, 56 mm SL, male.
</paragraph>
</caption>
<caption id="DF14660DFFC5FFBFFF4CF971410EF931" box="[151,1305,1703,1725]" httpUri="https://zenodo.org/record/245936/files/figure.png" pageId="8" pageNumber="209" targetBox="[151,1394,1188,1680]" targetPageId="8">
<paragraph id="8BD43685FFC5FFBFFF4CF971410EF931" blockId="8.[151,1305,1703,1725]" box="[151,1305,1703,1725]" pageId="8" pageNumber="209">
<emphasis id="B91FEA97FFC5FFBFFF4CF9714505F930" bold="true" box="[151,274,1703,1725]" pageId="8" pageNumber="209">FIGURE 6.</emphasis>
Photo of freshly collected specimen of
<taxonomicName id="4C6B4D06FFC5FFBFFD7EF97147ACF931" box="[677,955,1703,1725]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="multipapillus">
<emphasis id="B91FEA97FFC5FFBFFD7EF97147ACF931" box="[677,955,1703,1725]" italics="true" pageId="8" pageNumber="209">Glossogobius multipapillus</emphasis>
</taxonomicName>
, from Popondetta, photo G. Allen.
</paragraph>
</caption>
<paragraph id="8BD43685FFC5FFBFFF1CF93C45E3F83A" blockId="8.[151,1437,1770,1974]" pageId="8" pageNumber="209">
<emphasis id="B91FEA97FFC5FFBFFF1CF93C455EF88E" box="[199,329,1770,1794]" italics="true" pageId="8" pageNumber="209">Head pores</emphasis>
. nasal pore above and slightly behind posterior nostril; anterior interorbital pore present posterior interorbital pore present; postorbital pore behind eye present; infraorbital pore below postorbital present; lateral canal pore above preoperculum midway between eye and posterior preopercular margin present; lateral canal pore above posterior preopercular margin absent; terminal lateral canal pore above anterior operculum present; short tube above operculum, with pore at each end absent; 3 preopercular pores, upper in line with lower margin of eyes; widely separated from lower 2.
</paragraph>
<paragraph id="8BD43685FFC4FFBEFF1CFF4E47AEFDEC" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
<emphasis id="B91FEA97FFC4FFBEFF1CFF4E45ACFF3C" box="[199,443,152,176]" italics="true" pageId="9" pageNumber="210">Coloration in alcohol</emphasis>
. Head and body light brown. Head with scattered irregularly shaped brown spots; small dark brown spot behind eye above posterior end of preoperculum, much smaller than pupil diameter; slightly larger brown spot at dorsoposterior attachment of operculum; stripe from anteroventral margin of eye to anterior end of lips; lips with dark brown mottling, posterior tip of lips white. Body scales edged in dark brown (except ventrally on belly); some small spots and mottling dorsally; body with about six thin brown stripes; midline with large rectangular spots; first below first dorsal fin covering 5 scales longitudinally; second spot below second dorsal origin covering 2–3 scales, smaller spot covering 1–2 scales below middle of second dorsal fin; an elongate spot covering 2–3 spots below insertion of second dorsal fin; last spot on end of caudal peduncle covering two scale rows and connected to a thin vertical bar on caudal fin base. Pectoral base with a brown spot dorsally, rest of fin dusky to light brown. First dorsal fin with dark brown stripe covering basal half of fin; distal half whitish; second dorsal fin with small black spots forming 3 rows, distal third of fin whitish, distal tip brown; anal fin uniformly brown; caudal fin with 4–6 wavy brown bands, except ventrally; pelvic fin dark brown; mental fraenum with dark brown tips, rest of ventral surface of head with scattered melanophores.
</paragraph>
<paragraph id="8BD43685FFC4FFBEFF1CFDBA463BFCB4" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
<emphasis id="B91FEA97FFC4FFBEFF1CFDBA4655FD08" box="[199,578,620,644]" italics="true" pageId="9" pageNumber="210">Coloration when freshly collected</emphasis>
(
<figureCitation id="13502A00FFC4FFBEFD8AFDBA4681FD09" box="[593,662,620,645]" captionStart="FIGURE 6" captionStartId="8.[151,250,1703,1725]" captionTargetBox="[151,1394,1188,1680]" captionTargetId="figure@8.[151,1436,1188,1681]" captionTargetPageId="8" captionText="FIGURE 6. Photo of freshly collected specimen of Glossogobius multipapillus, from Popondetta, photo G. Allen." httpUri="https://zenodo.org/record/245936/files/figure.png" pageId="9" pageNumber="210">Fig. 6</figureCitation>
): Head, body, and pectoral-fin base yellowish tan with dark brown to nearly blackish markings as described above for preserved specimens; first dorsal fin with blackish stripe covering basal part of fin; distal half pale yellow to whitish with reddish apical tip; second dorsal fin with small black spots forming 3 rows on basal half, margin of fin broadly dusky reddish with white submarginal stripe; anal fin dusky brown to blackish, paler basally; caudal fin slightly yellowish to dusky grey brown with 4–6 wavy brown bands on central portion; pelvic fin dark grey.
</paragraph>
</subSubSection>
<subSubSection id="C371650EFFC4FFBEFF1CFC954766FB2C" pageId="9" pageNumber="210" type="distribution">
<paragraph id="8BD43685FFC4FFBEFF1CFC954622FC61" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
<emphasis id="B91FEA97FFC4FFBEFF1CFC95454EFCD0" bold="true" box="[199,345,835,860]" pageId="9" pageNumber="210">Distribution</emphasis>
. The species is restricted to the northeastern side of the Owen Stanley Range in Northern Province of
<collectingCountry id="F37C7615FFC4FFBEFEF6FCBE4601FC0C" box="[301,534,871,896]" name="Papua New Guinea" pageId="9" pageNumber="210">Papua New Guinea</collectingCountry>
, approximately between 8°00' and
<geoCoordinate id="EE5F5042FFC4FFBEFC1AFCBE4030FC0C" box="[961,1063,872,896]" direction="south" orientation="latitude" pageId="9" pageNumber="210" precision="925" value="-9.666667">9° 40' S</geoCoordinate>
latitude. It normally occurs in moderately-flowing, clear rainforest creeks and shallow rivers on sand or gravel bottoms. The
<typeStatus id="54D08827FFC4FFBEFB34FC5B4108FC29" box="[1263,1311,909,933]" pageId="9" pageNumber="210">type</typeStatus>
locality is situated in foothill terrain at an elevation of
<quantity id="4C939B60FFC4FFBEFD59FC7946DCFC4B" box="[642,715,943,968]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="9" pageNumber="210" unit="m" value="425.0">425 m</quantity>
, but the species also ranges to much lower altitudes closer to the coast, or about
<quantity id="4C939B60FFC4FFBEFE99FC02456CFC60" box="[322,379,980,1005]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="210" unit="m" value="50.0">50 m</quantity>
above sea level.
</paragraph>
<paragraph id="8BD43685FFC4FFBEFF1CFC214766FB2C" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
<emphasis id="B91FEA97FFC4FFBEFF1CFC214603FB9C" bold="true" box="[199,532,1015,1040]" pageId="9" pageNumber="210">Similarity to other species:</emphasis>
The unique papillae pattern, coloration, fin–ray counts and flat preopercular projection easily separate this species from other species of
<taxonomicName id="4C6B4D06FFC4FFBEFC98FBCA47CBFBB8" box="[835,988,1052,1076]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91FEA97FFC4FFBEFC98FBCA47CBFBB8" box="[835,988,1052,1076]" italics="true" pageId="9" pageNumber="210">Glossogobius</emphasis>
</taxonomicName>
. Only
<taxonomicName id="4C6B4D06FFC4FFBEFBF0FBCA417CFBB8" box="[1067,1387,1052,1076]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="circumspectus">
<emphasis id="B91FEA97FFC4FFBEFBF0FBCA417CFBB8" box="[1067,1387,1052,1076]" italics="true" pageId="9" pageNumber="210">Glossogobius circumspectus</emphasis>
</taxonomicName>
and some species from the Malili Lakes in Sulawesi have vertical rows of papillae on the cheek, but those species have an indistinct mental fraenum.
<taxonomicName id="4C6B4D06FFC4FFBEFE2EFBB2473BFBF0" box="[501,812,1124,1148]" class="Actinopterygii" family="Gobiidae" genus="Glossogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="multipapillus">
<emphasis id="B91FEA97FFC4FFBEFE2EFBB2473BFBF0" box="[501,812,1124,1148]" italics="true" pageId="9" pageNumber="210">Glossogobius multipapillus</emphasis>
</taxonomicName>
has the lobed mental fraenum, characteristic of most inland species in New
<collectingCountry id="F37C7615FFC4FFBEFE4FFB5145FEFB2C" box="[404,489,1159,1184]" name="Guinea-Bissau" pageId="9" pageNumber="210">Guinea</collectingCountry>
. Immediate relatives are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C371650EFFC4FFBEFF1CFB7A4651FAF4" pageId="9" pageNumber="210" type="discussion">
<paragraph id="8BD43685FFC4FFBEFF1CFB7A45A0FB64" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
<emphasis id="B91FEA97FFC4FFBEFF1CFB7A452AFB48" bold="true" box="[199,317,1196,1220]" pageId="9" pageNumber="210">Remarks:</emphasis>
Specimens from Safia average significantly fewer pectoral rays than do specimens from Kokoda and Popondetta (
<tableCitation id="C6E9033EFFC4FFBEFE88FB1945BEFB64" box="[339,425,1231,1256]" captionStart="TABLE 3" captionStartId="3.[151,239,538,561]" captionText="TABLE 3. Pectoral fin-ray counts of 3 species of Glossogobius." pageId="9" pageNumber="210">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BD43685FFC4FFBEFF1CFB224651FAF4" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
The broad rounded urogenital papilla of females is distinctive from the pointed papilla of males at around
<quantity id="4C939B60FFC4FFBEFA5BFB2244D4FABC" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="210" unit="mm" value="30.0">30 mm</quantity>
SL and larger. Of the known specimens approximately equal number of males and females were found (male to female ration of 1.2), with the largest male
<quantity id="4C939B60FFC4FFBEFD4FFAEA46F4FAD8" box="[660,739,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="210" unit="mm" value="56.0">56 mm</quantity>
SL and the largest female
<quantity id="4C939B60FFC4FFBEFBD0FAEA404DFAD8" box="[1035,1114,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="210" unit="mm" value="54.0">54 mm</quantity>
SL. Nothing is known of the biology or reproduction in the species.
</paragraph>
</subSubSection>
<subSubSection id="C371650EFFC4FFBEFF1CFA5245B6FA4C" pageId="9" pageNumber="210" type="etymology">
<paragraph id="8BD43685FFC4FFBEFF1CFA5245B6FA4C" blockId="9.[151,1437,151,1472]" pageId="9" pageNumber="210">
<emphasis id="B91FEA97FFC4FFBEFF1CFA524559FA10" bold="true" box="[199,334,1412,1436]" pageId="9" pageNumber="210">Etymology.</emphasis>
An arbitrary combination of letters referring to the multiple papillae rows on the cheek and treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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